Installing light rail transit across a floating roadway bridge presents many unique challenges, foremost of which is how to design the multidimensional moving joints at both ends of transition spans between the fixed and floating structures. A technical solution to this technical challenge has been proposed, analyzed, modeled, designed, and prototype tested in the context of the East Link Extension, a light rail transit project being carried out by Central Puget Sound Regional Transit Authority (Sound Transit). Described are the planning, execution, and results of a full-scale prototype testing program of the proposed Curved Element Supported Rail (CESuRa) Track Bridge System, based on the relationship of curved rail supports in two independent planes that adjust in response to the movements of the Interstate 90 (I-90) Bridge. The prototype CESuRa track bridges performed as expected; the information gathered and lessons learned will greatly benefit the final design, the production track-bridge fabrication, and the installation process. Also discussed is the anticipated performance of these track bridges on the I-90 structure in conjunction with the proposed direct fixation track system, fixed rail anchors, and sliding rail expansion joints to be strategically located across the 1.1-mi I-90 Bridge. Built in 1989, the I-90 Bridge is the fifth-longest floating bridge in the world and carries three westbound and two reversible lanes of traffic between Seattle and Mercer Island, Washington. The bridge will be reconfigured to carry four westbound lanes, and the reversible lanes will be converted to light rail transit.