Severe Grazing Research Articles

The impact of different grazing intensity and management measures on soil organic carbon density in Zhangye grassland

Studying the spatial and temporal changes of grassland soil organic carbon (SOC) is helpful in promote the management of regional ecosystem carbon sinks. Grazing is one of the main ways of rational utilization of grassland. Different grazing intensities will affect the change of SOC density. Under different grazing intensity and management measures in Zhangye grassland, this study uses the parameter localized CENTURY model to simulate the temporal and spatial variations of SOC density from 1970 to 2022. The results showed that long-term light grazing reduced the average SOC by 195.114 g·m−2 and 1.91%. Moderate and severe grazing, respectively, for a long time made the total SOC density loss of 5.21% and 17.69%. In a short period, mild and moderate grazing reduced total SOC first and then increased it. Under light grazing, total SOC density appeared higher relative changes in the southeast, and lower in the northwest and central. There was no significant difference in the relative changes of total SOC between steppe and desert grasslands under light grazing. The decrease range of steppe was gradually greater than that in desert grassland. Since different management measures were implemented in some sampling sites in 2017, we divided the study period into two periods, 1970–2016 and 2017–2022. The implementation of degraded grassland improvement, fallow grazing, and rotational grazing would increase the total SOC density and mild SOC density, rotational grazing > degraded grassland improvement > rest grazing. Rotational grazing and the improvement of degraded grassland increased the density of active and inert SOC, while resting grazing decreased the density of SOC.

Impacts of Grazing on Vegetation and Soil Physicochemical Properties in Northern Yinshan Mountain Grasslands

The grasslands at the northern foot of the Yinshan Mountains are an integral component of the northern grassland ecosystem in China. The ecosystem in this region has low stability and poor resistance to disturbance. In this study, experiments were conducted to evaluate the possible changes in vegetation community structure and soil physicochemical properties due to overgrazing in the grasslands. Completely randomized group experiments were designed with grazing intensity as the single-factor study was conducted using natural grassland located in Xilamuren (in Inner Mongolia, northern China) as the study area. Three blocks were created, each having four plots of different grazing intensities and each block having an area of 100 m × 100 m. The experiments were conducted to evaluate the possible variances both in the structure of the vegetation community and the soil physicochemical properties resulting from overgrazing in the grasslands at the northern foot of Yinshan Mountain. The results were as follows: The importance values of dominant species, such as Heteropappus altaicus and Artimisia gmelinii, exhibited varying degrees of change with an increase in the grazing intensity. The surface vegetation cover decreased significantly with an increase in the grazing intensity. The increasing grazing intensity led to a significant increase in the content of very coarse sand grains in the soil. Severe grazing increased the exposed surface area, intensified the effects of blowing wind and scouring action of water, and led to the coarsening of topsoil particles. At 0–5 cm depth, the bulk density of soil exhibited an increasing tendency with an increase in the grazing intensity. The organic matter content of the soil in the heavily grazed plots decreased by 11.74%, 11.00%, and 14.08%, respectively, when compared to that in the 0–40 cm soil layer with no grazing, light grazing, and moderate grazing. The results emphasized the importance of managing grazing intensity for soil and vegetation restoration. Thus, the effects of short-term grazing (for example, 5 years) on vegetation community composition and species diversity may be less pronounced. This study contributes to our understanding of pasture management and the restoration of grassland species diversity.

Effects of corn silage supplementation strategy and grazing intensity on herbage intake, milk production, and behavior of dairy cows

Effects of corn silage supplementation on milk production of grazing dairy cows depend in part on the substitution rate between the 2 forages, which may be influenced by grazing management. The aim of this study was to compare 2 grazing management strategies for measuring substitution rate between herbage and corn silage, in interaction with grazing intensity. Six treatments were compared, with 2 grazing intensities and 3 supplementation strategies investigated at both grazing intensities. The 2 grazing intensities were severe and light grazing, defined by either (1) herbage allowance (HA) of 15 (severe) or 30 (light) kg dry matter (DM)/cow per d at 3 cm above ground level or (2) postgrazing sward height, depending on the supplementation strategy. The 3 supplementation strategies were as follows: (U) an unsupplemented treatment, (A) 5 kg DM/d of corn silage offered at a similar HA as in U, and (H) 5 kg DM/d of corn silage offered at a similar postgrazing sward height as in U. Thirty-six multiparous Holstein cows were used in a randomized complete block design and divided in 2 groups for the entire experiment, one for each grazing intensity. Within each grazing intensity group, the corn silage supplementation strategy was studied using a 3 × 3 Greco-Latin square design, with 3 periods of 14 d. Supplementing cows with corn silage increased total DM intake only for severe grazing by 1.7 kg DM/d. The substitution rate between corn silage and grazed herbage was lower for severe than for light grazing, averaging 0.63 and 1.23, respectively. Herbage dry matter intake was lower by 1.2 kg/d for strategy H than A, leading to lower substitution rates (0.81 vs. 0.99, respectively), irrespective of grazing intensity. Milk production increased with silage supplementation for severe grazing (+1.0 kg/d milk) and was unaffected by silage supplementation for light grazing (-0.4 kg/d milk). The milk production response to corn silage supplementation averaged +0.23 and -0.08 kg of milk per kg DM of silage for severe and light grazing, respectively. Fat-corrected milk production tended to be lower by 0.4 kg/d for strategy H than A, leading to lower milk production response (+0.00 vs. +0.12 kg of milk per kg DM of silage, respectively). Milk protein concentration increased with silage supplementation for severe grazing (+1.0 g/kg) but decreased with silage supplementation for light grazing (-0.6 g/kg). Milk fat concentration did not differ among treatments. On average, daily grazing time (-47 min/d, i.e., -9%) and herbage intake rate (-4.9 g of DM/min, i.e., -14%) decreased when cows were supplemented, with greater grazing time reduction at severe than light grazing, and greater herbage intake rate reduction at light than severe grazing. In conclusion, the greater substitution rate and the lower 4% fat-corrected milk production when corn silage was provided at a similar postgrazing sward height rather than at a similar HA to those of unsupplemented cows explain why supplementing grazing dairy cows with conserved forages has no strong effect in practice from a production point of view.

Deferment associated to contrasting grazing intensities affects root/shoot biomass allocation in natural grasslands

AbstractQuestions(i) How does a strategic deferment influence both shoot and root biomass in South American Campos natural grassland after a long time of contrasting livestock grazing intensities? (ii) How do functional strategies of plants to tolerate or avoid grazing affect patterns of aboveground and belowground biomass?LocationA natural Campos grassland in the Central Depression of Rio Grande do Sul State, Southern Brazil (latitude 30°05′27″ S, longitude 51°40′18″ W, and altitude 46 m).MethodsWe studied the effects of deferments (spring or autumn) at moderate and severe livestock grazing intensities (maintained by adjusting contrasting forage allowances during 25 years) on the cover of dominant species, and aboveground and belowground biomass. The experiment was arranged in a completely randomized design with three replications of grazing exclusions, applied simultaneously at moderate and severe grazing intensities. Measurements of above‐ and belowground biomass and the coverage of the five dominant species were performed before and after grazing exclusion periods.ResultsUnder severe grazing intensity, erect‐growing grasses were replaced by prostrate species, which were able at least to maintain the belowground biomass found at moderate grazing intensity. In response to deferments, severe grazing intensity changed toward a significant increase in aboveground biomass and in the coverage of plants with erect growing habit. However, total belowground biomass remained unaffected by exclusions in both grazing intensities. The relation of root:shoot biomass was lower under moderate grazing intensity, and showed a negative response to deferment treatments at severe grazing intensity.ConclusionLong‐term contrasting grazing intensities applied in South American Campos grasslands determine several changes in above‐ and belowground relationships. Prevalence of prostrate plants under severe grazing is associated with a high partitioning capacity of assimilates toward the subterranean biomass, higher than in the tall species predominant under moderate grazing. The fast recovery of aboveground biomass, the maintenance of the underground biomass and the increase of tall grasses in response to deferments reveals the high resilience capacity of this ecosystem after a long‐term period of overgrazing.

Radiation use efficiency of the herbaceous layer of dry Chaco shrublands and woodlands: Spatial and temporal patterns

AbstractAimsMonitoring rangeland above‐ground net primary production (ANPP) requires information on the variation of radiation use efficiency (RUE). We studied the spatial and temporal variations of RUE of the herbaceous layer (RUEH) in woodlands and shrublands and its association with grazing regime, environmental, and vegetation variables.LocationWe worked in the Arid Chaco (Argentina), an ecotone region between the Chaco and Monte phytogeographic provinces. Woody and herbaceous species coexist in variable proportions in relation to a regional precipitation gradient and local edaphic variations.Methods and resultsOver eight growing seasons, we estimated RUEH at 12 sites, each with two contrasting grazing regimes (moderate and severe). To do this, we related estimated ANPPH from biomass harvests and APARH (absorbed photosynthetically active radiation of the herbaceous layer) calculated from the MODIS normalized difference vegetation index (NDVI). Then we linked the RUEH with precipitation, physiognomy, soil, and landscape data. Most of the spatial variability of RUEH was accounted for by the grazing regime. Moderately grazed areas had a three times greater RUEH than severely grazed ones (0.507 and 0.180 g MJ−1 respectively). They also exhibited different proportions of herbaceous functional types. Physiognomic characteristics of the woody layer explained a significant proportion of the spatial variation of RUEH not explained by grazing regime. RUEH was between 30 and 150% greater in woodlands than in shrublands depending on whether the sites were under moderate or severe grazing respectively.ConclusionGrazing regime and woody physiognomy accounted for most of the variation in herbaceous radiation use efficiency. Our results show that RUEH may be predicted with greater accuracy from a combination of vegetation maps and information on grazing regime. Thus, our models could be incorporated into web platforms that provide ANPPH (forage) monitoring services based on satellite data, in order to improve their estimates in woodlands, shrublands and savanna ecosystems.

Loss of fungal symbionts at the arid limit of the distribution range in a native Patagonian grass—Resource eco‐physiological relations

Abstract Crucial to our understanding of plant ecology is the consideration of the eco‐physiological responses and constraints of plant–fungal symbioses throughout the native distribution range of their host. We examined key eco‐physiological roles of two co‐occurring fungal symbionts [Epichloë endophytes and arbuscular mycorrhizal fungi (AMF)] in the endemic grass Hordeum comosum across a wide bioclimatic gradient and contrasting grazing severity. We sampled H. comosum plants along four humid‐to‐arid transects in Patagonia, Argentina, covering its entire distribution range and determined Epichloë presence, AMF root colonization, nitrogen and phosphorus concentration, intrinsic water‐use efficiency (iWUE, the ratio of photosynthesis to stomatal conductance) and 18O‐enrichment of cellulose in shoots. Root colonization by AMF increased with Epichloë presence. All plants hosted Epichloë in the humid range of the gradient, but symbioses occurrence decreased towards arid sites which also displayed severe grazing symptoms at site level. Symbiosis with Epichloë correlated positively with shoot nitrogen concentration in the centre of the distribution range, and with shoot phosphorus concentration across the entire distribution range. The site‐level relationship of AMF colonization with 18O‐enrichment and iWUE suggested that mycorrhiza boosted stomatal conductance in humid environments but curbed it in arid environments. While the interpretation of interactions and potential causalities from observational studies should be done with caution, this study demonstrates distinct correlations between plant–fungal symbiont associations and key resource parameters (phosphorus, nitrogen and iWUE vs. 18O‐enrichment). Such correlations may suggest particular functional roles for these symbionts in the ecology of their host plant. A free Plain Language Summary can be found within the Supporting Information of this article.

The South African perennial legume Lebeckia ambigua sustainably supports livestock production on infertile soils in rain-fed agro-ecosystems

Lebeckia ambigua is a perennial legume endemic to rangeland South Africa that is adapted to dry (< 400-mm annual rainfall), infertile and acidic (below pH 5.5Ca) soils. Although it is considered that the legume was grazed by mammalian ungulates during its evolution, little is known about its production, nutritive value for grazing animals or optimal management in an agricultural context, in either its centre of the origin or in Australia. Experiments were conducted to test the hypotheses that sheep would graze L. ambigua in autumn without adverse effects on either the animal or the plant. Lebeckia ambigua was established at two sites in Western Australia. In experiment 1, three grazing intensities were imposed by exclusion cages introduced during a single intensive grazing period, to provide differing residual pasture heights. The metabolisable energy and crude protein content of L. ambigua were more than 11.4 MJ/kg dry matter and 21%, respectively for all treatments. Un-grazed L. ambigua produced 1740 kg DM/ha while a single moderate or severe grazing reduced total biomass production by about 50% over 10 weeks post-grazing (P < 0.001). In experiment 2, continuous grazing of L. ambigua for 64 days had no negative impacts on biochemical health indicators of merino wethers. Grazing had no effect on plant density in either experiment indicating that L. ambigua is adapted to substantial grazing pressure in autumn. Lebeckia ambigua could become a valuable plant for sustainably increasing livestock production on infertile soils in rain-fed agro-ecosystems where other legumes fail.

Modern Pollen Assemblages in Typical Agro-Pastoral Ecotone in the Eastern Tibetan Plateau and Its Implications for Anthropogenic Activities

Long-term evolution of human-environment interaction in the Qinghai-Tibetan Plateau (QTP) has been discussed intensively in recent years. The identification of human-related pollen types may help explore the coupled process of climate change, ecological response and anthropogenic activities on the QTP. The aim of this study is to evaluate the impact of anthropogenic activities on surface pollen assemblages and identify pollen indicators associated with grazing and cultivation in typical agro-pastoral ecotone of the eastern QTP. Totally 84 surface samples were analyzed from five vegetational communities in the eastern QTP, which are identifiable based on surface pollen assemblages. Principal component analysis of 29 pollen types and two supplementary variables of human influences were used to assess the impact of anthropogenic activities on surface pollen assemblages in eastern QTP. Severe grazing led to increase of Rosaceae, Ranunculaceae, Fabaceae, Taraxacum-Type, Aster-type and Saussurea-Type in pollen assemblages. Highland agricultural cultivation can be identified by the content and morphological characteristic of Brassicaceae and Cerealia Poaceae pollen. The aerial dispersed distances of Brassicaceae and Cerealia Poaceae pollen are very short, and domesticated herbivores are important factors, which disperse the Cerealia Poaceae pollen into pastures in agro-pastoral ecotone of the QTP. Modern pollen indicators of cultivation and grazing activity will provide precise references for fossil pollen study of prehistoric human activities in the QTP.

Severe grazing pressure on an unpalatable plant, Primula japonica, and its potential chemical compound for grazing defence in a long-term deer grazing habitat

Severe grazing pressure on an unpalatable plant, Primula japonica, and its potential chemical compound for grazing defence in a long-term deer grazing habitat

Population Dynamics in Mixed Canopies Composed of Kikuyu-Grass and Tall Fescue

The current work was based on the central hypothesis that grazing management strategies modulate tillering dynamics of mixed canopies composed of kikuyu-grass (C4 perennial grass) and tall fescue (C3 perennial grass). Among the five grazing management conditions evaluated, three (7, 12, and 17 cm) represented the heights up to which the grasses were kept by mimicking a continuous stocking method throughout the experimental period (from May 2016 to October 2017), and the other two consisted of the heights 12 and 17 cm with a single grazing to 7 cm in mid-autumn. Nitrogen fertilization was applied only during winter–spring. The results showed that under severe grazing (7 cm), kikuyu-grass predominated in the area and that tall fescue population predominated in the moderately lenient (12 cm) and lenient (17 cm) grazed pastures, regardless of an occasional autumn grazing to 7 cm. After two years of evaluation, kikuyu-grass tillers were reduced to 6% of the total population in those pastures managed at the heights of 12 and 17 cm. Grazing management strategies modulate species proportions in a mixed canopy composed of kikuyu and tall fescue. Moreover, our data also suggest that it seems unlikely that there is a stable association of tall fescue and kikuyu-grass when no nitrogen fertilization is applied during summer.

Long-term influence of season of grazing and rainfall on vegetation in the eastern Karoo, South Africa

In the Nama-Karoo, South Africa, rainfall and grazing by livestock are two important drivers of botanical composition. Summer rains are advantageous particularly to the grass layer, whereas the characteristic dwarf shrubs survive better than grasses during drought, and benefit from rainfall during cool months. Heavy grazing by livestock during summer reduces the abundance of grasses, whereas winter grazing increases it. This study examined the influences of season of grazing (summer vs winter) and rainfall on botanical composition at two long-term grazing trials at Grootfontein in the eastern Karoo from 1949 to 1967. Severe summer grazing strongly reduced grass abundance and slightly increased the abundance of both palatable and unpalatable dwarf shrubs. High previous-season rainfall strongly promoted short-lived grasses. Dwarf shrubs and grasses declined in response to a decline in annual rainfall over the study period, except for the creeping grass Tragus koelerioides that increased in abundance, possibly in response to decreased competition and an increase in available gaps in the vegetation layer. Results indicate that grazing systems that exclude repeated summer-only grazing maintain ecological health, and that rainfall is the main driver of composition unless extreme grazing systems are employed.

Comportamiento productivo y calidad de pastos híbridos de Urochloa y estrella pastoreados con bovinos

En las zonas tropicales, las gramíneas son la principal fuente de alimentación para los rumiantes, sin embargo, las condiciones ambientales y el manejo de las praderas inciden directamente en el rendimiento y calidad de las mismas. El objetivo fue estudiar el efecto del rendimiento de los pastos híbridos de Urochloa y pasto Estrella (Cynodon plectostachyus) a diferentes frecuencias e intensidades de pastoreo, con bovinos en el trópico seco. Se evaluaron los pastos Cobra, Mulato II y Cayman (Urochloa) y pasto estrella en frecuencias de corte de 28 y 35 d, e intensidades severo y ligero de 10 y 15 cm, respectivamente, que se distribuyeron de forma aleatoria en un diseño de bloques al azar con arreglo factorial 2 x 2 con tres repeticiones. Se observó efecto del manejo en la acumulación de MS, siendo mayor la acumulación a menor frecuencia e intensidad ligera de pastoreo independientemente del pasto evaluado. El pasto Cobra, Cayman y Mulato II obtuvieron el mayor rendimiento en la frecuencia a 35 d e intensidad ligera con un promedio de 6 679 kg MS ha-1 mientras que, en ese manejo el pasto estrella obtuvo el menor rendimiento con 4 028 kg MS ha-1. La mayor proteína cruda en hoja, en todos los pastos se encontró en el pastoreo severo e intensidad a 28 d con 20, 20, 18 y 10% en Cobra, Mulato II, Cayman y Estrella, respectivamente (p&lt; 0.05). Se concluye que la mayor producción del forraje se obtuvo al cosechar a una intensidad ligera de 15 cm, cada 35 d y el mayor contenido de proteína se alcanzó cuando el forraje se cosecho a intensidad de 10 cm y una frecuencia de 28 d.

Does Ozone Alter the Attractiveness of Japanese White Birch Leaves to the Leaf Beetle Agelastica coerulea via Changes in Biogenic Volatile Organic Compounds (BVOCs): An Examination with the Y-Tube Test

Elevated ground-level ozone (O3) reduced C-based defense chemicals; however, severe grazing damages were found in leaves grown in the low O3 condition of a free air O3-concentration enrichment (O3-FACE) system. To explain this phenomenon, this study investigates the role of BVOCs (biogenic volatile organic compounds) as signaling compounds for insect herbivores. BVOCs act as scents for herbivore insects to locate host plants, while some BVOCs show high reactivity to O3, inducing changes in the composition of BVOCs in atmospheres with elevated O3. To assess the aforementioned phenomenon, profiles of BVOCs emitted from birch (Betula platyphylla var. japonica Hara) leaves were analyzed ex situ, and Y-tube insect preference tests were conducted in vitro to study the insect olfactory response. The assays were conducted in June and August or September, according to the life cycle of the adult alder leaf beetle Agelastica coerulea Baly (Coleoptera: Chrysomelidae). The Y-tube tests revealed that the leaf beetles were attracted to BVOCs, and O3 per se had neither an attractant nor a repellent effect. BVOCs became less attractant when mixed with highly concentrated O3 (&gt;80 ppb). About 20% of the total BVOCs emitted were highly O3-reactive compounds, such as β-ocimene. The results suggest that BVOCs emitted from the birch leaves can be altered by elevated O3, thus potentially reducing the attractiveness of leaves to herbivorous insects searching for food.

Late Quaternary Environmental History of the Horton Plains, Central Sri Lanka

Pollen and mineral magnetic analyses performed on two peat sequences from the elevated area at 2300 m above sea level (a.s.l.)in the central part of Sri Lanka provide an almost continuous succession of data on the vegetation, climate and land-use history of the area since 18,000 BP. An amelioration of the arid Late Pleostoceme environment is indicated by the development of herbaceous and summer forest communities. Xerophytic woodlands predominated at the termination of the Pleistocene, about 13,000 BP. At the very end of the Pleistocene an incerase in precipitatin is indentified by the predominance f a montane rain forest (12,000 - 11,000 BP). The Holecene vegetation changes reflect two significant phases of expansion and diversification in the rain forest, in addition an arid climatic phase from 6000 to 5000 BP was observed, and a short wet phase around 600 BP. The first indications of human impact in the pollen diagram, dated to approximately 14,000 BP, may be the result if severe deforestation, forest clearance and grazing. A pre-farming/pastrol culture existed from 14,000 to 10,000 BP, and changes in both human subsistence strategies and climate, and be recognized around 9000 BP, with the start of the first agricultural land-use (Hordeum sp.and Avena sp.). These activities continued until 6500 BP, whereas only limited agricultural activities and be identified during the period 6500 to 3000 BP. Thereafter, the area was abandoned until small-scale triticum cultivation started around 800 BP, lasting until 200 BP.

Grazing intensity impacts on soil carbon stocks of Western Himalayan Alpine paddocks

Overgrazing in the fragile Himalayan alpine grassland ecosystems is a major cause of substantial loss of soil organic carbon (SOC). The current study was conducted to analyze the effects of grazing on SOC in seven alpine grasslands of Kashmir. SOC was found to be greatly influenced by severe grazing and varied between 43.1 and 120 Mg/ha. Average SOC was found to be 70.5 Mg/ha; it was 90.6 Mg/ha in non-grazed areas (NGAs) and 50.4 Mg/ha in highly grazed areas (HGAs). The highest value of SOC was 139.1 Mg/ha (in NGA) whereas the lowest was 30.3 Mg/ha (HGA). An average of 1632.7 domestic animals reserved by 278 households were recorded per site. The study determined that an increased herd size results in a decreased SOC level. Grazing regime was a countering factor for SOC at all study sites and caused SOC loss of up to 103.3 Mg/ha. Besides intensive grazing, SOC was also found to be influenced by site physiognomy and microclimatic variations. Grazing management is urgently needed in the Western Himalayan alpine grasslands to enhance SOC levels.

Morphophysiology of forage peanut submitted to different intensities of defoliation on grazing with sheep.

The aim was to monitor the growth, the dry mass accumulation curve, and the structural photosynthetic components of forage peanut pasture, in different intensities of defoliation in sheep rotational stocking. Experimental area of Arachis pintoi (cv. Amarillo) had 0.29ha divided equally into 16 paddocks of 180m2 where they were assigned in a randomized block design with four repetitions; intensities of grazing were very lenient, lenient, moderate, and severe, represented by the residue heights of 11.2, 8.4, 5.9, and 3.2cm respectively, from 14cm (pre-grazing height). Weekly evaluations were carried out in the remaining period and in the pre and post-grazing period, with the forage cut to soil level for the determination of the dry mass available, separation of the structural components of pasture, and determination of the area of the leaflets. From this information, the values of leaf area index, leaf to stem ratio, and growth ratios were calculated. With these results, for the growth variables, regression curves were estimated as a function of the rest days offered by each treatment. Despite the lower mass of forage obtained when the leaf area is severely defoliated, the species presents even accumulation of biomass and foliar area. Severe grazing intensity resulted in greater accumulation of leaves, reflected in the leaf area ratio and leaf weight ratio. The lenient intensities provide less rest time between grazing, requiring a greater number of paddocks. Under the research conditions, severe defoliation provides less forage to the animals.

Precipitation and seasonality affect grazing impacts on herbage nutritive values in alpine meadows on the Qinghai-Tibet Plateau

AbstractAimsGrasslands used for animal husbandry are chosen depending on the nutritive values of dominant herbage species. However, the influence of grazing in combination with precipitation and growing season on the nutritive values of dominant species has not been explicated.MethodsTo unveil the influence of the different grazing intensities on the nutritional values, an ecological study was formulated, namely fencing (G0), light grazing (G1), moderate grazing (G2) and high grazing (G3). This ambitious study was undertaken on the nutritive values of the four dominant species of herbage in an alpine meadow on the Qinghai-Tibet Plateau (QTP) during growing season (June–September) for two successive years, namely 2015 (rainy year) and 2016 (droughty year).Important FindingsWe found that (i) the nutritive value of Kobresia capillifolia, Polygonum viviparum and Caragana sinica was noticeably increased by grazing, but negligible effect on Potentilla fruticosa nutritive value was recorded. (ii) During the rainy year (2015), compared with G0, Polygonum viviparum and Potentilla fruticosa displayed 5.4 and 1.5% increases in the crude protein (CP) content and 8.5 and 2.4% increases in vitro true digestibility (IVTD), respectively, while the neutral detergent fibre (NDF) decreased by 13.5 and 0.9%, respectively. During the droughty year (2016), compared with G0, C. sinica and Potentilla fruticosa showed increases in the CP content by 4.3 and 1.3% and increases in the IVTD by 10.7 and 0.4%, respectively, during G3, while the NDF decreased by 6.0 and 1.0%, respectively. (iii) The nutritive values of all species were higher in the years when the rains were good. However, the nutritive values suffered heavily during drought conditions. Besides, the highest and lowest values of nutrition were detected in June and in September, respectively. The inter-seasonal and the inter-annual changes in the nutritional values of species were higher for K. capillifolia and Polygonum viviparum than for Potentilla fruticosa and C. sinica, suggesting that Potentilla fruticosa and C. sinica had higher water-use efficiency. (iv) Grazing clearly reduced the drought tolerance of three species and showed no effects on Potentilla fruticosa. (v) Grazing clearly increased the inter-month variation in the nutritional value of K. capillifolia and Polygonum viviparum but showed no effects on Potentilla fruticosa or C. sinica. Evidently, the grazing effects impacting the nutritional value of the dominant species of herbage exhibited conspicuous inter-annual and seasonal variations with species-specific influences and responses. Our findings are expected to have far-reaching implications enabling the authorities to arrive at strategic decisions and designing of relevant policies for the efficient management of the ecosystems ensuring the speed restoration of the QTP under severe grazing and extreme climatic circumstances.

Terrestrial Gastropod Grazing on Macrolichens in a Northern Broadleaf–Conifer Forest

Herbivory by terrestrial gastropods, particularly Arion spp. (a slug), can alter epiphytic lichen communities; however, little is known about this interaction in forests of North America. We used 3 lines of evidence to explore this interaction: field grazing assessments on lichen thalli, a 10-y re-measure of gastropod abundance, and gastropod feeding trials in a montane forest at Hubbard Brook Experimental Forest (HBEF) in northern New Hampshire. Grazing damage by terrestrial gastropods was widespread, though few sites had severe grazing. Grazing damage was significantly higher on flatter terrain and on broadleaf trees. Slug densities were significantly lower in 2016 than in earlier surveys (1997–2006) on 4 of 6 plots. In feeding trials, 2 common lichens, Hypogymnia physodes and Platismatia glauca, were grazed more heavily by both native and non-native slugs than other lichen species. However, the Succineidae (amber) snails preferred Lobaria pulmonaria, a lichen that has been declining at HBEF in the last decade. Overall, lichen communities in the HBEF were moderately impacted by terrestrial gastropod grazing, but potential effects of the non-native slugs at higher elevations and impacts on lichen health of widespread, moderate grazing deserve further study.

Interactions of grazing and rainfall on vegetation at Grootfontein in the eastern Karoo

Rainfall and grazing are primary drivers of vegetation composition in the Nama-Karoo. Increased rainfall increases grassiness, to where Nama-Karoo transitions to grassland. Severe grazing treatments (e.g. continuous or summer-only) increase abundance of grazing-tolerant dwarf shrubs and annual grasses, and decrease perennial grasses. Grootfontein, which is ecotonal between the Nama-Karoo and Grassland Biomes, houses long-term grazing trials. The area has experienced higher-than-average rainfall in recent decades. Plant basal cover data from the 1960s and 2010s allow several hypotheses to be addressed: (1) historical severe grazing (until 1985) will limit subsequent grassiness (grazing legacy effect); (2) severe grazing will preclude increases in grassiness, independent of rainfall (herbivore trap effect); and (3) historically leniently-grazed sites will transition to grassland with increased rainfall (biome shift effect). Rainfall was lower from 1957–1966 (350 mm) than from 2003–2012 (490 mm). The grazing legacy effect was supported based on the abundance of Aristida diffusa, despite all sites becoming much grassier. The herbivore trap effect was not supported. The biome shift effect was supported in that shifts to grassland sometimes occurred. Results suggest that increasing rainfall has prompted a shift to much increased grassiness and decreased abundance of dwarf shrubs, and that grazing had a smaller secondary effect.

Testing rangeland health theory in the Northern Great Plains

Abstract Correctly assessing whether rangeland ecosystem services are stable, improving, or degrading is of global importance. Soil aggregate stability (SAS) is widely used to infer rangeland health, partly because high SAS is thought to reduce run‐off by increasing infiltration. We studied the sensitivity of SAS to grazing and other disturbances, the effects of SAS on infiltration, and the utility of alternative indicators of infiltration in the Northern Great Plains. To test grazing effects on SAS, we compared SAS between paired areas that were lightly to moderately grazed or excluded from grazing for 6 years. Additionally, we compared SAS between grazed and not‐grazed plots of a 2‐year controlled grazing experiment with moderate and severe grazing. We also applied herbicide, mowing, and fungicide treatments to test SAS responses to disturbances more generally, as well as effects of SAS and other factors on infiltration. To more generally test for a SAS–infiltration relationship, we performed a meta‐analysis of our data combined with other data from the region. Grazing often reduced stability of small macroaggregates (0.25–1 mm) in the controlled grazing experiment but not the paired grazing area experiment. Grazing had no detectible effect on SAS of larger macroaggregates (1–2 mm). Herbicide tended to reduce SAS, and mowing sometimes increased SAS. Infiltration exhibited high plot‐to‐plot variation and was not significantly affected by treatments. Variation in infiltration was best explained by plant community composition variables and was not explained by either SAS or other soil properties. Our meta‐analysis revealed no general SAS–infiltration relationship. Synthesis and applications. Our findings counter prevailing expectations that soil aggregate stability (SAS) is consistently sensitive to rangeland disturbance(s) and a leading indicator of soil water transport. Plant community composition properties were better predictors of infiltration. Our findings support the theory that excessive grazing increases the prevalence of a grazing tolerant species, which was associated with low levels of infiltration irrespective of SAS.