The Western Cordillera and forearc of the Northern Andes (north of 5° S) are constructed from allochthonous terranes floored by oceanic crust. A paucity of accurate geochronological data from the ultramafic and mafic basement sequences of the Western Cordillera and Coastal region of Ecuador has lead to distinct differences in published tectonic models, with different allochthonous units and contrasting timelines of accretion and collision. We present new zircon UPb and groundmass 40Ar/39Ar dates of the basement units, which have been combined with new whole rock geochemical and isotopic data to constrain their tectonic histories prior to and during their accretion to South America. Ultramafic and mafic rocks with oceanic plateau affinities can be divided into two plateau sequences, where an inboard, younger plateau (85–92 Ma; Pallatanga Block) is sutured against South America, and was modified by a pre-accretionary intra-oceanic arc (Pilatón-Pujilí arc) that was terminated by the accretionary process. An older, outboard oceanic plateau sequence (∼120 Ma) forms the basement to a series of coastal blocks within the present forearc (Piñon, San Lorenzo, Pedernales and probably the Esmeraldas blocks). The outer plateau also hosts a pre-accretionary intra-oceanic arc (Pascuales – San Lorenzo arc), which remained active during and post-accretion (San Lorenzo – Rio Cala arc). Coeval changes in palaeomagnetic declination, the onset of rapid exhumation (>1 km/My) of the Early Cretaecous continental margin and sedimentation within the newly formed retro-arc foreland basin and foreland region supports a model where both oceanic plateau sequences accreted to South America in a single accretion event at 75–70 Ma. Thus, it is likely that the accreted oceanic plateau was composite, and included both the younger and older plateau sequences prior to accretion. We test an alternative model where the inner (younger) and outer (older) oceanic plateaus accrete in sequence, although this fails to account for the timing of block rotations in the forearc.