Reptilian Remains Research Articles

Palaeobiological and geochemical aspects of reptilian coprolites from a Maastrichtian Deccan volcano-sedimentary intertrappean deposit of central India

Cretaceous-Paleogene coprolite (fossilized faecal matter) records are significant in terms of providing direct palaeobiological evidence (from inclusions) in order to understand the diet of producer animal(s). In the past >150 years, research investigations (in India) have focussed on the Maastrichtian Type-A coprolite morphotype. Consequently, little information is available on the overall assemblage of Indian Maastrichtian vertebrate coprolites in terms of their morphological diversity, chemical composition, biotic-abiotic inclusions in the context of producer linkage(s) and geographic distribution within the Deccan volcano-sedimentary infra- and intertrappean deposits of India. Therefore, we present a detailed record of a coprolite assemblage from the Maastrichtian intertrappean deposits of Lotkheri, central India. This coprolite assemblage comprises five morphotypes based on their geometry, surface, and internal textures. Morphological considerations and biotic inclusions suggest that reptiles were the most likely producers of these ichnofossils. The associated faunal remains of reptiles (mainly chelonians and crocodilians) support this hypothesis. Dentalites or bite traces of Garfish (genus Lepisosteus) observed on the external surface of a few coprolite specimens (studied herein) are a rarity in the global fossil records. The analytical evidence confirm the phosphatic composition of the coprolites with the unique presence of three distinct morphologies (i.e. spherical, rods, and needles) of apatite crystals. Finally, we provide the chemical nature and plausible mechanism of apatite crystal formation via the chemical processes undergoing during the biomineralization of these crystals observed within coprolites.

Diet of Rusty-spotted Cat Prionailurus rubiginosus (I. Geoffroy Saint-Hilaire, 1831) (Mammalia: Carnivora: Felidae) in Sanjay Gandhi National Park, Mumbai, India

The 103.68 km2 Sanjay Gandhi National Park (SGNP), Mumbai, exists amidst human densities that figure among the highest in the world. The rich biodiversity of SGNP includes the Rusty-spotted Cat Prionailurus rubiginosus, endemic to India, Sri Lanka, and Nepal, and categorised as ‘Near Threatened’ on the IUCN Red List. Little is known about its ecology and the dynamics of its coexistence with the other small carnivores in SGNP. We conducted a study with citizen volunteers to explore the diet of the Rusty-spotted Cat and other sympatric small carnivores in SGNP and in the adjoining human-dominated areas of Yeur village, Shivaji Nagar, Dahisar Quarry, and Aarey Milk Colony. After initial training, the volunteers collected scat samples from all forest ranges in SGNP and the surrounding areas outside, following defined protocols. Seventy-eight scat samples were analysed for species assignments using standardised molecular techniques, felid-specific primers, and DNA sequencing, and 24 were identified as of the Rusty-spotted Cat. The contents of the samples were examined under a microscope to identify prey remains. Results were presented as the mean number of scat samples containing remains of specific taxa with 95% Confidence Intervals. Diet estimated from 22 Rusty-spotted Cat scat samples and 52 samples of other small carnivores revealed rodents to be the major prey of the entire group. However, a higher proportion of Rusty-spotted Cat scat samples had remains of rodents (95%) and reptiles (6%) as compared to samples of other small carnivores, i.e., 79% with rodent remains and none with remains of reptiles. On the other hand, a lower proportion of Rusty-spotted Cat scat samples had remains of insects (14%), plant matter (9%), and birds (5%) than samples of other small carnivores (40% plant matter, 38% insects, 17% birds). Our results highlight the role of small carnivores, especially Rusty-spotted Cat in regulatory services through pest control.

Callovian Marine Reptiles of European Russia

Our knowledge of marine reptiles of the Callovian age (Middle Jurassic) is majorly based on the collections from the Oxford Clay Formation of England, which yielded a diverse marine reptile fauna of plesiosaurians, ichthyosaurians, and thalattosuchians. However, outside of Western Europe, marine reptile remains of this age are poorly known. Here, we survey marine reptiles from the Callovian stage of European Russia. The fossils collected over more than a century from 28 localities are largely represented by isolated bones and teeth, although partial skeletons are also known. In addition to the previously described rhomaleosaurid and metriorhynchids, we identify pliosaurids of the genera Liopleurodon and Simolestes; cryptoclidid plesiosaurians, including Cryptoclidus eurymerus, Muraenosaurus sp., and cf. Tricleidus, and ophthalmosaurid ichthyosaurians, including the iconic Ophthalmosaurus icenicus. These findings expand the ranges of several Callovian marine reptile taxa far to the Eastern Europe, and support the exchange of marine reptile faunas between Western and Eastern European seas in the middle to late Callovian. However, some specimens from the lower Callovian of European Russia show differences from typical representatives of the middle Callovian Oxford Clay fauna, possibly representing the earlier stages of evolution of some of these marine reptiles not yet recorded in Western Europe or elsewhere.

Crocodile Rock! A Bioarchaeological Study of Ancient Egyptian Reptile Remains from the National Museum of Lithuania

Remnants of what was believed to be a single baby crocodile, originating from ancient Egypt and curated in the National Museum of Lithuania, have been recently assessed using noninvasive and nondestructive techniques. These had been donated in 1862 to the then Museum of Antiquities by the prominent Polish-Lithuanian collector Count Michał Tyszkiewicz. After careful investigation of the three mummified reptile fragments available, the authors were able to identify at least two individuals based on morpho-anatomical characteristics. This indicates that the two small crocodiles originally described in historic records are still present within the collection and that none of these items was lost during the different lootings perpetrated throughout the museum’s history. Information regarding the post-mortem treatment of these animals was also obtained. This is the first scientific study of animal mummies in the Baltic States, and it should be followed by proper conservation and display of these findings.

Osteology and phylogenetic relationships of a new archosauriform reptile from the Middle Triassic (Anisian) of Germany

ABSTRACT Skeletal remains of archosauriform reptiles from the Lower and lower Middle Triassic Buntsandstein Group in the Central European Basin are rare. This paper reports on a partial, almost completely disarticulated skeleton of a previously unknown archosauriform reptile from the lower Middle Triassic (Anisian) Röt Formation of Rotfelden in Baden-Württemberg (Germany). Marcianosuchus angustifrons gen. et sp. nov. is distinguished by the following combination of features: posterodorsal process of premaxilla slender, slightly inclined posterodorsally, and with rounded apex; posterodorsal surface of frontal covered by slightly diverging, fine longitudinal grooves; squamosal with distinct lateral ridge extending for entire length of element; teeth with proportionately small, weakly recurved crowns; humerus considerably shorter than femur; distal shaft of ischium rod-like, not plate-like; pubis with flat distal portion forming ‘pubic apron’: neural spines of posterior cervical and anterior dorsal vertebrae with transversely greatly expanded apices that are Y-shaped in anterior or posterior view; dorsal vertebrae with centra taller dorsoventrally than long anteroposteriorly; dorsal osteoderms more or less rectangular in outline, longer anteroposteriorly than wide mediolaterally, with slightly rounded anterior and concave posterior margins and bearing dorsal ridge or eminence; and dorsal surfaces of osteoderms with unsculptured anterior region and posterior region bearing keel or eminence and covered by sculpturing of radially arranged grooves and pits. Marcianosuchus angustifrons represents the first definitive record of a non-archosaurian archosauriform from the Buntsandstein Group of Germany. Its body plan most closely resembles that of Euparkeria capensis among well-known non-archosaurian archosauriform reptiles, but the two taxa are clearly distinct.

New small reptile remains from the Late Cretaceous of Patagonia increase morphological diversity of sphenodontids (Lepidosauria)

Sphenodontids are a group of reptiles that were diverse and global for much of the Mesozoic but today they are only represented by the New Zealand tuatara. Here we describe new sphenodontid remains coming from the Cretaceous (Maastrichtian) Los Alamitos Formation, at Río Negro province, Argentina. Previous reports in the stratigraphical unit included an indeterminate sphenodontid and the enigmatic Kawasphenodon. The new material here reported includes an isolated and incomplete dentary and a palatine bone. The dentary belongs to a small taxon having quadrangular teeth and an interlocked mandibular symphysis. The palatine shows compressed teeth and a fang-like canine. These specimens probably belong to two new taxa. These, together with Kawasphenodon, suggest that Maastrichtian sphenodontians from northern Patagonia were at least as diverse as those reported from Cenomanian beds. This contrasts with the poorer record of lizards in the same beds. However, the record of Maastrichtian sphenodonts in southern Patagonia is restricted to a single finding of a Sphenodontine sphenodontid. In Australasia the sphenodont record is also restricted to Cenozoic sphenodontines, very similar to extant Sphenodon species. The currently available fossil record suggests that northern Patagonian rhynchocephalians were more morphologically diverse than those of southern Patagonia and Australasia during the Cretaceous, probably reflecting another faunistic particularity of the 'Weddelian Bioprovince'.

Northernmost occurrences of plesiosaurs and turtles in the Upper Cretaceous of Eurasia

Here we describe the first findings of reptile remains in the Upper Cretaceous marine strata of Siberia, which represent the northernmost occurrences of plesiosaurians, ?mosasaurids and turtles in the Cretaceous of Eurasia (66–72° N palaeolatitudes). The specimens come from the upper Cenomanian, Turonian and Coniacian of the Pyasina River basin, Santonian and Maastrichtian of the Tanama River and upper Santonian–lower Campanian of the Kheta River. They include remains of polycotylid and elasmosaurid plesiosaurians, as well as indeterminate small and osteologically immature plesiosaurians, indicating a somewhat typical Late Cretaceous plesiosaurian assemblage at the Arctic Polar Circle and implying that these shallow-water regions possibly used by plesiosaurians as a birth and nursery areas during the polar summer. In addition to plesiosaurian remains, a partial carinate ?mosasaurid tooth, and two turtle shell fragments, assigned to either non-marine Macrobaenidae or to marine Chelonioidea, and to non-marine Testudinoidea, are discovered, providing first direct evidence for the presence of these reptile groups in the Late Cretaceous of Northern Siberia. The findings of turtles are the northern most records of this group in the Cenomanian and Coniacian of Eurasia, which parallels previous records from the upper Cenomanian of Arctic Canada, and further supports warm climate conditions during the Cretaceous Greenhouse at high latitudes. Therefore, the Cenomanian–Turonian Boundary Event associated to the climate warming and to the beginning of the Cretaceous thermal maximum is well-recognized in Northern Siberia showing an expression of this “global” event over the Arctic region.

“A Crocodile Spirit, Crocodile-Faced”: Discovery of Crocodile Remains in the Early Middle Kingdom Tombs of the North Asasif Necropolis in Western Thebes (Egypt)

Abstract The recent discovery of Nile crocodile remains in the mortuary complexes of two high-ranking courtiers of Nebhepetra Mentuhotep II, located in the early Middle Kingdom necropolis in the valley of North Asasif, opened the way to an exploration of the role of reptile remains in funerary contexts. The skeletal remains, which were not mummified, consisted of fragments of the skull and mandible, loose teeth, and osteoderms. This paper explores the association that may have existed between the deceased and the crocodile god Sobek, whom the ancient Egyptians identified with pharaonic power, inundation and fertility. From the Middle Kingdom, Sobek, who was believed to have risen from the Primeval Waters, was merged with the sun-god Ra, and in the solar form of Sobek-Ra was made part of the eternal journey of the sun from the east to the west. This association was also reflected in the Spells of the Coffin Texts, in which the deceased became Sobek.

Early Triassic ichthyopterygian fossils from the Russian Far East

Ichthyopterygia is a major clade of reptiles that colonized the ocean after the end-Permian mass extinction, with the oldest fossil records found in early Spathian substage (late Olenekian, late Early Triassic) strata in the western USA. Here, we describe reptilian remains found in situ in the early Spathian Neocolumbites insignis ammonoid zone of South Primorye in the Russian Far East. Specimen NSM PV 23854 comprises fragmentary axial elements exhibiting a combination of morphological characteristics typical of Ichthyopterygia. The cylindrical centra suggest that the specimen represents a basal ichthyopterygian, and its size is comparable to that of Utatsusaurus. Specimen NSM PV 24995 is represented by a single limb bone, which is tentatively identified as an ichthyopterygian humerus. With a body length of approximately 5 m estimated from the humeral length, NSM PV 24995 represents one of the largest specimens of early Spathian marine reptiles known to date. Such size variation among the earliest ichthyopterygians might suggest an explosive diversification in size immediately after the end-Permian mass extinction. Both vertebrae and humerus specimens exhibit an extremely cancellous inner structure, suggesting a high degree of aquatic adaptation in ichthyopterygians, despite their short history of evolution in the ocean.

Novel findings of late Cenomanian-Turonian Pachyophiid snakes, fishes and plants in the SE Bosnia-Herzegovina

New fossils from the Late Cenomanian-Turonian locality in the Bileca area are reported. The fossils include a diverse assemblage of snakelike reptile remains, fishes and plants. While fossil snakes have already been found in this area, the remains of fish and plants are reported for the first time. Based on the state of fossil preservation, the following were identified: the snake Pachyophis (Simoliophiidae), the pychnodont fish Coelodus sp., and plant remains of the flowering plants groups Magnoliidae exc. Lilianae, Arecaceae (palm trees), and Cycadopsida.

Identification of a large anguimorph lizard (Reptilia, Squamata) by an articulated hindlimb from the upper Maastrichtian (Upper Cretaceous) of Basturs-1 (Lleida, Spain)

Appendicular remains of squamate reptiles are barely described in the fossil record due to their low preservational potential and generally poor diagnostic information. Not many squamate fossil individuals preserve appendicular bony elements, these being mainly restricted to the rare articulated specimens found in a limited number of localities with specific conditions that favor exceptional preservation. Detailed descriptions of these bones, especially tarsals and metatarsals, are thus scarce in the literature due to the lesser relevance given to these elements in most anatomical descriptions. In this study we analyze an unpublished fossil specimen from the Maastrichtian of Basturs-1 (Lleida, Catalonia, Spain) corresponding to several articulated appendicular pes bones of a possible member of Varaniformes. We also provide detailed insights on the anatomy of the tarsalia and metatarsalia, particularly in anguimorphs. The fossil specimen here described, with an estimated snout-vent length (SVL) of ∼581 mm, reveals the putative varaniform from Basturs-1 as one of the largest Mesozoic terrestrial lizards, and possibly the largest from the European fossil record. Previous observations of an association between large lizards and dinosaur nesting sites are further supported by the find of this giant form in a locality known for the presence of numerous dinosaur eggs.

Fossil vertebrates from a unique marine bonebed of the Upper Cretaceous Smoky Hill Chalk, western Kansas, U.S.A.: new insights into the paleoecology of the Niobrara Formation

ABSTRACT The Smoky Hill Chalk Member of the Upper Cretaceous Niobrara Formation is a sedimentary deposit formed in the Western Interior Seaway of North America. Here, we examined a fossiliferous bonebed from an uncertain horizon within the Smoky Hill Chalk in Gove County, Kansas, U.S.A. Over 40,450 taxonomically identifiable specimens were collected, consisting of at least four chondrichthyans, 24 actinopterygians, and two tetrapods, including taxa such as Nanocorax, Squalicorax, Rhinobatos, pycnodontids, Lepisosteidae, Protosphyraena, Pachyrhizodus, ichthyodectiforms, plethodids, Albulidae, Apateodus, Cimolichthys, Enchodus, Acanthomorpha, Platecarpus, and Hesperornithiformes. The diversity of these vertebrates suggests the presence of a very complex ecosystem when the bonebed formed, which is determined to be mid- or late Santonian in age based on biostratigraphic, geographic, and geologic evidence. Based on published literature and this new study, at least 117 vertebrate taxa are now known from the Smoky Hill Chalk of Kansas, comprising 18 chondrichthyans, 63 actinopterygians, and 36 tetrapods. Comparisons of faunal data from the bonebed with databases on vertebrate fossils from the Niobrara Formation of Kansas in seven major museum collections indicate that the Niobrara fossil record based on museum collections is biased towards large reptilian remains compared with fish fossils. In particular, this study indicates that Enchodus spp. were especially common when the bonebed formed. A major corollary from this study is that a more accurate reconstruction of the paleoecology of the Western Interior Seaway requires the need for more analyses on smaller vertebrate taxa.

Late Cretaceous marine reptiles from Malyy Prolom in Ryazan Oblast, Central Russia

During the Cenomanian–Turonian transition (~94 Ma), what is today Central Russia formed part of the northern epicontinental margin of the Tethys Ocean. Diverse marine vertebrate faunas inhabited these palaeoenvironments, but their fossils are incompletely documented. Here, we report the discovery of marine reptile remains, recovered together with pterosaur, chondrichthyan, and actinopterygian fish material from a basal-most glauconitic sand and gravel layer of the Dmitrov Formation. These strata are exposed in an active quarry near the village of Malyy Prolom in the Shatsky District of Ryazan Oblast, Central Russia. The Dmitrov Formation deposits are middle–upper Santonian, but unconformably contact the underlying lower–middle Cenomanian Yakhroma Formation via a condensed boundary horizon that contains the vertebrate fossils with bivalve shell fragments and siliceous and phosphatic clasts. Such sedimentary characteristics indicate a high-energy shoreface setting where the vertebrate teeth and bones were likely reworked during cyclical regressions commencing in the latest Cenomanian–early Turonian. Time-averaging is also evidenced by the mixed occurrences of brachauchenine pliosaurids, elasmosaurid and polycotylid plesiosauroids, ophthalmosaurid ichthyosaurians similar to Pervushovisaurus, and a possible yaguarasaurine mosasauroid. These taxa are typical of Cenomanian–Turonian assemblages from across the northern peri-Tethys, and represent components of what were probably palaeobiogeographically widespread marine reptile faunas.

Late Paleogene herpetofaunas from the crossroads between two continents – new amphibian and reptile remains from the Oligocene of southern Balkans and Anatolia

We describe here new amphibian and reptile remains from three Oligocene localities of Turkey. Two of the localities (Kavakdere and Kocayarma) are situated in southeastern Europe and the other one (Kargi 2) in Anatolia, both areas where Oligocene herpetofauna is practically almost unknown. The material consists of albanerpetontids, pelobatid anurans, turtles, crocodylians, lacertids, scinciformatans, anguines, and “tropidophiids”. Albanerpetontids are for the first time identified in southeastern Europe, with the material being reminiscent of the younger species Albanerpeton inexpectatum Estes & Hoffstetter, 1976; the material potentially represents the oldest record of that species. Pelobatids, scinciformatans, and “tropidophiids” represent the oldest occurrences of these clades in the northeastern Mediterranean. The anguine genus Ophisaurus Daudin, 1803 is identified for the first time in the Paleogene of Eastern Europe. The “tropidophiids” are referred to two genera, Falseryx Szyndlar & Rage, 2003 and tentatively also to Platyspondylia Rage, 1974, with the latter having been so far exclusively known from western and central Europe. The role of a potential southern dispersal route of taxa among Asia and Europe, involving the area of southern Balkans and Anatolia, similarly to what has been recently demonstrated for mammals, is highlighted also for amphibians and reptiles.

The amphibians and reptiles from the Early Pleistocene of Coste San Giacomo (Anagni Basin, Italy)

ABSTRACT The site of Coste San Giacomo, in the Anagni Basin, is renowned for its mammal fauna of Early Pleistocene (Gelasian) age, including 32 taxa of small and large mammals. Adding to these, a small amount of remains of amphibians and reptiles was also recovered during recent excavations focused on collecting small vertebrates as well. The analysis of these remains resulted in the identification of a total of at least six different taxa: three anurans (Bufonidae indet., Pelophylax sp., and Rana sp.) and three reptiles (Lacertidae indet., Pseudopus sp., and Natrix sp.). This assemblage is comprised of taxa that are common in the Italian Quaternary, but also presents the first occurrence of the large anguid Pseudopus from the Apennine Peninsula in this period. On the whole, the herpetofauna indicates the presence of humid conditions in CSG at the moment of deposition of the bone bed that yielded the vertebrate remains.

Rare and Endemic Elements in Triassic Tetrapod Assemblages of Obshchii Syrt Highland (Eastern Europe)

The historical sequence of Early Triassic tetrapod communities in the Obshchii Syrt area includes eight consecutive faunal assemblages: this degree of detail in the identification of evolutionary stages is unique on a global scale. Temnospondyl amphibians are the dominant elements in these tetrapod assemblages, with three capitosauromorph groups (capitosaurids, benthosuchids, and trematosaurids) characterized by the widest distribution. Reptile remains are much less frequent. The paper provides a brief overview of the findings of rare and endemic tetrapod groups known from the Lower Triassic of Obshchii Syrt. These include amphibians (qantasids, lonchorhynchids, selenocarine capitosaurids, rhytidosteids, plagiosaurs, some forms of benthosuchids and trematosaurids, and chroniosuchians, the relict anthracosaurs) and reptiles (eosauropterygians, theriodonts, individual procolophonid, protorosaurian, and thecodontian genera). The peculiarity of the systematic composition of the Early Triassic tetrapod communities of Obshchii Syrt and Southern Cis-Urals with respect to that of the coeval fauna of terrestrial vertebrates of the more northern regions of Eastern Europe is emphasized, as is the legitimacy of considering these two areas as a specific region characterized by the distinctive character of their zoogeographical connections.

A small theropod dinosaur from the Lower Jurassic Lias Group of Charmouth, Dorset

Terrestrial reptile remains are rare in Early Jurassic deposits from the UK but include important records of early dinosaurs and pterosaurs. Here we report an isolated limb bone from the Charmouth Mudstone Formation (upper Sinemurian) of Charmouth, Dorset that was previously misidentified as a referred specimen of the pterosaur Dimorphodon macronyx. Comparisons with other Early Jurassic tetrapods indicate that this element is the fibula of a neotheropod dinosaur, representing one of only a handful of theropod remains from the Lias Group. Although the material is limited the fibula is distinct from other penecontemporaneous European theropods and might represent a new – though currently undiagnosable – taxon. This, alongside other isolated discoveries, indicates that there might have been a higher diversity of theropod taxa in the Early Jurassic of Europe than is generally appreciated.

The first tetrapod track recorded from the Rhaetian in the British Isles

Terrestrial vertebrate trace fossils are relatively abundant in mid-to-late Triassic and early Jurassic deposits in the British Isles but to date none at all have been recorded from the Rhaetian, the final stage of the Triassic. This represents a persistent gap in the terrestrial ichnological record. We present the first Rhaetian track to be recognised in the British Isles, found at Aust Cliff on the south bank of the Severn Estuary near Bristol in SW England. This locality is well known for disarticulated remains of Rhaetian fossil reptiles including some terrestrial species but in 2006 a track (TECMAG0161) was found for the fist time. Although the specimen was found ex-situ the palynological data from the surrounding matrix confirms a Rhaetian age. The track was examined with CT scanning and photogrammetry. We tentatively assign the track to the ichnogenus Procolophonichnium based on size and digit proportions. The isolated nature of the specimen offers little concrete information about the track maker but such tracks have previously been attributed to parareptiles or therapsid trackmakers. The specimen adds a datapoint to an otherwise ichnologically empty period of time in the British Isles. The track also provides solid evidence for a [locally] terrestrial environment in a sequence that is otherwise considered predominantly marine or estuarine. This discovery suggests that there may be more such tetrapod tracks of Rhaetian age preserved, at least at Aust, and further searching will hopefully lead to the current minimal dataset being expanded.

Tanystropheus and other archosauromorph reptile remains from the Middle and Late Triassic of Villány (Villány Hills, Hungary)

Tanystropheus and other possible archosauromorph fossils have been discovered from Middle to Upper Triassic shallow marine sedimentary sections in Villany (Villany Hills, southern Hungary). Four fragmentary cervical vertebrae can be assigned to Tanystropheus sp. based on characteristic features including the strongly elongate and hollow vertebral body with extremely reduced neural spine. Besides the cervicals, various teeth, classified into four different morphotypes including longitudinally striated, carinated and ziphodont ones, are thought to belong to archosauromorphs, since they markedly differ from the frequently found teeth of fish and sauropterygians. In addition, three enigmatic cranial bones that might represent some circumorbital elements, have been found as well, and are referred to here as Sauropsida indet. These fossils, originated from the same tectonic unit as those from the Anisian of Bihor (Romania), are of great importance for a better understanding of the poorly known semi-aquatic to terrestrial vertebrate fauna of the Middle to Late Triassic of central Europe.

Le dauphin à dents de requin Squalodon (Cetacea: Odontoceti) du remarquable assemblage de vertébrés marins de Montagna della Majella (Formation de Bolognano, Italie centrale).

The extinct family Squalodontidae consists of heterodont, medium-sized odontocetes, featuring a long rostrum that houses large, procumbent incisors and heavily ornamented postcanine teeth carrying accessory denticles, hence their vernacular name, dolphins. These longirostrine toothed whales are often seen as bridging the anatomical gap between archaic Oligocene odontocetes and their late Miocene to Holocene relatives. Possibly among the major marine predators of their time, the shark-toothed dolphins are important components of several lower Miocene marine-mammal assemblages from the North Atlantic and Mediterranean/Paratethysian realms. In the present work, a partial skull of Squalodontidae is described from the strata of the Bolognano Formation cropping out in the northeastern sector of the Montagna della Majella massif (Abruzzo, central Italy), which has previously yielded a rich lower Miocene marine-vertebrate assemblage, including eleven taxa of elasmobranchs as well as subordinate teleosts and very fragmentary remains of marine reptiles and mammals. The specimen consists of the anterodorsal portion of a rostrum, preserving parts of both premaxillae and left maxilla, and the anteriormost seven upper left teeth. This partial skull is here identified as belonging to the genus Squalodon, whose presence in the Montagna della Majella vertebrate assemblage had already been tentatively proposed on the basis of two fragmentary teeth. The paleontological significance of this find is discussed in the broader framework of the Euromediterranean record of Squalodon.