Growth Of Scots Pine Research Articles

Assessing the ecological complexity and uncertainty of predicting forest ecosystem services under climate change

Climate change affects Central European forest ecosystems in different ways and, consequently, these changes result in different feedbacks on the provision of forest ecosystem services. Regarding the complexity and the variability in climate-forest interactions outcome, forest decision makers necessitate reliable information about changes in the forest ecosystem services for planning and adaptation purposes. However, forest productivity predictions incorporate multiple levels of uncertainty that have to be regarded to ensure building realistic expectations in forest decision-making. Besides the chosen forest simulation model, uncertainties come from the climate change data represented by a set of representative concentration pathways (RCP), within the underlying ensemble of global circulation and regional climate models (GCM-RCM), and further in the treatment of the CO2-fertilization effect. We considered the mentioned uncertainties in a framework on simulating forest growth and water services for two forest sites, a Sessile oak and a Scots pine stand in Rhineland-Palatine, Germany. The framework revealed a high variability in future forest ecosystem services. Particularly, the variability among the selected GCM-RCM models within the same Representative Concentration Pathway (RCP) was higher than the variability among different RCPs (RCP2.6 and RCP8.5 representing the low and high CO2-emission scenarios, respectively). Sessile oak productivity increased under all scenarios, whereas Scots pine growth declined in the lower end of the RCP8.5 scenario. Water services remained mostly stable at both sites. Moreover, we applied a panel data model to estimate what climate indices caused changes in the forest ecosystem services. We found that Scots pine is more sensitive to a multitude of climate indices, such as temperature changes and Sessile oak showed strong response to the CO2-fertilization. We propose applying this framework to evaluate forest management options under climate change.

Repeated Lophodermium needle cast has a cumulative impact on the growth of Scots pine for up to 12 years

Repeated Lophodermium needle cast has a cumulative impact on the growth of Scots pine for up to 12 years

Tree growth history, legacy of large-scale forest management and climatic variability in North Finland

Tree growth history was investigated in North Finland to study the recently observed reduction in Scots pine growth. Tree-ring records from the National Forest Inventory (NFI) were compared to changes that have occurred in forest management activities and to variations in the North Atlantic Oscillation (NAO), as similar comparisons have not been previously made with NFI data. A phase of increased growth was dated to the early 1970s when the forest management activities were intensified on a large scale. More recently, growth has notably declined since the early 2000s which coincides with reduced activity to clean the ditches to ensure they are draining the substrate adequately. The growth reduction appeared greater for trees from stands with higher basal area. Trees representing sites where the first cleaning of the stand (thinning) had been carried out did not show reduced growth. NAO indices correlated positively with growth, especially during the cold season. An interpretation on the value of snow conditions in a dynamic interaction with management history/stand density is put forth to explain the growth decline observed in NFI data over the past two decades.

The effect of climate change on the radial growth of Pinus sylvestris l. and Quercus robur l. in the stands of Kharkiv green zone

Purpose. To identify the characteristics of the reaction of the radial growth of Scots pine (Pinus Sylvestris L.) and Scots oak (Quercus robur L.) to climate change in the forest-steppe zone of Ukraine. Methods. Dendrochronological, dendroclimatological, and statistical methods are applied. COFECHA and ARSTAN programs were used. Results. The study was conducted in 100-year-old stands of English oak and Scots pine in the Southern Forestry (Left-bank Forest-steppe). The reaction of the radial growth of trees to climate changes was compared for 1960-1987 and 1988-2016. For the dendroclimatic analysis, the de Martonne hydrothermal indices, the Selyaninov hydrothermal coefficient, the forest aridity index, and the O1 hydrothermal coefficient were used. Relative humidity indicators, which are derived from temperature and precipitation, were also used. It was found that in the second period, the sensitivity of the reaction of the radial growth of trees to climate variations increased, which indicates a certain weakening of the stands due to the increase in temperature. Dendroclimatic analysis of Pinus sylvestris L. and Quercus robur L. showed that when comparing the response of the radial growth of trees to climate variations in 1960-1987 and 1988-2016, the condition of both pine and oak weakened, as evidenced by an increase in significant coefficients correlations between growth indices and climatic factors. Conclusions. Pine was found to be more sensitive to climate change compared to oak. At the same time, these species of these trees at this stage were able to adapt to climate change, as evidenced by tree-ring chronologies, which show stable growth during 2010-2016.

Auxins differentially affect growth in Scots pine and Norway spruce in spring and autumn

Trees adapt to changing environmental conditions largely by regulating growth processes, and ongoing climate aridization puts pressure on growth processes during spring and summer. However, trees can be limited in their ability to utilize favourable early-season and late-season conditions for growth, and many of these limitations can be due to shortages in auxin transport and/or signalling. We studied whether leaf spraying with auxins in spring and autumn can aid in the growth of Pinus sylvestris and Picea abies saplings. In spring, spruce demonstrated prominent positive effects of auxin supplementation on the growth of current-year shoot and xylem growth, with increased xylem cell formation, whereas a limited effect was observed in pine. Autumnal auxin treatment had prominent positive effects on the growth of the root system and xylem area in pine, activating xylem cell enlargement and increasing carbon deposition in cell walls. In spruce, the effects of auxin spraying on xylem growth were less prominent, and the effects on root growth were absent compared to those in pine. The metabolome analysis of pine organs demonstrated that the activation of growth processes in the xylem and roots was accompanied by a decrease in nonstructural carbohydrates in the cambial zone but an increase in roots, indicating that auxin influenced the link between growth and carbon balance in an organ-specific manner. The major amino acid content was not influenced by auxin treatment. The observed results indicate that changes in the auxin balance during the autumnal period can activate growth processes to increase plant adaptation to changing climate conditions.

Stand structural effects on growth distribution and growth efficiency in Scots pine and Mediterranean pine in Spain

AbstractUnderstanding the growth efficiency of individual trees, or growth per unit of resource utilization, can inform silvicultural management strategies to maximize tree and stand growth. Stand structure—the size and spatial distributions of trees within the stand—strongly influences water, light, and nutrient availability, as well as the resource-use efficiency of each tree. Key silvicultural tools for stand management include manipulating tree density, size distribution, and arrangement by controlling natural regeneration, artificial seeding, planting seedlings, and/or subsequent thinning of established trees. We analyzed two sets of plots from even-aged stands of common coniferous species in central Spain, 106 pure Scots pine (Pinus sylvestris) and 92 pure Mediterranean pine (Pinus pinaster), to examine the dynamics of the dominance hypothesis, the relationship between stand structure and growth, and the relationship between structure and growth efficiency. Our main findings revealed a negative impact of size-class uniformity on stand growth in both Scots pine and Mediterranean pine, while the positive effect of tree size on growth efficiency was supported for Mediterranean pine stands but uncertain for Scots pine. At the operational level, our results highlight how thinning intensity is more important than the thinning method in Mediterranean pinewoods and how thinning can benefit the provision of multiple ecosystem services. We also recommend integrating dominance effects on growth into individual tree modelling.

Effects of Planting Position, Seedling Size, and Organic Nitrogen Fertilization on the Establishment of Scots Pine (Pinus sylvestris L.) and Norway Spruce (Picea abies (L.) Karst) Seedlings

The forest regeneration phase in Sweden commonly involves mechanical soil preparation followed by the planting of Scots pine (Pinus sylvestris L.) or Norway spruce (Picea abies (L.) Karst) seedlings. The prepared soil offers planting positions with different properties, including reduced damage by pine weevils (Hylobius abietis L.). Nitrogen fertilization can be applied at the time of planting to aid establishment of the seedlings. In this study, we compared the effects of different planting positions, organic nitrogen fertilization, and different seedling sizes on the early survival and growth of Scots pine and Norway spruce seedlings. The main planting positions were capped mound, hinge, and mineral soil. Seedlings planted close to organic material were categorized as being in “low-quality positions”, since proximity to organic material increases pine weevil attraction. Higher mortality rates related to pine weevil damage were recorded for the seedlings planted in the low-quality positions, regardless of seedling size or N fertilization. Pine weevil attack rates increased with increasing seedling size. Growth was, in general, lowest in the mineral soil positions. The effect of organic N fertilization on growth was positive for the spruce regardless of the planting position or seedling size, while it depended on the planting position and seedling size for the pine, indicating that the effects of organic N fertilization depend on the seedling species, seedling size, and planting position.

Формирование морфологии деревьев культивируемых сосновых древостоев

The aim of this study is to assess the qualitative characteristics of the crowns, stems and growth of Scots pine (Pinus sylvestris L.) trees in cultivated stands of a dry forest on gently sloping hills thinned in a regular-uniform manner, as well as to develop proposals for production based on the data obtained. The stands under study grow in the southwestern part of the ribbon forests of the Altai Territory in the territory of the Klyuchevskoye forestry. The thinning was carried out in 2016 as part of the planned forest maintenance activities. The field stage of the research was completed in 2022. The sample plots laid had a rectangular shape and a square of at least 0.25 ha. The assessment of the qualitative characteristics of the trees has been carried out in accordance with the classification proposed by G.A. Chibisov et al. The significance of differences (or their absence) in the tree distribution by quality categories in the plots has been established using the χ2 test for comparing two distributions. The article presents the taxation characteristics of the main forest element of the stands under study, the quality indices for the assessed tree indicators, and the graphs of tree distribution by diameter and quality classes. It has been found that most of the trees have poor crowns (27–45 %) and growth (14–41 %). Such indicators are a consequence of an excessively high density of the stands and the lack of timely thinning. Regular-uniform thinning of moderate and moderately high intensity has contributed to an increase in the quality class of the trees. The impact of high and very high intensity thinning on the stands has been negative. Based on the conducted research, it is recommended to carry out thinning at an earlier age of the stand, abandon the regular-uniform thinning in favor of selective thinning using the grassroots method (if possible) and carry out thinning along the largest exploitable diameter of 10 cm in the studied forest stands and similar ones. The study can be continued by establishing the optimal exploitable diameter in younger plantations and in other types of forests, as well as determining the intervals of optimal density of pine stands of different ages and different types of forest. The results obtained can be useful in designing economic activities in the study area.

Elevation-dependent tree growth response to climate in a natural Scots pine/downy birch forest in northern Sweden.

Forests dominate the landscape at high latitudes in the boreal regions and contribute significantly to the global carbon stock. Large areas are protected and provide possibilities to analyze natural forest dynamics including resilience to climate change. In Fennoscandia, Scots pine (Pinus sylvestris L.) and downy birch (Betula pubescens Ehrh.) often coexist in natural forests close to the limits of their ecological ranges. Tree growth in these forests is generally thought to be limited by temperature, and changes in growth trends can therefore serve as early indicators of the impact of global warming on natural ecosystems. We sampled 592 Scots pine and downy birch trees along two elevational gradients spanning the transition from the forest zone to the coniferous treeline in Tjeggelvas nature reserve, northern Sweden. Based on the tree-ring data, we compared annual basal area increment (BAI) trends from 1902 to 2021, analyzed the ring-width indices (RWI) in relation to local climate data, and investigated trends in climate-growth relationships. We found that the mean annual growth of both species was higher in more recent years than at the beginning of the 20th century. The RWI were positively correlated with summer temperatures, however, we found a much stronger relationship for Scots pine than downy birch. We noticed a decrease in the importance of summer temperature for Scots pine growth, whereas the importance of late spring temperatures increased over the 120-year-long study period. Due to strongly positive BAI trends combined with a decrease in temperature sensitivity, the overall conclusion of our study is that the influence of increasing temperatures is still positive and outweighs the negative impacts of climate change on Scots pine growth in natural forests in northern Sweden, particularly at higher elevations. Natural forests are important natural experiments that contrast the managed forests and are key to understanding the latter.

Increasing Winter Temperatures Stimulate Scots Pine Growth in the North German Lowlands Despite Stationary Sensitivity to Summer Drought

More than half of the forest area of the North German Lowlands is stocked with Scots pine-dominated forests, mostly plantations. Climate change suggests a declining suitability of Europe’s temperate zone for conifer plantations, but only a few studies have examined the long-term growth trends of Scots pine in relation to environmental and site factors in this region. We studied the radial growth patterns of Scots pine over the last 60 years at ten sites along a precipitation gradient (830–530 mm mean annual precipitation) from an oceanic to a subcontinental climate, analyzing the spatial and temporal variability of the climate sensitivity of growth to identify the main climatic factors influencing pine growth across this gradient, which covers a large part of the species’ tolerated precipitation range. Annual radial increment was sensitive to late-winter temperatures (February, March) and summer drought and heat (June–August), with sensitivity increasing from the oceanic to the drier continental sites. Warmer late-winter periods apparently have stimulated growth during the last decades, while the sensitivity to summer-drought has remained fairly stable. Until recently, the negative impact of warming summers on growth has been compensated by the positive effect of late-winter warming, resulting in stable (or increasing) growth trends. However, our comparison of the climate sensitivity across sites suggests that the drought effect compensation through winter warming will in future be limited by increasing drought exposure. Thus, future productivity declines are likely in the northern German lowlands despite warming winters, discouraging large-scale pine plantations in the face of climate warming.

100 years of change in an old-growth Scots pine forest in Hamra National Park: insights from permanent plots established in central Sweden in 1922

ABSTRACT Hamra National Park in the province of Dalarna in central Sweden was established in 1909 to protect a unique unlogged old-growth forest. In 1922, Henrik Hesselman set up two permanent plots in Hamra to be able to follow the development of the forest over time. In 2022, we re-inventoried the Scots pine-dominated plot and found that (1) the number of trees in the plot almost doubled from 593 trees in 1922 to 1013 trees in 2022, (2) Norway spruces are starting to take over the initially Scots pine-dominated plot, (3) the basal area of dead trees has increased by 286% and (4) the variability in annual Scots pine growth has increased over the studied time period. Given this strong deviation from the original state of the forest, we suggest restoring part of the forest to its early-1900s state while allowing the permanent plot to evolve naturally.

Comparing the effects of ground cultivation and protection against browsing upon the natural regeneration of Scots pine and birch in a Caledonian pinewood

Abstract The Caledonian pinewoods of northern Scotland represent the remnants of once extensive forests found at the western edge of the natural range of Scots pine (Pinus sylvestris L.). The surviving pinewoods are of international significance and are of high conservation and cultural value. However, for many decades, there has been concern about their long-term future because of a lack of regenerating seedlings and young trees. This problem is thought to be driven by high deer populations resulting in intensive browsing and severe damage to young trees, as well as a lack of favourable micro-sites for seedling germination. An experiment was established on the Mar Lodge estate in 2007–08 adjacent to native pinewood stands to test the effects of protection and site cultivation upon seedling numbers and growth. The experiment was located within a regeneration zone where intensive deer control had reduced numbers to <5 animals km−2. Ten years after initiation, occurrence and numbers of both Scots pine and birch seedlings were substantially increased in cultivated treatments compared with uncultivated controls. There were no effects of protection upon seedling numbers. Protection improved height growth of Scots pine, but not of birch. Cultivation had no effect upon the growth of pine seedlings but did influence the height of birch seedlings due to the rapid growth of some pre-existing seedlings in the uncultivated control. Vegetation cover had increased from ~60%to at least 80% cover 6 years after cultivation. The results suggest that site cultivation may be desirable to boost seedling germination within a Caledonian pinewood, but sustained measures to control browsing pressure are a prerequisite for satisfactory establishment of these seedlings.

Natural regeneration and early development of Scots pine seedlings after gap cutting in northern Finland

ABSTRACT In the northern boreal region, tree growth, timber yields and economic returns are low, meaning low regeneration costs are the basis of profitable forestry. If natural regeneration is successful, it may be favoured over expensive planting or seeding. We studied the regeneration success and early growth of Scots pine in gaps in terms of seedling density and height 10 years after gap cuttings in central Finnish Lapland. Three gap sizes (diameters of 20, 40, and 80 m) were studied on patch scarified xeric and sub-xeric sites in six random blocks and a total of 18 replicates of each. The number of pine seedlings was high across the gap sizes. The proportion of empty regeneration plots (size 5 m2) was ∼ 2%. Site preparation substantially increased the number of seedlings. The growth of seedlings was faster in larger gaps, but a high proportion of exposed mineral soil decreased it. The results suggest that all studied gap sizes regenerated naturally well, and that soil scarification exposing 10–20% of the surface area or even less can be enough to achieve regeneration goals. Gaps of a diameter of 40 m or more are required to achieve an optimal balance between seedling density and growth.

Climate Sensitivity and Drought Legacy of Tree Growth in Plantation Forests in Northeast China Are Species- and Age-Dependent

The occurrence, frequency, and severity of drought are accelerating due to global warming. Understanding the vulnerability of plantation forests to climate change, particularly to drought events, is critical to revealing the underlying mechanisms of tree resilience, recovery, and acclimation, which are important for plantation management. How the stand age affects the climate sensitivity of tree growth, as well as the direction, magnitude, and duration of the drought legacy, in plantation forests in northeast China is still unclear. In this study, we used MODIS-derived NDVI time series with gridded climate data from 2000 to 2020 to fill this knowledge gap. The selected plantation forests were dominated by four coniferous species: Korean pine (Pinus koraiensis), Scots pine (Pinus sylvestris), Japanese larch (Larix kaempferi), and Dahurian larch (Larix gmelinii). The results show that the climate sensitivity of tree growth differed among species and age groups. The growth of Korean pine and Scots pine was mostly dependent upon precipitation, while the growth of Japanese larch and Dahurian larch was determined primarily by temperature. Old Japanese larch (21–40 years) and Dahurian larch trees (31–60 years) were more sensitive to temperature and precipitation than young conspecifics, whereas old Korean pine (41–60 years) and Scots pine (31–60 years) were less sensitive to precipitation and temperature than young conspecifics. Furthermore, the legacy of drought lasted one year for Korean pine, Japanese larch, and Dahurian larch and over three years for Scots pine. Old trees were more severely affected by drought, particularly Scots pine and Dahurian larch. The findings of the study can help improve plantation forest management for better adaptation to future climate change.

Effects of Climate Change on Scots Pine (Pinus sylvestris L.) Growth across Europe: Decrease of Tree-Ring Fluctuation and Amplification of Climate Stress

From an economic perspective, Scots pine (Pinus sylvestris L.) is one of Europe’s most important tree species. It is characterized by its wide ecological adaptability across its natural range. This research aimed to evaluate the forest structure, productivity and especially radial growth of heterogenous pine stands in 16 research plots in the Czech Republic, Poland, Spain and Great Britain. The study assessed the tree-ring formation and its relationship to climate change for each country, using 163 dendrochronological samples. The stand volume of mature pine forest ranged between 91 and 510 m3 ha−1, and the carbon sequestration in the tree biomass was 40–210 t ha−1. The stands had a prevailing random distribution of trees, with a high vertical structure close to selection forests (forest stands with typical very diverse height, diameter and age structure). Spectral analyses showed a substantial decrease in fluctuations in the tree-ring index and a loss in natural growth cyclicity in the last thirty years. The results also evinced that mean air temperature was the most important factor influencing the radial growth compared to precipitation totals. Pine thrives in precipitation-stable locations, as shown by the results from Great Britain. The conclusions of this study confirm the fundamental effect of ongoing global climate change on the dynamics and growth of pine forests in Europe.

Major tree species of Central European forests differ in their proportion of positive, negative, and nonstationary growth trends

AbstractTemperate forests are undergoing significant transformations due to the influence of climate change, including varying responses of different tree species to increasing temperature and drought severity. To comprehensively understand the full range of growth responses, representative datasets spanning extensive site and climatic gradients are essential. This study utilizes tree‐ring data from 550 sites from the temperate forests of Czechia to assess growth trends of six dominant Central European tree species (European beech, Norway spruce, Scots pine, silver fir, sessile and pedunculate oak) over 1990–2014. By modeling mean growth series for each species and site, and employing principal component analysis, we identified the predominant growth trends. Over the study period, linear growth trends were evident across most sites (56% increasing, 32% decreasing, and 10% neutral). The proportion of sites with stationary positive trends increased from low toward high elevations, whereas the opposite was true for the stationary negative trends. Notably, within the middle range of their distribution (between 500 and 700 m a.s.l.), Norway spruce and European beech exhibited a mix of positive and negative growth trends. While Scots pine growth trends showed no clear elevation‐based pattern, silver fir and oaks displayed consistent positive growth trends regardless of site elevation, indicating resilience to the ongoing warming. We demonstrate divergent growth trajectories across space and among species. These findings are particularly important as recent warming has triggered a gradual shift in the elevation range of optimal growth conditions for most tree species and has also led to a decoupling of growth trends between lowlands and mountain areas. As a result, further future shifts in the elevation range and changes in species diversity of European temperate forests can be expected.

Fertilisation with potato starch wastewater effect on the growth of Scots pine (Pinus sylvestris L.) forest in Poland

Fertilisation is often used to increase plant productivity in agriculture but has also been used in forestry. In our study, Scots pine forest growing in a nitrogen-poor environment was fertilised with NPK post-production wastewater from a potato starch factory. Our research aimed to investigate the dependence of tree growth on different NPK concentrations. Cell characteristics such as cell wall thickness (CWT), lumen diameter (LD) and tree-ring features such as ring width (RW), total number of cells in annual growth (nTotal), earlywood (EW) and latewood (LW) were investigated. Twenty-six years of regular fertilisation of the forest with different doses of wastewater rich in NPK elements have affected the anatomical structure of Scots pine trees. It is presumed that the reduction in CWT and LD on the fertilised site was due to deficiencies in plant water conductivity, which may have occurred due to physiological drought. The influence of nitrogen on unfertilised site from the wastewater area could contribute to the CWT thickening. The results confirm that the use of NPK in excessive doses is detrimental to trees' conductive system.

Global warming leads to growth increase in Pinus sylvestris in the Kazakh steppe

Forest community structure and function in climate-sensitive regions exhibit uncertain changes under the influence of global warming. Especially in northern Central Asia, the response of pine growth to climate change remains unclear. Here, we identified changes in growth trends and climate response in a total of 646 trees for the abundant species Scots pine (Pinus sylvestris) at 30 sites in the Kazakh steppe. We used basal area increment, regional curve standardization and negative exponential curve detrended chronologies to comprehensively analyze trends in pine growth, and linear mixed effects and random forest models were calculated to explore growth-climate relationships. An increase in the overall growth of Scots pine under the influence of global warming, expressed in this region by increasing temperatures and precipitation, was observed during the period 1950–2018. Spatial differences in climate response between the sub-regions were identified showing that pine growth is more influenced by summer precipitation in the eastern compared to the western and central Kazakh steppe. Warm spring temperatures and more precipitation are important common climatic factors affecting pine growth in northern Central Asia. Moreover, increased drought intensity and longer drought duration negatively impact pine growth in this region. The Pacific Decadal Oscillation and North Atlantic Oscillation were identified to be the main atmospheric circulation patterns that influence the spatial differentiation in pine growth in this steppe. Albeit pine growth in northern Central Asia shows an increasing trend currently but also in future climate scenarios, the risks and challenges posed by drought should not be underestimated.

The effects of arginine phosphate (ArGrow® Granulat) on growth of Scots pine and Norway spruce seedlings planted in varying soil layer structures simulating site preparation

The effects of arginine phosphate (ArGrow® Granulat) on growth of Scots pine and Norway spruce seedlings planted in varying soil layer structures simulating site preparation

Short-term effect of thinning on inter- and intra-annual radial increment in Mediterranean Scots pine-oak mixed forests

Thinning treatments along with the establishment of mixed forest stands have been put forward as possible adaptation strategies to cope with climate change, although the effectiveness of combining these two measures has scarcely been studied and may vary depending on stand conditions and the thinning regime employed. The aim of this study was to better understand the effect of commercial thinning and of the different growth behavior of two coexisting species on their inter- and intra-annual cumulative radial increment patterns. For this purpose, we studied radial increment in a Scots pine-Pyrenean oak (Pinus sylvestris L.-Quercus pyrenaica Willd.) Mediterranean mixed forest in north-west Spain over two climatically contrasting years (2016–2017). The data came from a thinning trial consisting of a randomized latin square design with a control and two commercial thinning treatments from below; one moderate and the other heavy, removing 25% and 50 % of initial basal area, respectively, of both species. The radial increment was analyzed based on bi-weekly readings from band dendrometers installed on 90 oak and pine trees. A non-linear mixed model based on double-Richards curve was fitted to explore the differences between thinning treatments and species response in the intra-annual cumulative radial increment patterns. Inter-annual basal area increments for each species at stand level were quantified by aggregating the tree estimates obtained from the model fitted in the first step. Tree and stand level growth were greater in Scots pine, which also showed a greater growth response to early spring droughts than the Pyrenean oak. Heavy thinning increased radial increment in trees of both species at the expense of decreased total stand basal area. At species level, basal area growth in Scots pine decreased through thinning, whereas for Pyrenean oak, the heavy thinning intensity resulted in the same basal area growth as the control. Thus, heavy thinning induced a trade-off between total stand growth and tree-level response to climatic conditions for Scots pine but with no loss in productivity in the case of the Pyrenean oak. Hence, heavy thinning may be an appropriate measure to attain productive stability of the oak coppice in the studied mixed forest as well as to adapt tree growth to future droughts associated with climate change.