Pleistocene Foragers Research Articles

The foodplant fitness landscape of Wonderboom and possible implications for Middle Pleistocene foragers of the Magaliesberg, South Africa

With this contribution, we provide a species and edible part list for the foodplant population of the Wonderboom landscape at the eastern end of the Magaliesberg range in Gauteng, South Africa, as a current proxy to hypothesise about past foraging behaviours. The greater Magaliesberg region is an ancient, relatively stable ecotone between the Grassland and Savanna Biomes, with its roots in the plateau uplift at ~ 4 Ma. We present the foraging landscape (foraging-scape) in a three-tier model at intervals of ~ 12.5 km, ~ 35 km, and ~ 70 km to assess proportional increases in foodplant species and edible parts when the foraging range is increased. We demonstrate that foraging within a ~ 12.5 km radius from the site provides an exceptionally rich foodplant landscape today. Thus, if the foodplant fitness potential during the Middle Pleistocene was roughly similar, it is reasonable to hypothesise that the Wonderboom foragers may have had little reason—apart from perhaps famine or drought—to go beyond a daily range for collecting plant foods. The Wonderboom ~ 12.5 km radius surface area is broadly similar to the footprint of the Cradle of Humankind ~ 40–70 km to its southwest. Directly comparing their foodplant species inventories shows that currently, 185 more foodplants grow around Wonderboom than in the Cradle. Our results suggest that the ecotone locality of Wonderboom, between the Grassland Biome to the south and the Savanna Biome to the north, may have had adaptive advantages in terms of its foodplant foraging potential.

Pigs as Pets: Early Human Relations with the Sulawesi Warty Pig (Sus celebensis).

The Sulawesi warty pig (S. celebensis) is a wild and still-extant suid that is endemic to the Indonesian island of Sulawesi. It has long been theorised that S. celebensis was domesticated and/or deliberately introduced to other islands in Indonesia prior to the advent of the Neolithic farming transition in the region. Thus far, however, there has been no empirical support for this idea, nor have scientists critiqued the argument that S. celebensis was a pre-Neolithic domesticate in detail. Here, it is proposed that early foragers could have formed a relationship with S. celebensis that was similar in essence to the close association between Late Pleistocene foragers in Eurasia and the wild wolf ancestors of domestic dogs. That is, a longstanding practice of hunter-gatherers intensively socialising wild-caught S. celebensis piglets for adoption into human society as companion animals ('pets') may have altered the predator-prey dynamic, brought aspects of wild pig behaviour and reproduction under indirect human selection and control, and caused changes that differentiated human-associated pigs from their solely wild-living counterparts.

Finding Fairness: From Pleistocene Foragers to Contemporary Capitalists By Justin Jennings. Gainesville: University Press of Florida 2021. Pp. 332. $90. ISBN 9780813066745 (cloth).

<i>Finding Fairness: From Pleistocene Foragers to Contemporary Capitalists</i> By Justin Jennings. Gainesville: University Press of Florida 2021. Pp. 332. $90. ISBN 9780813066745 (cloth).

Moving on from Here: Suggestions for the Future of “Mobility Thinking” in Studies of Paleolithic Technologies

Beginning in the early 1990s, the theme of mobility became central to many studies of Paleolithic lithic technology. Models relating group movements to the design, production, and treatment of stone artifacts have been widely employed both to explain features of Paleolithic assemblages, and as a source of information about the ecology of Pleistocene foragers. However, many of the questions that “mobility thinking” was originally adopted to answer are no longer at the center of inquiry in Paleolithic research. To remain relevant to new research agendas, new approaches to mobility and lithic technology should be developed. In keeping with the theme of this special issue, this paper discusses three areas where relatively simple changes in theory and practice could help to integrate “mobility thinking” into emerging research questions of broad, cross-disciplinary relevance: these include territories and ranging patterns, social networks, and intra-group variation in mobility.

Human exploitation of carnivores in Pleistocene China: A case study of the faunal remains from Shuidonggou Locality 7

Hominin-animal interactions at Pleistocene archaeological sites have been the main subject of research during human evolution. However, in contrast to that of ungulates, little is known about the evidence of human processing of carnivore species in this time interval of China. In this paper, we present the first case report of an interactive relationship between carnivore species and hunter-gathers in Pleistocene China, based mainly on a detailed taphonomic study of the faunal remains from Shuidonggou Locality 7. It seems highly probable that hominins butchered some individuals of carnivores and roasted bone elements with adhering meat at the site; whereas it’s equally feasible that these animals were exploited by hominins mainly for their pelt or fur. In either way, evidence from the site indicates novel strategies in adaptive behaviors adopted by Late Pleistocene foragers in harsh environments of Northwest China.

An abundance of developmental anomalies and abnormalities in Pleistocene people

Diverse developmental abnormalities and anomalous features are evident in the Pleistocene Homo fossil record, varying from minor but rare dental, vertebral, and carpal variants to exceptional systemic disorders. There are currently 75 documented anomalies or abnormalities from 66 individuals, spanning the Pleistocene but primarily from the Late Pleistocene Middle and Upper Paleolithic with their more complete skeletal remains. The expected probabilities of finding these variants or developmental disorders vary from <5% to <0.0001%, based on either recent human incidences or relevant Pleistocene sample distributions. Given the modest sample sizes available for the skeletal or dental elements in question, especially if the samples are appropriately limited in time and geography, the cumulative multiplicative probability of finding these developmental changes is vanishingly small. These data raise questions regarding social survival abilities, differing mortuary treatments of the biologically unusual, the role of ubiquitous stress among these Pleistocene foragers, and their levels of consanguinity. No single factor sufficiently accounts for the elevated level of these developmental variants or the low probability of finding them in the available paleontological record.

Rockshelters and the impact of the Laacher See eruption on Late Pleistocene foragers

Predictive modelling has identified rockshelter sites to the north-east of the Laacher See volcano in western Germany. These will be excavated to investigate the impacts of volcanic eruption on Late Pleistocene foragers.

A fuzzy logic methodology for assessing the resilience of past communities to tephra fall: a Laacher See eruption 13,000 year BP case

While volcanologists are experienced in assessing present and past volcanism, and while archaeologists are experts in understanding past societies, the study of how ancient volcanic activity has impacted contemporaneous communities remains little systematised. We here present a fuzzy logic-based methodology for bringing together expert assessments in evaluating the vulnerability and, by extension, the resilience of a group of late Pleistocene foragers to the Laacher See eruption, a large explosive eruption that affected continental Europe 13,000 years ago. Based on attributes assessing human health, shelter, food supply, and water supply our analyses suggest community resilience falls from 1.0 to 0.75 – 0.8 under a 5 cm tephra fall, to 0.45 – 0.50 with a 10 cm fall, and to 0.17 – 0.2 with an 18 – 20 cm tephra fall. Our explicit assessment of different experts’ evaluation of the different attribute’s relative importance facilitates a rigour in formulating such impact scenarios. The assessment methodology is rapid and can then be matched against existing evidence or, importantly, be used to also assess contemporary communities’ potential for loss under different tephra fall conditions. The methodology can be readily transferred between case studies and, in principle, between hazards, and could contribute significantly to the design of realistic disaster scenarios, which in turn serve to build resilience in at-risk communities.

A possible Late Pleistocene forager site from the Karaburun Peninsula, western Turkey

The ‘Karaburun Archaeological Survey’ project aims to illuminate the lifeways of Late Pleistocene and Early Holocene foragers in western Anatolia. A recently discovered, lithic-rich site on the Karaburun Peninsula offers new insights into a currently undocumented period of western Anatolian prehistory.

Assessing the proposed pre-last glacial maximum human occupation of North America at Coats-Hines-Litchy, Tennessee, and other sites

Genomic studies indicate that the first Pleistocene foragers who entered North America diverged from ancestral populations in Beringia sometime after the Last Glacial Maximum (LGM); however, several archaeological sites in North America have been proposed to predate the LGM. We present the results of our excavation and analysis of one such site, Coats-Hines-Litchy, Tennessee, which show that this site is a paleontological locality containing a geofact assemblage that pre-dates the LGM. Other sites in North America that purportedly predate the LGM occur in geomorphic contexts that are also conducive to the formation of geofact assemblages. As such, we propose that the reported artifacts from these sites were created by natural processes. No sites in North America currently provide credible evidence of a pre-LGM occupation.

A Unique Assemblage of Engraved Plaquettes from Ein Qashish South, Jezreel Valley, Israel: Figurative and Non-Figurative Symbols of Late Pleistocene Hunters-Gatherers in the Levant.

Three engraved limestone plaquettes from the recently excavated Epipaleolithic open-air site Ein Qashish South in the Jezreel Valley, Israel comprise unique evidence for symbolic behavior of Late Pleistocene foragers in the Levant. The engravings, uncovered in Kebaran and Geometric Kebaran deposits (ca. 23ka and ca. 16.5ka BP), include the image of a bird—the first figurative representation known so far from a pre-Natufian Epipaleolithic—along with geometric motifs such as chevrons, crosshatchings and ladders. Some of the engravings closely resemble roughly contemporary European finds interpreted as "systems of notations" or "artificial memory systems"–records related to timing of seasonal resources and associated aggregation events of nomadic groups. Moreover, similarly looking signs and patterns are well known from the context of the local Natufian—a final Epipaleolithic culture of sedentary or semi-sedentary foragers who started practicing agriculture. The investigation of the engravings found in Ein Qashish South involves conceptualizations developed in studies of European and local parallels, a selection of ethnographic examples and preliminary microscopic observations of the plaquettes. This shows that the figurative and non-figurative images comprise a coherent assemblage of symbols that might have been applied in order to store, share and transmit information related to social and subsistence realms of mobile bands. It further suggests that the site functioned as a locality of groups' aggregation and indicates social complexity of pre-Natufian foragers in the Levant. While alterations in social and subsistence strategies can explain the varying frequency of image use characterizing different areas of the Late Pleistocene world—the apparent similarity in graphics and the mode of their application support the possibility that symbol-mediated behavior has a common and much earlier origin.

Facing climatic hazards: Paleolithic foragers and Neolithic farmers

Food security intertwined with the need to maintain biological survival of a demographically viable population is the basic long-term policy for all societies. This paper compares selected cases of successful and failed Paleolithic groups of hunter-gatherers, as recorded archaeologically across Eurasia and Africa, in responding to the impacts of abrupt climatic changes. The complex prehistory of Pleistocene foragers is briefly compared to strategies adopted by Neolithic farmers. The major difference between forager and farmer economies is not their social organization as much as the access options to alternative territories and food resources. Open Pleistocene landscapes across Eurasia and Africa allowed for movements of Paleolithic groups over large distances with or without adopting new exploitation techniques. Successes and failures that we measure in a chronological macro-scale left archaeological evidence (lithic assemblages, faunal remains, occasional flimsy dwellings, etc.) in various regions, recorded by systematic and comprehensive surveys and excavations. However, we miss the chronological micro-scale of most of the Paleolithic period that could inform us about extinctions. We can identify only survival stories explained as successful adaptations. When farming communities were established during the course of the Holocene, variable modes of social and economic interactions and group resilience evolved in order to secure survival in years of bad harvests. Interactions with foragers and herders, competition, raids, village abandonment, migration into others' territories, were among the optional strategies. Due to the difference in chronological scales between 2.6 Ma of the Paleolithic period and the 12 Ka of the Holocene (or Anthropocene), we can more easily recognize the role of abrupt climatic changes among prehistoric societies during the Terminal Pleistocene and the Holocene and evaluate the success and failure of both, hunter-gatherers and farmers.

Neandertals, early modern humans, and rodeo riders

In 1995 Berger and Trinkaus (J. Archaeol. Sci. 22, 841–852) proposed that the anatomical distribution of Neandertal trauma, with a predominance of upper body lesions, reflected close-quarter ambush hunting as dictated by the available Middle Paleolithic weaponry (the “Rodeo rider” hypothesis). The necessity for mobility among these Late Pleistocene foragers, as a factor possibly reducing the number of preserved lower limb injuries, was considered as an alternative explanation. The accumulating data on Upper Paleolithic injuries and Middle Paleolithic weaponry, considerations of differential skeletal susceptibility of minor trauma, and evidence of interhuman violence, plus the importance of mobility for Late Pleistocene human existence, suggest that hunting injuries may explain only part of the pattern. The purpose of this note is not to resolve to ultimate factors behind the anatomical distribution of traumatic lesions among the Neandertals (or early modern humans). It is 1) to emphasize that there are multiple probable contributing factors other than close-quarter ambush hunting due to the limitations of Middle Paleolithic weaponry, and 2) to open the discussion to alternative interpretations.

Afromontane foragers of the Late Pleistocene: Site formation, chronology and occupational pulsing at Melikane Rockshelter, Lesotho

This paper provides a preliminary chronostratigraphic and palaeoenvironmental framework for the Late Pleistocene archaeological sequence at Melikane Rockshelter in mountainous eastern Lesotho. Renewed excavations at Melikane form part of a larger project investigating marginal landscape use by Late Pleistocene foragers in southern Africa. Geoarchaeological work undertaken at the site supports in-field observations that Melikane experienced regular, often intensive, input of groundwater via fissures in the shelter's rear wall. This strong hydrogeological connection resulted in episodic disturbances of the sedimentary sequence, exacerbated by other processes such as bioturbation. Despite this taphonomic complexity, a robust chronology for Melikane has been developed, based on tightly cross-correlated accelerator mass spectrometry (AMS) 14C with acid-base-wet oxidation stepped-combustion (ABOx-SC) pretreatment and single-grain optically stimulated luminescence (OSL) dating. The results show that human occupation of Melikane was strongly pulsed, with episodes of Late Pleistocene occupation at ∼80, ∼60, ∼50, ∼46–38 and ∼24 ka. At least three additional occupational pulses occurred in the Holocene at ∼9 ka, ∼3 ka and in the second millennium AD, but these are dealt with only briefly in this paper. Implications of the Late Pleistocene pulsing for the colonisation of high elevations by early modern humans in Africa ahead of dispersals into challenging landscapes beyond the continent are discussed.

Carcass transport decisions in Homo antecessor subsistence strategies

Pleistocene foragers used several prey acquisition and processing strategies. These strategies and their associated decisions are elucidated by taphonomic studies that cover animal transport, modifications by different agents and archaeological remains. Interpretative models of archaeological sites are by necessity based on natural and experimental observations. Ethno-archaeological data shows that several factors influenced decisions about carcass transport from the kill site to the home site. These factors often have little archaeological visibility. Díez et al. (1999) has previously interpreted the general characteristics of the macro-mammal remains from Gran Dolina Level TD6-2 (Sierra de Atapuerca, Burgos, Spain) as the result of anthropic accumulation, in which the anatomical profiles appeared to be the result of selective transport based on the animals’ weight. Recent taphonomic analysis has shown that carcasses with different weights may be subject to similar transport strategies, suggesting that other factors influenced these choices. The hominins that occupied TD6-2 (the TD6-2 hominin group), at least sometimes, transported large carcasses to the cave in their entirety, implying participation by groups of individuals in hunting parties. These individuals delayed their consumption of large amounts of food, instead moving it to Gran Dolina, where it was shared with other group members. These decisions are evidence of social cooperation and food sharing amongst early European hominins.

People of the ancient rainforest: Late Pleistocene foragers at the Batadomba-lena rockshelter, Sri Lanka

Batadomba-lena, a rockshelter in the rainforest of southwestern Sri Lanka, has yielded some of the earliest evidence of Homo sapiens in South Asia. H. sapiens foragers were present at Batadomba-lena from ca. 36,000 cal BP to the terminal Pleistocene and Holocene. Human occupation was sporadic before the global Last Glacial Maximum (LGM). Batadomba-lena’s Late Pleistocene inhabitants foraged for a broad spectrum of plant and mainly arboreal animal resources (monkeys, squirrels and abundant rainforest snails), derived from a landscape that retained equatorial rainforest cover through periods of pronounced regional aridity during the LGM. Juxtaposed hearths, palaeofloors with habitation debris, postholes, excavated pits, and animal and plant remains, including abundant Canarium nutshells, reflect intensive habitation of the rockshelter in times of monsoon intensification and biome reorganisation after ca. 16,000 cal BP. This period corresponds with further broadening of the economic spectrum, evidenced though increased contribution of squirrels, freshwater snails and Canarium nuts in the diet of the rockshelter occupants. Microliths are more abundant and morphologically diverse in the earliest, pre-LGM layer and decline markedly during intensified rockshelter use on the wane of the LGM. We propose that changing toolkits and subsistence base reflect changing foraging practices, from shorter-lived visits of highly mobile foraging bands in the period before the LGM, to intensified use of Batadomba-lena and intense foraging for diverse resources around the site during and, especially, following the LGM. Traces of ochre, marine shell beads and other objects from an 80 km-distant shore, and, possibly burials reflect symbolic practices from the outset of human presence at the rockshelter. Evidence for differentiated use of space (individual hearths, possible habitation structures) is present in LGM and terminal Pleistocene layers. The record of Batadomba-lena demonstrates that Late Pleistocene pathways to (aspects of) behavioural ‘modernity’ (composite tools, practice of symbolism and ritual, broad spectrum economy) were diverse and ecologically contingent.

Civilizing Cooperation: Paul Seabright and the Company of Strangers

Paul Seabright is the first to clearly identify a major puzzle about human social evolution: the expansion of cooperation in the more complex societies of the Holocene. Identifying that problem is a major achievement, but in this paper I give a somewhat different account of the nature of the problem and a somewhat different account of the social world of Pleistocene foragers. So, we agree that there is a problem, but not on its nature or solution.

The first Africans: African archaeology from the earliest tool makers to most recent foragers

1. Introducing the African record 2. Frameworks in space and time 3. First tool users and makers 4. Early Pleistocene foragers 5. Mid-Pleistocene foragers 6. Transitions and origins 7. The Big Dry: the archaeology of marine isotope 4-2 8. Hunting, gathering, intensifying: the mid-Holocene record 9. Foragers in a world of farmers 10. The future of the first Africans' past.

LATE PLEISTOCENE AND EARLY HOLOCENE FORAGER ORGANIZATIONS IN SOUTHEAST ASIA] The Late Pleistocene to Early Holocene foragers of northern Luzon

LATE PLEISTOCENE AND EARLY HOLOCENE FORAGER ORGANIZATIONS IN SOUTHEAST ASIA] The Late Pleistocene to Early Holocene foragers of northern Luzon

LATE PLEISTOCENE AND EARLY HOLOCENE FORAGER ORGANIZATIONS IN SOUTHEAST ASIA] Hoabinhian macrobotanical remains from archaeological sites in Vietnam: Indicators of climate changes from the Late Pleistocene to the Early Holocene

LATE PLEISTOCENE AND EARLY HOLOCENE FORAGER ORGANIZATIONS IN SOUTHEAST ASIA] Hoabinhian macrobotanical remains from archaeological sites in Vietnam: Indicators of climate changes from the Late Pleistocene to the Early Holocene