Rhaetian Research Articles

FIRST EVIDENCE OF PALAEO-TETHYAN UPPER TRIASSIC CALCAREOUS NANNOFOSSILS IN NORTH DOBROGEAN OROGEN (ROMANIA)

This study presents the analysis of Upper Triassic sediments from five locations in North Dobrogea(Romania) and the Black Sea. Microfacies analyses on thin sections from the Frecăţei log reveal a shallowing trend and likely an increase in energy upward into the section. The oldest part of the log is characterised by deposition of mudstones transitioning to wackestones, with an increase of bivalves. Similar wackestones are observed in the Izvoarele and Rândunica logs, i.e. microfacies with abundant bivalves, some foraminifera and echinoderms. The offshore boreholes 816 and 817 Lebăda Vest (core CM 9 and CM 31) seem to have been deposited in a basinal or distal marine shelf environment indicated by the presence of mudstones with rare bioclasts. In contrast, sample CM 10 from borehole 816 LV is a micritised grainstone suggesting a deposition in a shallower, higher energy environment.Scanning electron microscope observations reveal a moderate diagenetic alteration in all studied samples, mainly due to dissolution. Two calcareous nannofossil species: Prinsiosphaera triassica triassica and Eoconusphaera zlambachensis were commonly observed in sample F of the Frecăţei log, allowing for assignment of the sediments to the Rhaetian age. Two coccoliths were also observed in sample F of Frecăţei log and sample CM 9 of the borehole 816 LV indicating the presence of coccolithophorids within the study region. This discovery constitutes the first confirmed record of well-preserved, determinable Upper Triassic calcareous nannofossils in the Palaeo-Tethys Ocean.

Biosediment assemblages reveal disrupted silica cycling and redox conditions throughout the Rhaetian Stage: Evidence for a precursor event to the end-Triassic mass extinction

The Norian–Rhaetian boundary (Late Triassic) represents an important precursor extinction event to the end-Triassic mass extinction, but the biotic and geochemical shifts are not well-understood due to poor stratigraphic constraints. Here we examine the microfossil record for metazoans and protists on a Panthalassan carbonate ramp (Gabbs Formation, Nevada, U.S.A.) during the late Norian to mid-Rhaetian, and correlate changes in these assemblages with macrofossil shifts and geochemical data (strontium and carbon isotopes). In the latest Norian, demosponge spicules represent a small proportion of shallow marine biosediments. Demosponges are joined in the earliest Rhaetian by increasingly abundant hypersilicified sponge spicules and silica-limited hexactinellid sponge spicules synchronous with a negative strontium isotope excursion indicating increased hydrothermal or volcanic activity. Common carbonate microfossils such as echinoderm ossicles and ostracods are typically silicified in these deposits as well, suggesting increased silicic porewater. The source for increased dissolved silica in shallow marine systems is suggested to be hydrothermal vent degassing, likely associated with increased tectonic rifting activity. Mid-Rhaetian microfossil assemblages exhibit evidence for intermitted anoxia in reducing conditions, supporting a scenario in which environmental stress was a prolonged feature of much of the Rhaetian Stage, rather than a short-term event in the terminal Rhaetian. While there is no marine sedimentary record of volcanism recognized for this interval, biosedimentary assemblages may serve as proxies for geochemical conditions associated with rifting.

A possible Norian–Rhaetian boundary in the high-latitude continental Junggar Basin indicated by the δ 13 C org record

Abstract In contrast to the Triassic–Jurassic boundary, there is no consensus on the definition and age of the Norian–Rhaetian boundary (NRB), which hampers the global correlation of Rhaetian strata and thus interpretation of the patterns and driving mechanism of the end-Triassic mass extinction. Recent works show that a significant negative carbon isotope excursion (N-CIE) occurs in the NRB interval ( c. 205.7 Ma), which probably provides a physical marker for the NRB. However, no such records have so far been reported from high-latitude continental deposits. Here we present high-resolution bulk organic carbon isotope data from the Upper Triassic continuous lacustrine-to-fluvial deposits in the Haojiagou section of the high-latitude continental Junggar Basin in northwestern China. A significant N-CIE is shown at an age of c. 205 Ma, which is most probably the expression of the NRB N-CIE in the Junggar Basin owing to their very close occurrence times. The NRB is thus probably along the N-CIE in the Haojiagou section, and a short Rhaetian stage is then supported. The Late Triassic climate in the high-latitude Junggar Basin was consistent with the global changes in climate, characterized by a long-sustained cooler and humid climate interrupted by the late Norian and latest Rhaetian warm events.

Upper Carnian to Rhaetian (Upper Triassic) conodont and carbon-isotope stratigraphy of the Csővár borehole succession, Hungary

A total of 125 samples from core from the Csővár borehole (north-central Hungary) were investigated for conodont bio- and carbon-isotope chemostratigraphy. This 620 m thick continuous pelagic carbonate succession comprises strata from the upper Carnian to upper Rhaetian, including both, the Carnian-Norian and the Norian-Rhaetian stage boundaries. The mass occurrence of the Metapolygnathus communisti conodont group followed by the occurrence of Primatella gulloae mark the base of the Norian in the lowermost part of the section, which allows good correlation with the Pizzo Mondello section (Sicily, Italy). The lower Norian conodonts of the core belong predominantly to the genus Ancyrogondolella. Subdivision of the middle Norian interval is hampered by the lack of fully adult specimens. The upper Norian shows a greater diversity than the rest of the Norian, and the rich assemblages give an insight into the evolutionary transition from Misikella hernsteini to Mi. posthernsteini (sensu stricto). The first occurrence of Mi. ultima in the core changes the previous views of the thickness of the upper Rhaetian in the Csővár area and emphasizes the necessity of re-calibration of the isotope stratigraphic results of previous studies. The carbon-isotope results show distinct fluctuations and trends. Some of them are similar to literature data, others are different. It can be stated from the new data that a ∼ 2 ‰ long-term δ13C background decline throughout the Norian is most probably global in extent. Negative carbon-isotope excursions occur in Norian substages: (upper) Alaunian, lower Sevatian, middle Sevatian and upper Sevatian. Distinct positive δ13C excursions are present across the Alaunian-Sevatian boundary and the Norian-Rhaetian boundary, and the climax of the latter is located in the lower part of the posthernsteini Zone. This last isotope event is particularly important because at the Global Stratotype Section and Point (GSSP) candidate for the base of the Rhaetian stage at Pignola (Southern Apennines, Italy) a distinct negative δ13C excursion has been reported for this stratigraphic level.

A new fossil insect locality from the Upper Triassic of Jiyuan, Henan Province, Central China

The Triassic insects are relatively abundant from the Anisian to Rhaetian stages. Several well-known Triassic insect localities are distributed worldwide. Examples include the Anisian Grès à Voltzia in France, the Ladinian–Carnian Madygen Formation in Kyrgyzstan, Uzbekistan, and Tajikistan, the Cow Branch Formation of the latest Triassic in the USA, and the Carnian Molteno Formation in South Africa (Grimaldi & Engel, 2005).

Palynological investigation of an Upper Triassic sequence in Nanjing area, eastern China, with paleoecological implications

The end-Triassic is characterized by a significant biotic crisis in both marine and terrestrial realms. However, terrestrial records in eastern Tethys are relatively limited, especially in eastern China. The Fanjiatang Formation represents the Upper Triassic sequence in southern Jiangsu and Anhui provinces of eastern China, yielding a rich plant and bivalve fossil record. The present study provides the first detailed palynological investigation for the Upper Triassic of Nanjing area, eastern China. Six of twenty processed palynological samples are productive, and more than 70 terrestrial spore and pollen fossil taxa were identified. A palynological assemblage, namely, the Concavisporites–Dictyophyllidites–Kyrtomisporis–Classopollis Assemblage was established, characterized by abundant fern spores, mainly including Concavisporites, Dictyophyllidites and Kyrtomisporis, and much less abundant gymnosperm pollen grains (important taxa include Alisporites, Chasmatosporites, and Classopollis), indicating a Late Triassic Norian–Rhaetian age. In conjunction with previously reported macro-plant data, the palynofloral analysis reveals a vegetation dominated by ferns, with diverse cycadophytes, less abundant lycophytes, bryophytes, equisetales, ginkgophytes and seed ferns, and a few conifers. Paleovegetation and semiquantitative palynological analysis permit a paleoecological interpretation, suggesting a lowland groundcover-dominated vegetation under (sub)tropical warm and humid climate in Nanjing area during this time interval. More detailed palynological and macro-plant data through the whole Upper Triassic in this region are necessary in the future to better reveal the Late Triassic paleoecological settings and variations on land prior to the end-Triassic mass extinction in eastern Tethys.

Magnetostratigraphy of the Mercia Mudstone Group (Devon, UK): implications for regional relationships and chronostratigraphy in the Middle to Late Triassic of Western Europe

Global synchronization of environmental change in terrestrial successions in deep time is challenging owing to the paucity of dating methods, a case also applicable to the Middle to Upper Triassic Mercia Mudstone Group in Britain. Using coastal cliff sections, magnetostratigraphy was evaluated at 263 horizons, defining 53 magnetozones. Magnetozones from the lower 140 m of the group demonstrate correspondence to those from the mid-Ladinian to early Carnian polarity timescale, dating that is compatible with magnetostratigraphy from the underlying Sherwood Sandstone Group. Magnetostratigraphy of the Dunscombe Mudstone Formation, and associated palynological data, suggest a late Carnian to earliest Norian age, and a dramatically lower accumulation rate than for adjacent formations. The polarity record demonstrates coeval flooding events, evaporite deposits and intervals of sand supply between the Wessex Basin and the Central European Basin in the Carnian. This is the result of linked climatic and eustatic changes between these separate basins, related to aeolian dust supply and the shrinkage of hyper-arid source regions for the fine material. Magnetostratigraphy from the Branscombe Mudstone and Blue Anchor formations demonstrates their Norian and early Rhaetian age. These and other data suggest an alternative synchronization of marine and non-marine polarity records for the Norian polarity timescale. Supplementary material : Section details and detailed logs of the sampled sections and inferred sequence boundaries, magnetic mineralogy data, demagnetization behaviour and mean directions, summary of virtual geomagnetic pole data and a comparison with other European poles, construction of other composite reference sections and revised polarity scales, an Excel sheet of magnetic data, statistically evaluated correlation models and age models are available at https://doi.org/10.6084/m9.figshare.c.6613788

Uppermost Triassic Halstätt-like cephalopod limestone (Lilu Facies) and Foraminifera, Timor-Leste

Ammonoid-rich wackestone beds forming a ca 2 m section between shallow-marine limestones of the Bandeira Group (Norian to possibly Rhaetian) and a wackestone–calcareous mudstone succession belonging to the Lower Jurassic Wailuli Group are recorded from the western part of Timor Leste. They are included in the Halstätt-like Lilu Facies of the Bandeira Group. Age-diagnostic involutinid foraminiferal species, not previously recorded from Timor, include Involutina liassica, Trocholina sp. ex. gr. Trocholina turris and Lamelliconus permodiscoides, which collectively indicate a Rhaetian age, possibly ranging down to the Sevatian. The conodont Norigondolella steinbergensis, recorded low in the section, is consistent with this age determination. The Lilu Facies was deposited at a water depth of a few tens of metres, as suggested by the presence of extensive cyanobacterial borings into shell fragments and the presence of microbial laminations including the presence of stromatolites and bacinellid-like fabrics. Among macrofossils, common ammonoids are present in the section, and marine reptile bones are conspicuous in the lower beds. Other biogenic components of the rock observed in acetate peels and thin sections are abundant echinoderm debris (including pentacrinoid columnal plates), bivalve fragments and micro-gastropods, rare siliceous sponge spicules, solitary corals, and brachiopod debris. In the studied region, a major drowning of a shallow-marine carbonate platform took place probably in the Rhaetian or possibly latest Norian. Similar ‘ammonitico rosso limestone’ interbedded with thick-bedded limestones of a shallow-marine carbonate platform is present in the uppermost Triassic of Seram. More detailed biostratigraphical comparisons and correlations of similar units in Timor and other nearby islands may indicate a general marine drowning of Late Triassic carbonate platforms, at least in the northern part of the East Gondwana Interior Rift system, during the Rhaetian or possibly latest Norian.

Lost Norian fluvial tracks: Sedimentology and stratigraphy of the Upper Triassic coarse-grained deposits in Kamienica Śląska (Upper Silesia, southern Poland)

In the Triassic‒Jurassic boundary profiles of the Upper Silesian region, there are locally developed coarse clastic deposits, commonly known as the Połomia Gravels, Połomia Beds or “Połomia Formation’’ (informal name). Due to the lack of reliable stratigraphic tools, the chronostratigraphic position of these deposits remains one of the most controversial aspects of the Triassic‒Jurassic lithostratigraphy in the region. Sparse biostratigraphic data from the overlying and underlying deposits indicate a wide range of ages, from the Late Triassic to Early Jurassic. This paper presents the results of sedimentological, petrographical and palynological analyses of the coarse-grained deposits from the Kamienica Śląska gravel pit, which is currently one of the biggest facilities of this type in Upper Silesia. The outcrop section, with a total thickness of up to 20 m, is dominated by friable, light grey to beige and locally dark grey, large-scale planar to trough cross-stratified conglomerates and coarse-grained sandstones with subordinate thin interbeds of grey and reddish brown mudstones. Three facies associations have been distinguished, representing a main channel belt, secondary channels and floodplains subenvironments. Facies analysis points to a braided river tract with localised floodplain sediments. The grain composition of the Kamienica Śląska gravel/conglomerate is less diversified than that of the polymictic typical Połomia Beds of the Myszków area and resembles oligomictic conglomerates known from the Grabowa Formation of the Norian age and/or conglomerates of the Gorzów Beds of Rhaetian age, which also occur on the studied region. Palynological analysis of mudstone interbeds within the conglomeratic deposits shows the presence of miospores guiding and characteristic for subzone c of the Corollina meyeriana zone of the late Norian‒early Rhaetian age. The appearance of the coarse-grained deposits in late Norian could be associated with the development of a long-reach braided fluvial tract, draining the S and SE part of the Sudetian-Malopolska-Lublin land (S-M-L land) in response to the tectonic rearrangement in the source area and gradual climatic change from semi-arid to humid in the Rhaetian.

The Upper Triassic deposits of the west Bangong-Nujiang suture zone and their paleogeographic implications

The Bangong-Nujiang Suture Zone (BNSZ) of Tibet (Xizang) has been interpreted to represent a relic of the Bangong-Nujiang Ocean. However, the existence of this ocean during Triassic time remains a point of contention. A sedimentary succession spanning the Upper Permian through Triassic described from the central BNSZ suggests that the Lhasa and South Qiangtang terranes were contiguous thus negating the existence of a terrane-separating ocean during Triassic time. However, the apparent lack of Triassic deposits in the west BNSZ has called into question the existence of Triassic deposits in the central region of the BNSZ. Our biostratigraphic work in the Wuga Formation of the Gaize area has yielded abundant Norian conodonts thus confirming the existence of Upper Triassic deposits in the west BNSZ. The clastic deposits of the Wuga Formation are herein interpreted to be of Rhaetian age. Moreover, intercalated limestone and chert are termed the Dongnale Formation of Norian age. The Norian to Rhaetian succession can be correlated with strata of the central BNSZ as well as with deposits of the Lhasa Terrane and the South Qiangtang Terrane. Similar stratigraphies among these regions through the Late Triassic suggests a shared depositional setting and that the BNSZ was not an ocean in Norian and Rhaetian time.

Dinosaur fossil records from India and their palaeobiogeographic implications: an overview

The complex palaeogeographic history of India involving a gradual transition from Gondwana to Laurasia with an intervening phase of prolonged physical isolation, and the biotic signatures of this complex history as preserved in India’s Mesozoic fossil record are of much current interest and continue to be debated. Seen in this context, the fossil record of dinosaurs from India provides a unique opportunity to study their diversity and palaeobiogeographic distribution in time and space. The Indian fossil record, as currently documented, is patchy and restricted mainly to three intervals of the Mesozoic era: Late Triassic, Early/Middle Jurassic and Late Cretaceous. The Late Triassic–Jurassic record, representing a Pangean setting, is known primarily from the Gondwana formations of Pranhita–Godavari (P–G) Valley in the southern Indian state of Andhra Pradesh, although sporadic Jurassic occurrences are also known from Kutch (Gujarat) and Rajasthan. The earliest Late Triassic dinosaur fauna of India comes from the rhynchosaur–dominated Lower Maleri Formation of Carnian age. Known from fragmentary and isolated specimens, the Late Triassic dinosaur fauna is currently represented by the sole species Alwalkeria maleriensis, which is possibly a basal saurischian with uncertain relationships. A slightly younger dinosaur fauna from the archosaur–dominated Upper Maleri Formation of late Norian–earliest Rhaetian age consists of a more diverse assemblage including the two named basal sauropodomorphs (Nambalia roychowdhurii and Jaklapallisaurus asymmetrica). In contrast to the Late Triassic, the Early Jurassic record of Indian dinosaurs described from the Upper Dharmaram and Lower Kota formations of P–G Valley, is far more abundant, diverse and based on more nearly complete material that is currently referred to four named taxa of stem sauropodomorphs or basal sauropods (Lamplughsaura dharmaramensis, Pradhania gracilis, Kotasaurus yamanpalliensis, Barapasaurus tagorei) plus an ornithischian (Ankylosauria). Kotasaurus, one of the earliest known sauropods, is more primitive than Barapasaurus and shared numerous plesiomorphic characteristics with prosauropods. Together, the Late Triassic and Early Jurassic sauropods dinosaurs of India document the early radiation of this group. Amongst the other important records of Jurassic dinosaurs in India is the oldest known camarasauromorph sauropod whose identification is based on a metacarpal, a first pedal paw and a fibula from the Middle Jurassic (Bajocian) strata of Khadir Island, Kutch. Fragmentary postcranial skeletal material of an unidentified Middle Jurassic dinosaurs is also known from Kuar Bet (Patcham Island) in the Rann of Kutch and the Jumara area of Kutch Mainland. Post–Gondwana, the Late Cretaceous dinosaurs of India occur in a different geodynamic setting in which the Indian Plate, as traditionally considered, was a northward drifting island continent in the middle of the Indian Ocean. Apart from the solitary record of a Cenomanian–Turonian sauropod from Nimar Sandstone, Cretaceous dinosaurs from India are documented mainly by skeletal remains and eggs/eggshells from the Maastrichtian infratrappean (=Lameta Formation) and intertrappean deposits in the Deccan Volcanic Province of eastern, western and central peninsular India, and from broadly coeval Kallamedu Formation of Cauvery Basin, southern India. Skeletal remains of the Lameta dinosaurs belong to two major groups, titanosaur sauropods and abelisaurid theropods, plus a possible ankylosaur, whereas the Cauvery records include fragmentary titanosaur bones and a solitary tooth of a troodontid theropod. Apart from bones and teeth, a number of dinosaur egg–bearing nesting sites are also known to occur in the Lameta Formation of east–central and western India, extending for more than 1,000 km across the states of Madhya Pradesh, Gujarat and Maharashtra. Close phylogenetic relations of the Lameta titanosaurs and theropods with corresponding taxa from the Maastrichtian of Madagascar (Vahiny, Majungasaurus) and the rare occurrence of Laurasian elements such as a troodontid, pose interesting palaeobiogeographic problems in the context of India’s supposed oceanic isolation, especially after its separation from Madagascar at ~ 88 Ma.

The timing of rifting events in the easternmost Mediterranean: U-Pb dating of zircons from volcanic rocks in the Levant margin

ABSTRACT The Levant basin, at the easternmost Mediterranean, formed during the opening of the Neo-Tethys at Permian to Jurassic times, as recorded by subsidence and magmatism along its SE passive margin. While the subsurface extensional structure was reconstructed in detail, the spatial and temporal distribution of igneous activity is not well known. Here we present new SIMS U-Pb dating of zircons from rifting related volcanic sequences recovered from four deep boreholes in the Levant margin of coastal Israel. The Asher volcanics, a deeply buried 2.5 km thick basaltic sequence, which accumulated within fault-bounded basin, was previously dated by K-Ar and Ar-Ar to early Jurassic times. Zircons (n=46) separated from the bottom and top of the volcanic sequence yielded 206±2 Ma and 205 to 202 Ma, respectively, indicating that Asher volcanics are of Late Triassic (Rhaetian) age. Zircon xenocrysts indicate Permian (289±2 Ma) felsic magmatism. Rifting activity in the Levant margin included several Permian-Triassic pulses of faulting and subsidence, with a prolonged magmatism-bearing rifting event at Late Carnian to Rhaetian times (~230-201 Ma). Pulsed rifting is also indicated in the conjugate, northern, margin of the southern Neo-Tethys, and the timing of subsidence and magmatic pulses well correlate to the Levant margin.

New data on the palynology of the Triassic–Jurassic boundary of the Silves Group, Lusitanian Basin, Portugal

New evidence is presented on the Triassic–Jurassic boundary in the northern Lusitanian Basin, Portugal, based on miospore assemblages from a composite Upper Triassic to Lower Jurassic succession of the Silves Group. The latter comprises, from base to top, the Conraria, Penela, Castelo Viegas and the Pereiros formations. Three informal palynological zones have been documented and compared with coeval palynozones from West and South Europe providing new biostratigraphic data to detail the age of the lower and upper formations of the Silves Group and to review previous age attribution.A Norian, possibly earliest Rhaetian age, is documented for the Conraria Formation on the basis of a palynological assemblage referable to the Classopollis meyerianus–Granuloperculatipollis rudis (CG) zone. The Penela and Castelo Viegas formations did not allow a palynostratigraphic revision, due to the not promising lithology for palynological studies. The Pereiros Formation is dated on the basis of microflora assemblages referable, from bottom to top, the Ischyosporites variegatus–Kraeuselisporites reissingeri (IK) zone of late Rhaetian–earliest Hettangian age and Pinuspollenites minimus (Pm) zone of Hettangian age. The discontinuity between the underlying Castelo Viegas Formation and the overlying Pereiros Formation did not allow to define the lower boundary of the IK palynozone. The Triassic–Jurassic boundary lies in the lower part of Pereiros Formation within the IK zone. The microflora assemblages from the Lusitanian Basin show close affinity to those of eastern N America and western Tethys areas.

Rhaetian (Late Triassic) ostracods (Crustacea, Ostracoda) from the offshore prolongation of the North Dobrogean Orogen into the Romanian Black Sea shelf

The present study evaluates the significance of Triassic ostracods from the Romanian Black Sea shelf as part of a project for the understanding of the palaeoceanographical evolution of the offshore extension of the North Dobrogean Orogen. The drill core CM31, sampled from the borehole 817 Lebăda Vest drilled on the western Romanian portion of the Black Sea shelf, contains sediments of Rhaetian, Late Triassic age. The taxonomy of ostracods obtained is discussed and adds to the scientific understanding of marine ostracods during the Rhaetian stage. We document 72 species, of which seven are new: Histriabairdia pontuseuxinusensis Forel gen. et sp. nov., Bairdiacypris argonautaii Forel sp. nov., Ceratobairdia? akhilleusi Forel sp. nov., Isobythocypris atalantella Forel sp. nov., Petasobairdia amazonella Forel sp. nov., Paracypris ovidi Forel sp. nov. and Pseudomacrocypris? kerabani Forel sp. nov. Histriabairdia Forel gen. nov. is introduced to accommodate species of the Triassic–Jurassic interval previously attributed to the modern genus Anchistrocheles. These assemblages point to an outer shelf environmental with relatively normal marine oxygenation. The oldest occurrence of Pokornyopsis, found in this material, indicates that this forerunner of modern troglobitic ostracods may not have been troglobitic in the Triassic.

A new ammonite from the Penarth Group, South Wales and the base of the Jurassic System in SW Britain

AbstractTwo ammonites have been found in porcelaneous limestones of the White Lias Formation traditionally considered to be of Rhaetian (Late Triassic) age at Lavernock Point, south Wales (ST 181 681). These ammonites, named here Neophyllites lavernockensis. sp. nov, are the earliest recorded from the UK. This horizon is located directly above a major negative δ13C isotope anomaly (CIE1) in the Upper Cotham Member that marks the top of the Triassic System and below another negative δ13C isotope anomaly (CIE3). Both correlate with negative δ13C isotope anomalies in the Triassic–Jurassic global boundary stratotype section and point (GSSP) at Kuhjoch, Austria. This establishes the base of the tilmanni Chronozone, Hettangian Stage and Jurassic System in SW Britain within the White Lias Formation at Lavernock Point.

The first tetrapod track recorded from the Rhaetian in the British Isles

Terrestrial vertebrate trace fossils are relatively abundant in mid-to-late Triassic and early Jurassic deposits in the British Isles but to date none at all have been recorded from the Rhaetian, the final stage of the Triassic. This represents a persistent gap in the terrestrial ichnological record. We present the first Rhaetian track to be recognised in the British Isles, found at Aust Cliff on the south bank of the Severn Estuary near Bristol in SW England. This locality is well known for disarticulated remains of Rhaetian fossil reptiles including some terrestrial species but in 2006 a track (TECMAG0161) was found for the fist time. Although the specimen was found ex-situ the palynological data from the surrounding matrix confirms a Rhaetian age. The track was examined with CT scanning and photogrammetry. We tentatively assign the track to the ichnogenus Procolophonichnium based on size and digit proportions. The isolated nature of the specimen offers little concrete information about the track maker but such tracks have previously been attributed to parareptiles or therapsid trackmakers. The specimen adds a datapoint to an otherwise ichnologically empty period of time in the British Isles. The track also provides solid evidence for a [locally] terrestrial environment in a sequence that is otherwise considered predominantly marine or estuarine. This discovery suggests that there may be more such tetrapod tracks of Rhaetian age preserved, at least at Aust, and further searching will hopefully lead to the current minimal dataset being expanded.

Halorina cryptica nov. ichnogen., nov. ichnosp., mass-occurrence of Upper Triassic crustacean microcoprolites from neptunian dikes and sills cutting the Dachstein-type carbonate platform and their paleoenvironmental significance (Northern Apuseni Mountains, Romania)

The new crustacean microcoprolite Halorina cryptica nov. ichnogen., nov. ichnosp., is reported from cryptic cavities cutting the Upper Triassic (Carnian to lower Rhaetian) Dachstein-type limestones from the Northern Apuseni Mountains, Romania. The new ichnotaxon is extremely abundant in cavities, neptunian dikes and sills filled with red ferruginous carbonate sediment. The associated microfauna consists of ostracods and rare foraminifera. The microfacies is represented by bioclastic coprolite-bearing wackestone-packstone to grainstone. The red ferruginous carbonate fillings are strongly bioturbated. The neptunian sill located at the top of the studied section contains a rich ichnofauna associated with brachiopod accumulations. It is dominated by the dimerelloid rhynchonellid Halorella, indicating a late Norian-early Rhaetian age. Although neptunian dikes and sills are rather common in the Dachstein-type carbonate platform that extended on the northwestern Tethyan margin, we report here the first record of mass-occurrence of crustacean microcoprolites in neptunian dikes and sills filled with red carbonate sediments of marine origin. They are interpreted here as cryptic cavities with specific palaeoenvironmental features (e.g., lack of light, abundant nutrients, intensive microbial activity) where crustacean arthropods thrived.

"Short" or "long" Rhaetian ? Astronomical calibration of Austrian key sections

The establishment of the Late Triassic time scale has given rise to considerable controversy, particularly regarding the Rhaetian duration and the inferred absolute age models. In this respect the astronomical polarity time scale (APTS) established from the continental successions of the Newark Basin (eastern North America) is considered as a reference record, although its completeness is questioned. Numerous magnetostratigraphic correlation schemes have been proposed between the APTS and biostratigraphically well-constrained Tethyan marine sections. This has led to two main contrasting options: a “short” Rhaetian duration (about 4–5 myr), and a “long” one (about 8–9 myr). Astronomical calibration of the Tethyan Rhaetian and estimate of its duration is necessary to help advance this debate. We have undertaken a cyclostratigraphic analysis of a Rhaetian composite record built from four overlapping Austrian reference sections. Magnetic susceptibility variations of the 131.5 m thick record are astronomically paced by the precession and 405-kyr orbital eccentricity cycles. 405-kyr orbital tuning allows to establish a floating time scale, and thus to suggest a minimum duration of 6.69 myr of the Rhaetian stage. Given the well-established radioisotopic age of the Rhaetian-Hettangian boundary of 201.36 Ma, an age no younger than 208.05 Ma for the Norian-Rhaetian boundary can be proposed. This result will contribute to the refinement of the Late Triassic time scale, but it does not solve the long-standing debate on bio-magnetostratigraphic correlations between the continental Newark APTS and the Tethyan marine sections, nor the question of the completeness of the Rhaetian Newark Basin. (Less)

STRATIGRAFIA E SEDIMENTOLOGIA DEI "CALCARI E MARNE A <em>RHAETAVICULA CONTORTA</em> AUCTT. (NORICO-RETICO) DELLA TOSCANA NORD-OCCIDENTALE - NUOVA SUDDIVISIONE FORMAZIONALE

Nine sections from the "Limestones and marls with Rhaetavicula contorta" Auctt. (Norian-Rhaetian) of the Tuscan Nappe in eastern Liguria and northwestern Tuscany, have been examined for their stratigraphic and sedimentological characteristics. The results obtained have been integrated with data pertaining to the previously studied Pania di Corfino section.
 Two formations were identified and studied in detail: that of Pania di Corfino and that of La Spezia.
 Where present, the Pania di Corfino Formation accounts for the lower portion of the sections. It consists mainly of thick beds and massive layers of dolomites, dolomitic limestones and limestones. Thinly-bedded dark limestones appear less frequently, while layers rich in clay are rare. The top of the formation is surely of Rhaetian age, in some sections, as indicated by the presence of Triasina hantkeni Majzon; in the others, it is Norian-Rhaetian. The age of the bottom portion can be regarded as Norian-Rhaetian in all the sections; the presence of considerably thick deposits beneath the established Rhaetian layers in some sections, however, suggests an Early Rhaetian or even Late Norian age.
 The depositional environment of the Pania di Corfino Formation may be envisaged as an epicontinental carbonate platform with predominantly restricted water circulation on which fine carbonate sediments are deposited beneath the wave base; features suggesting a tidal flat are rare. The environment of deposition was, for brief periods only, that of a platform with open circulation in which the effects of wave-motion were prominent; features strictly relatable to a true platform edge have not been found.

Anatomy of a Late Triassic Bristol fissure: Tytherington fissure 2

The Late Triassic and Early Jurassic fissures around Bristol and South Wales are famous as sources of excellent fossils of early mammals, dinosaurs, sphenodontians, and other tetrapods. However, the ways in which these fissures filled with sediment have not often been documented. Moreover, systematic faunal sampling up a working face exposure has rarely been attempted for the UK Triassic fissures. Here we show in detail how a complex fissure in Tytherington Quarry, north-east of Bristol, filled in multiple steps, and document both sedimentary facies and faunas. Tytherington Quarry is a key Late Triassic Bristol sauropsid fissure site, having previously yielded abundant Thecodontosaurus remains and a new genus of sphenodontian, Diphydontosaurus. Our findings comprise specimens of terrestrial reptiles, including sphenodontians and archosaurs. However, abundant remains of fishes and marine invertebrates were also recovered, indicating a significant marine influence during parts of the fissure-filling process. Comparisons of our results with those of previous studies on Tytherington and other Late Triassic SW UK fissure sites indicate some consistent patterns of faunal association and provide further evidence of a Rhaetian age for the Late Triassic palaeo-island fissures north of Bristol. Our analysis supports a model of fissure formation and filling entailing punctuated episodes of sedimentation through a relatively short time.