In the cableless seafloor observation networks (SONs), the links among network nodes rely on underwater acoustic communication (UAC). Due to the energy constraint and the high-reliability requirement of the cableless SONs, the noncoherent UAC has been a preferred choice, even though a noncoherent UAC scheme generally suffers from low spectral efficiency. In this paper, we propose a high-spectral-efficiency noncoherent UAC transmission scheme which is implemented as an orthogonal frequency-division multiplexing (OFDM) system adopting the on-off keying (OOK) modulation. To simultaneously achieve high performance at a low energy consumption, an irregular recursive convolutional code (IrCC) is employed and an accumulator (ACC) is introduced to achieve a modulation with memory at the transmitter side. The ACC enables a turbo iteration between the soft demapper called the ACC-OOK demapper and the soft decoder on the receiver side, and also reduces the decoding error floor. To account for the unknown signal-to-noise ratio (SNR), an iterative threshold estimation (ITE) algorithm is proposed to determine a proper decision threshold for the ACC-OOK demapper. The IrCC is designed to match the extrinsic information transfer (EXIT) curve of the ACC-OOK demapper, lowering the SNR threshold of the aforementioned turbo iteration. Simulations and experimental results verify the superiority of the proposed noncoherent UAC scheme over conventional ones.