The understorey of tropical rainforests is characterized by a low energetic level and a uniform climate : low light intensity (1 % or less of the incident radiation reaches ground level), lack of wind, atmospheric humidity near saturation, and almost constant temperature. However, the many species of understorey plants display a great diversity of growth habits. This energy-poor environment implies economies in the elaboration of non assimilating organs such as roots and stems, whereas leaf blades tend to never overlap. Stem thickening is reduced, as cambial activity is low (in shrubby Dicots), or nil (in Ferns, Monocots, and herbaceous Dicots). This goes along with a low branching level and a single stem, either unique and perennial, or replaced by basal repetition from a collar-like zone. Contrary to what is usually said, orthotropic and plagiotropic stems are equally frequent in shaded environments. The present study deals with the arrangement of leaves on stems, the whole leafy surface of an individual plant composing its «assimilatory set ». In a tropical rainforest understorey plant, the form of this assimilatory set tends to remain almost the same throughout the life cycle of the individual, old leaves and stems being replaced by new ones. Field observations carried out all over the tropical World led the author to identify a limited number of elementary global forms of the assimilatory set of understorey plants. Some are simple geometric forms (discs, tori, cylinders, and cones) with an axial symmetry ; others (ovals and laminas) exhibit a helicoidal or a bilateral symmetry. Sheets and pendant elements are asymmetric forms. Twenty elementary global forms are described : stemless discs, elevated discs, cupola of discs, superposed discs, stemless sheaf tori, stemless crown tori, funnel shaped tori, ascending sheaf tori, elevated horizontal ovals, inclined ovals, horizontal creeping laminas, ascending oblique laminas, vertical appressed lami¬ nas, elevated horizontal laminas, superposed horizontal laminas, helices, cylinders, cones, pendant elements, and sheets. Each of these forms can be further divided, according to the variety of growth habits able to lead to the same global form. Exemples are given for each mode of elaboration of the various forms. The convergences in global forms between Ferns, Monocots, and Dicots are emphasized. The same global form can indeed be achieved by different growth habits. Reciprocally, the same growth habit can also lead to different global forms. The position of the sexual organs does not play any role in the elaboration of a global form, the latter corresponding to the way the plant takes the best possible advantage of its energy-poor environment. The preferential habitats of each elementary global forms are specified.
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