Planktonic Foraminiferal Assemblages Research Articles

Environmental perturbations around the Badenian/Sarmatian (Middle Miocene) boundary in the Central Paratethys: Micropalaeontological and organic geochemical records

The boundary between the two regional Middle Miocene stages (the Badenian and Sarmatian) of the Central Paratethys is associated with the largest faunal turnover event in the Paratethys realm. A recent study performed in the northern Polish Carpathian Foredeep revealed that the Badenian/Sarmatian boundary is coeval with a major change in benthic and planktonic foraminiferal assemblages that occurs a few metres below the beginning of the Anomalinoides dividens Interval Zone in our section. The lowermost Sarmatian biozone is the Elphidium angulatum Partial Range Zone, and its lower boundary corresponds to the disappearance of almost all Badenian foraminiferal species. Detailed analysis of a fully cored borehole in southern Poland using micropalaeontological tools (foraminifera and palynology) and a range of (inorganic and organic) geochemical methods indicated that during the late Badenian and almost the entire profile of Anomalinoides dividens Zone, oxic or suboxic sedimentary conditions prevailed. In the Elphidium angulatum Zone and the lowest part of the subsequent Anomalinoides dividens Zone, suboxic and anoxic conditions dominated the sea bottom. This research indicated that the sudden disappearance of marine Badenian species in the Polish Carpathian Foredeep was caused by a rapid change in basin chemistry although not necessarily it implies open marine conditions changing to brackish water environments.

Geochemical and Biostratigraphical Evaluation of Lokpanta Shale in Abakaliki Anticlinorium, Anambra Basin Province, Southeast Nigeria

A geochemical and biostratigraphic method has been applied to explore the quality of organic matter, thermal evolution, potential hydrocarbon source and type of fossils of Lokpanta Shale from the Abakaliki Anticlinorium, Anambra Basin Province, Southeast Nigeria. Data obtained reveal that the total organic carbon (TOC) (2.18wt%) of the shale constitutes that of good to excellent source rock with gas-prone kerogen indicated by Rock-Eval S2/S3 (2.32). The low oxygen index (OI) (27.20 mgCO2g-1TOC) suggest deposition under low energy environments. The plots of HI against Tmax classified the organic matter as Type III kerogen. The poor concentration of OM is thought to account for its current hydrogen index (31.47 mgHCg-1TOC). The predominance of Type III kerogen in the Lokpanta shale suggests their potential to generate gas in the deeply buried sections. The Tmax values from the pyrolysis of the shales of the Lokpanta ranges from 428 to 442oC corresponding to maturity levels within the oil formation. The lamination of the sediments of the Lokpanta area with no burrows and the occurrence of dominantly planktonic foraminiferal assemblages also indicate a quiet water anoxic marine condition.

Biostratigraphy of Shiranish Formation from Selected Wells, Central of Iraq

The Quantitative high-resolution planktonic foraminiferal analysis of the subsurface section in three selected wells in the Ajeel Oil Field (Aj-8, Aj-12, and Aj-15) in Tikrit Governorate, Central Iraq has revealed that Shiranish Formation deposited in Late Campanian- Latest Maastrichtian age. This formation consists mainly of marly and marly limestone yielding diverse planktonic foraminiferal assemblages and calcareous benthic foraminifera, with a total of 46 species that belong to 23 genera, Three zones and four subzones, which cover the Late Campanian to the Latest Maastrichtian, were identified based on the recorded planktonic foraminifera and their ranges. They are as follows:1. Globotruncana aegyptiaca Zone that dated to be Late Campanian 2. Gansserina gansseri Zone that refers to Early Maastrichtian 3. Abathomphalus mayaroensis Zone that indicates Late Maastrichtian, in this section, the Abathomphalus mayaroensis Zone is divided into four subzones, namely in that order from bottom to top: Racemiguembelina fructicosa (Early Late Maastrichtian), Pseudoguembelina hariaensis (Middle-Late Maastrichtian), P. palpebra ( Late Late Maastrichtian), and Plummerita hantkeninoides (latest Maastrichtian).

Biostratigraphy of the Early Turonian- Late Campanian pelagic limestone (Kometan and Gulneri formations) near Sayid Sadyq Town, Kurdistan Region, NE-Iraq.

This study aims at the presentation of a planktonic foraminiferal biozonation of the Late Cretaceous pelagic carbonate section in the east of Sulaymaniyah governorate Kurdistan region, Northeast of Iraq. The zonation proved the occurrence of the Kometan and Gulneri formations which belong to the Early Turonian – Late Campanian age. This is achieved by taking fifty-two (52) samples from the lower part of the section to the upper one with an interval of two meters between successive samples. From these samples, sixty-two (62) thin sections are prepared and (43) species from (11) genera of planktonic foraminifera are indicated and identified in the two formations of the studied section. According to the First and Last occurrence of identified planktonic foraminiferal assemblage, the studied section is divided into seven biozones, from older to young. These biozones are Whitenella archaeocretacea-Heterohelix moremani Total Range Zone, Marginotruncana sigali Partial Range Zone, Dicarinella concavata Interval Range Zone, Dicarinella asymetrica Total Range Zone, Globotruncanita elevata Partial Range Zone, Globotruncana ventricosa Interval Range Zone, Radotruncana calcarata Total Range Zone. The age of the two Formations is assigned as Early Turonian- Late Campanian based on the planktonic foraminiferal ranges from the lower part of the section to the upper part. The boundary of the Gulneri and Balambo formations is gradation while there is a gap between the former formation and Kometan Formation.

Towards an Integrated and Accurate Planktonic-Foraminiferal-Deduced Bio-Chrono-Stratigraphic Framework of Late Quaternary Mediterranean Marine Cores

The late Quaternary is a key stratigraphic interval as it encompasses the Late Glacial to Holocene transition, which is characterized by a series of pronounced centennial climate oscillations and subsequent short-term events of paleoceanographic variability. Across this stratigraphic interval, significant turnovers and faunal changes in the composition and abundance of planktonic foraminiferal assemblages are well-documented through their high-resolution quantitative analysis performed in the south Aegean core NS-18. The identification of 10 synchronous bioevents among the Mediterranean sub-basins allows accurate inter-basinal correlations over the study time interval, thus contributing to the development of a robust chronostratigraphic framework for this setting. Moreover, the identification and timing of additional 20 diachronous bioevents, in conjunction with the already established bio-ecozonation scheme of the Aegean Sea, provide a continuous record of faunal changes (in terms of species-specific distributional abundances) which can be used as an additional locally expressed biochronological tool for the eastern Mediterranean deep-sea cores. The present study certainly indicates that the cause-and-effect relationships between the paleoceanographic/paleoclimatic perturbations and biological response require a highly resolved regional chronostratigraphy.

British–Irish Ice Sheet and polar front history of the Goban Spur, offshore southwest Ireland over the last 250 000 years

Deep Sea Drilling Program (DSDP) Site 548 was cored in 1984 at a water depth of 1256 m on the Goban Spur, offshore southwest Ireland. Coring retrieved a ~100‐m‐thick Pleistocene contourite sequence. This study uses planktonic foraminiferal assemblage and benthic foraminiferal oxygen isotope analyses to establish an age model for the upper 40 m of this core. This site's multidisciplinary analyses of planktonic foraminiferal assemblages, lithic grains, facies and calcium carbonate concentration reveal a 250 000‐year record of the North Atlantic polar front variability and British–Irish Ice Sheet (BIIS) history. The sequence is characterized by alternations of ice rafted debris (IRD) laden pelagic mud facies with calcium carbonate‐rich silty sand contourite facies that track glacial/interglacial cycles. The polar front migrated southward across the area several times during glacial maxima and stadial periods, while warmer Mediterranean Outflow Water (MOW) flowed northward across the region during interglacial and interstadial periods depositing contourites. Lithic analyses reveal a complex history of IRD deposition associated with iceberg calving from the Laurentide Ice Sheet and northwest European ice sheets, mainly the BIIS. Comparison between the Goban Spur (DSDP Site 548) and the Celtic Margin (MD03‐2692) and central North Atlantic Integrated Ocean Drilling Program (IODP) Site U1308 suggests differences between the ‘non‐Laurentide Ice Sheet’ Heinrich Events (HE) 6 and 3 at the Goban Spur, with IRD from the BIIS being prominent during HE 6 and IRD from other European ice sheets north of the BIIS likely being more dominant during HE 3. The nature of lithics in IRD‐rich horizons during Terminations 3, 3A, 2 and 1 suggests significant iceberg calving episodes preceding BIIS retreat during the onset of interstadial intervals.

Dynamics of the subpolar gyre and transition zone of the North Atlantic during the last glacial cycle

Past studies of marine sediments of the North Atlantic documented millennial-scale sea-surface cooling related to the Dansgaard-Oeschger events that influenced the oceanic fronts and surface hydrology during the last glacial cycle. Most studies have focused on using ice-rafting, single or multiple species of planktonic foraminifers' abundances, and geochemistry to assess environmental changes. However, detailed planktonic foraminiferal assemblage data in the North Atlantic Transition Zone (NATZ) and subpolar gyre (SPG) for the full last glacial cycle are scarce. Here we report planktonic foraminiferal abundances, oxygen isotopes, and ice-rafted detritus (IRD) counts from two sediment cores, Hu9007-08 and Hu71022-377, collected from the Milne seamount of southwestern SPG (SW SPG) and NATZ, respectively. Further, we have used the published planktonic foraminiferal assemblage, IRD, and isotopic data from the eastern and western SPG to provide an integrated reconstruction of changes in the surface water masses and dynamics of the Polar, Arctic, and subarctic fronts (PF, AF, and SAF, respectively). Despite the overall warmth of the SW SPG (near the Newfoundland Basin) compared to the eastern SPG during marine isotope stage (MIS) 5, the North Atlantic-wide cooling events C24 and C21 were still identified in the SW SPG. Further, the planktonic foraminiferal abundances in cores EW9303-37JPC (i.e., scarce Neogloboquadrina pachyderma and abundant Globigerinoides inflata) and Hu9007-08 reveal a relatively warm MIS 5e compared to the Holocene. However, the decrease in the abundance of G. inflata suggests that the SW SPG was influenced by the seasonal presence of subpolar waters and possibly a southward shift in the oceanic frontal systems between ∼126 and ∼120 ka. During the Heinrich ice-rafting events, when meltwater flooded the surface North Atlantic, the PF, AF, and SAF shifted southward, and the polar/subpolar water masses briefly invaded the western and eastern SPG and NATZ. In the early part of MIS 3 (60–40 ka), the oceanic frontal system shifted northward from its position during MIS 4. The SPG and NATZ gradually warmed, and the subpolar species Globigerina bulloides in cores Hu9007-08, Hu71022-377, and CH69-K09 increased in the middle of MIS 3 (i.e., 38–30 ka), suggesting continuous sea surface warming while the oceanic frontal systems continued to move northward. This hydrological environment rapidly cooled in the late MIS 3-MIS 2 (30–19 ka), with the PF shifted southward to the mid-latitude (∼45°N) during the Last Glacial Maximum. During the last deglaciation-Holocene, the regions of cores Hu9007-08 and Hu71022-377 were mainly influenced by the warm North Atlantic Current, and the SAF was located north of both cores. This study presents the first large-scale water mass movements and frontal changes in the SPG and NATZ to better understand the dynamics of the surface circulation in the North Atlantic during the last glacial cycle.

Surface and thermocline ocean circulation intensity changes in the western Arabian Sea during ∼172 kyr

The relationship between monsoon-induced productivity and ocean circulation over millennial timescales is crucial. However, a study regarding the circulation of the western Arabian Sea (WAS) needs to be conducted. Understanding the seasonal, surface, and thermocline ocean circulation throughout the glacial-Interglacial period in the WAS is also elusive and essential. The long distant Antarctic Intermediate Water (AAIW) flow, and Sub-antarctic Mode Water (SAMW) penetration to the north side of the equator in the Indian Ocean needs to be understood and underlined using the micropaleontological (planktonic foraminiferal assemblages) and geochemical analysis such as oxygen isotope of Globigerinoides ruber and Neogloboquadrina dutertrei (δ18OG.ruber and δ18ON.dutertrei) in this study. Furthermore, understanding the variations in paleoproductivity, upper oceanic water column stratification, surface mixing, and thermocline layer variability in the WAS is also helpful. According to the results, the variation in the upwelling intensity during Glacial-Interglacial and monsoon variation suggest the upper water stratification/vertical mixing in the WAS. Here, we provide new evidence of poorly ventilated water mass in our region. Our results indicate that enhanced northward expansion of AAIW coincides with the weakening of summer monsoon during glacial periods except mid-MIS4, reflected by isotopic study and assemblages. In this context, depleted δ18ON. dutertrei value and high concentration of Globorotalia menardii (G. menardii) were due to lack/absence of AAIW ventilation and SAMW; therefore, warming occurred in the mid-MIS4.

Links between precipitation patterns over eastern tropical South America and productivity in the western tropical South Atlantic Ocean during the last deglacial

The possibility of a collapse (or marked weakening) of the Atlantic Meridional Overturning Circulation (AMOC) within a few centuries boosted the interest of the scientific community on the responses of different compartments of the climate system to past AMOC collapses (or marked weakening). The most recent examples of such periods are Heinrich Stadial 1 (HS1,18-15 ka BP; near-collapsed AMOC) and the Younger Dryas (YD, 12.9-11.7 ka BP; markedly weakened AMOC). Here, we present data from elemental ratios, spectral reflectance, organic biomarkers, sea surface temperatures and planktonic foraminiferal assemblage from two marine sediment cores recovered from the western tropical South Atlantic Ocean spanning the last ca. 20 kyr. The recovered sediments record the responses of eastern tropical South American hydroclimate, western tropical South Atlantic productivity and sea surface temperatures to changes in AMOC strength associated with HS1 and the YD. Our data show that both periods were characterized by wetter conditions over eastern tropical South America associated with increased continental runoff compared to the Last Glacial Maximum. High sea surface temperatures during HS1 and the YD are in line with marked AMOC decreases that led to heat accumulation in the surface layer of the western South Atlantic Ocean, facilitating increased precipitation over eastern tropical South America. Our data demonstrate for the first time that intense runoff during HS1, and hence nutrient transfer to the ocean, affected the pelagic ecosystem, likely increasing primary productivity in the otherwise oligotrophic western tropical South Atlantic Ocean. Our study demonstrates a tight coupling between marine and terrestrial processes in the eastern South American realm during periods of strong climate instability.

Micropaleontological and Isotopic Perspective of Surface Water Mass Variability in the NE Atlantic from MIS 6 to 5e (188 to 115 ka)

The penultimate glaciation (marine isotope stage (MIS) 6) is considered regionally extreme compared to the last glacial maximum, in which the European ice sheets had a vast areal extent. In contrast to the last deglaciation (19–7 ka), the penultimate deglaciation (140–130 ka) hosts one of the most rapid oceanographic changes of the late Pleistocene. In this study, we reconstructed changes in the near-surface and thermocline in the central to northeast Atlantic by analyzing sediments from two Integrated Ocean Drilling Program Expedition 306 sites. Sites U1313 (41°00.6′ N, 32°57.4′ W) and U1314 (56°21.9′ N, 27°53.3′ W) were drilled on the eastern flank of the mid-Atlantic ridge and Gardar Drift of the eastern subpolar North Atlantic, respectively. We analyzed planktonic foraminiferal assemblages, ice-rafted debris (IRD), and oxygen isotopes in two planktonic foraminifers, Globigerina bulloides, and Globorotalia inflata, from MIS 6 to 5e (185–115 ka). Warmer and colder sea-surface conditions were marked by a change in the relative abundance of polar, subpolar, and transitional planktonic foraminifers. Oxygen isotopes in G. bulloides and G. inflata suggest that the thermocline deepened at the subtropical Site U1313 during MIS 6. The lack of Globorotalia inflata prevented us from profiling the mixed layer and thermocline at the subpolar Site U1314. In contrast to MIS 6, the mixed layer and thermocline were re-stratified during the last interglacial. The lack of major IRD events at both sites suggests the stability of the Laurentide ice sheet during MIS 6 compared to the subsequent glaciation. The presence of Heinrich event 11 indicates the discharge of freshwater that freshened the sea surface, resulting in mixing between the mixed layer and thermocline. Our results were placed into a broader context using published data that shed light on the sensitivity of freshwater discharge to the North Atlantic and the following changes with a transition from a penultimate glacial to an interglacial period in surface circulation.

Planktonic Foraminiferal Biostratigraphy of the Kometan Formation from Kurdistan Region, NE Iraq

The Kometan Formation is widely spread in northeastern Iraq and is mostly composed of deep marine carbonates, with a predominance of planktonic foraminiferal assemblages throughout its sequences. Two outcrop sections, Smaquli and Tabin, were chosen to study the biostratigraphy of the Kometan Formation. The study identified 32 planktonic foraminiferal species belonging to 12 genera. According to stratigraphic distribution and the relative distribution of planktonic foraminifera assemblages, the study demonstrated the possibility of subdividing the sequence of the Kometan Formation into five foraminiferal biozones, including: Marginotruncana schneegansi Partial Range Zone, Dicarinella primitiva Interval Zone, Dicarinella concavata Interval Zone, Dicarinella asymetrica Total Range Zone, Globotruncanita elevata Partial Range Zone. The Planktonic zones were correlated with other zonal schemes inside and outside Iraq. They are considered to be extending from Late Turonian to Early Campanian.

Late Miocene biostratigraphy and paleoenvironment of the Sais Basin (southern Rifian Corridor, Morocco): New insights from the Moulay Yakoub area

During the Late Miocene, the Rifian Corridor was one of the two main corridors allowing exchanges between the Atlantic and the Mediterranean Seas. Sedimentary records accumulated along this corridor and specifically, in the Sais Basin, constitute case-study material for understanding the impact of the South Rifian Corridor on the evolution of the fauna, phytoplankton and environment during the Late Miocene. In order to reconstruct the evolution of marine paleoenvironments in the Moulay Yakoub region and to compare this evolution with other areas of the Sais basin, we conducted an integrated biostratigraphic and palynological study of the Upper Miocene of borehole KA 102. High-resolution biostratigraphy based on planktonic foraminifera revised and defined the chronostratigraphic framework of the Upper Miocene sedimentary series of the Moulay Yakoub region providing an age of 7.8 Ma to 6.52 Ma (Late Tortonian to Early Messinian). In the Late Tortonian, the planktonic foraminiferal assemblages suggest an open and relatively deep marine environment. At the Tortonian/Messinian boundary, planktonic foraminiferal and dinoflagellate associations, as well as distal indices, suggest a relatively deep and neritic distal marine environment with a decrease in diversity and an increase in reworked taxa. Surface waters are warm with a minor cooling trend observed just below this boundary. In comparison to previously studied sections in the Sais Basin, the Lower Messinian is characterized by a deeper and more distal marine environment. However, proximal trends and warm surface water conditions persisted in all studied areas from the basin during this period.

Planktonic foraminiferal assemblages reflect warming during two recent mid-latitude marine heatwaves

Under future climate scenarios, ocean temperatures that are presently extreme and qualify as marine heatwaves (MHW) are forecasted to increase in frequency and intensity, but little is known about the impact of these events on one of the most common paleoproxies, planktonic foraminifera. Planktonic foraminifera are globally ubiquitous, shelled marine protists. Their abundances and geochemistry vary with ocean conditions and fossil specimens are commonly used to reconstruct ancient ocean conditions. Planktonic foraminiferal assemblages are known to vary globally with sea surface temperature, primary productivity, and other hydrographic conditions, but have not been studied in the context of mid-latitude MHWs. For this study, the community composition and abundance of planktonic foraminifera were quantified for 2010-2019 along the Newport Hydrographic Line, a long-term monitoring transect at 44.6°N in the Northern California Current (NCC). Samples were obtained from archived plankton tows spanning 46 to 370 km offshore during annual autumn (August – October) cruises. Two MHWs impacted the region during this timeframe: the first during 2014-2016 and a second, shorter duration MHW in 2019. During the 2014-2016 MHW, warm water subtropical and tropical foraminifera species were more prevalent than the typical polar, subpolar, and transitional species common to this region. Cold water species were abundant again after the first MHW dissipated in late 2016. During the second, shorter-duration MHW in 2019, the assemblage consisted of a warm water assemblage but did not include tropical species. The foraminiferal assemblage variability correlated with changes in temperature and salinity in the upper 100 meters and was not correlated with distance offshore or upwelling. These results suggest that fossil foraminiferal assemblages from deep sea sediment cores may provide insight into the magnitude and frequency of past MHWs.

Planktonic foraminiferal biostratigraphy of the upper Turonian-lower Santonian succession in the Lorestan Zone of the Zagros sedimentary Basin, SW Iran

The Surgah Formation was studied in four sections (Tang-e-Shirin Ab, Kuh-e-Surgah Holostem Valley, and Sarkan) from the NW-SE of the Zagros sedimentary Basin. Forty-one planktonic foraminiferal species belonging to eighteen genera were recognized. These species were used to identify three zones across the Surgah Formation from the bottom to the top: Dicarinella primitiva – Marginotruncana sigali Zone, Dicarinella concavata Zone, and Dicarinella asymetrica Zone. Based on these data, the Surgah Formation was assigned to the Late Turonian-Santonian age. The base of the Coniacian Stage projects within the lower part of the Dicarinella concavata Zone and is traced at the first occurrence (FO) of Pseudotextularia nuttalli. Owing to the absence of Pseudotextularia nuttalli at the Sarkan well, we utilized the extinction of Whiteinella aprica to estimate the base of the Coniacian Stage. The Coniacian/Santonian (C/S) boundary is placed within the basal part of the Dicarinella asymetrica Zone based on the FO of Globotruncana linneiana. The base of the Santonian Stage denotes marked radiation in the globotruncanids among the planktonic foraminiferal assemblage. Great lithofacies and thickness variations have been observed within the D. concavata Zone. These variations likely indicate that the shoreline was located northwest of the Lorestan province during the Coniacian. In contrast, the southwest extreme of the Lorestan province was characterized by deep-water settings.

Agulhas leakage extension and its influences on South Atlantic surface water hydrography during the Pleistocene

The inter-ocean transfer of warm and salty water through the Agulhas leakage is regarded as a significant factor influencing the South Atlantic's surface and intermediate water hydrography. To evaluate the extension of Agulhas leakage for the past 1.2 Ma, planktonic foraminiferal assemblage from Ocean Drilling Program Hole 1088B, within the vicinity of AL, and Hole 1090B, located south of the significant Agulhas retroflection region, were generated. The Hole 1088B shows higher abundance of tropical and sub-tropical assemblage. The combination of Globigerinoides ruber, Trilobatus sacculifer, Globigerinoides conglobatus and Globorotalia menardii has previously been defined as the Agulhas leakage fauna [ALF]. The pulse-like amplification of the ALF record at major glacial-interglacial transitions indicates these were periods of strengthened Agulhas leakage with subsequent shedding of Agulhas rings and filaments as far as ∼41°S. From ∼450 ka, the frontal zone exhibits strong ALF variations with evident increase at glacial-interglacial transitions of Marine Isotopic Stages (MIS) 12, 10, 8 and 6, suggesting vigorous eddy mixing and a shift in the leakage zone towards the subantarctic waters. The migration of the Subtropical Convergence and Neogloboquadrina pachyderma (s) coiling ratios during these glacial-interglacial transitions supports the southward expansion of Agulhas leakage. The ALF record is supported by the δ18O record of Globigerina bulloides from hole 1088B, which identify a major anomaly at MIS 10, with depleted δ18O values towards the glacial termination. Simultaneously, the results from Hole 1090B show restricted presence of tropical-subtropical assemblage with their appearance reported only at glacial-interglacial transitions of the MIS 16, 12, 10 and 8. The curtailed ALF index and dominance of the subpolar species N. pachyderma (s) indicate the termination of Agulhas leakage before reaching the Subantarctic zone. The ODP sites chosen for the study reveal the intensification of Agulhas leakage and southward shift of the major leakage zone into the subantarctic zone during the Pleistocene glacial-interglacial transitions.

Middle Eocene–early Miocene planktonic foraminiferal biostratigraphy, chronostratigraphy, sea‐level reconstruction and sequence biostratigraphy at N. ElFaras‐1Xwell, Qattara Depression, Western Desert, Egypt

The planktonic foraminiferal assemblages are tracked in the middle Eocene–early Miocene succession within N. El Faras‐1X Well, Qattara Depression, Egypt. This succession is composed mainly of three lithostratigraphic units the upper part of the Apollonia Formation (middle to late Eocene), Dabaa Formation (late Eocene and Oligocene) and the lower part of the Moghra Formation (early Miocene). Biostratigraphically, the recorded 77 planktonic foraminiferal species enabled the recognition of 12 planktonic foraminiferal biozones; 4 biozones belong to middle to late Eocene, 7 biozones belong to the Oligocene and 1 belongs to the early Miocene. The proposed biozones were compared with their equivalents in other Egyptian regions and other schemes. Eleven Lowest Occurrence and two Highest Occurrence planktonic foraminiferal bioevents were also recognized and correlated all over the world. The chronostratigraphy and the stage boundaries were mainly delineated according to the diagnostic planktonic foraminifera. The middle–late Eocene boundary (Bartonian–Priabonian boundary), The Eocene–Oligocene boundary (Priabonian–Rupelian boundary), the early Oligocene–late Oligocene boundary (Rupelian–Chattian boundary) and the Oligocene–Miocene boundary (Chattian–Aquitanian boundary) were accurately determined. Based on the % P, the palaeobathymetry was estimated and hence the sea‐level fluctuation. The reconstructed sea‐level curve revealed two major transgressive–regressive cycles with minor oscillation within them. These events were globally correlated and match in most parts with that of the worldwide sea‐level curves. The sequence stratigraphy framework consists mainly of two transgressive–regressive (T–R) depositional sequences (DS1 and DS2). The DS1 includes the TST1 below which is capped by mfs1 and the HST1 above. The DS2 involves the TST2 below that was capped by mfs2 and the HST2 above.

Nutrients and hydrography explain the composition of recent Mediterranean planktonic foraminiferal assemblages

The composition of planktonic foraminiferal assemblages in surface sediments of the Mediterranean Sea shows a pronounced zonal gradient. This gradient is different from what is observed in the open ocean, indicating that the assemblage composition may respond to environmental variables other than the otherwise dominant surface water temperature. Here we make use of a dataset of census counts of 18 taxonomic categories in 124 surface-sediment samples extracted from the ForCenS database to understand the main environmental factors affecting the composition of recent Mediterranean planktonic foraminiferal assemblages. The tested explanatory variables include temperature, salinity, chlorophyll, nitrate and phosphate concentrations in the surface waters, temperature, nitrate and phosphate concentrations at depth and thermal, salinity and density vertical gradients. The composition of the assemblages is aligned along two environmental gradients and redundancy analysis reveals that the tested variables explain a large portion (>70%) of the variance in the compositional data. The first environmental gradient reflects the nutrient content in the deep waters, affecting nutrient availability in the productive zone, while the second gradient is driven by higher (northern area) and lower (southern area) upward nutrient advection due to differences in density stratification, conditioned by the regional hydrography. Although sea surface temperature is the most reconstructed environmental variable from fossil assemblages of planktonic foraminifera, it seems to play a secondary role in the Mediterranean Sea. This, has implications for reconstructions of past oceanographic conditions in this region, opening up the possibility to reconstruct productivity instead, or next to, temperature.

Planktonic foraminiferal biostratigraphy of the Farrokhi Formation and determination of the Campanian/Maastrichtian boundary in Central Iran

The Cretaceous Farrokhi Formation is located in the Khur area of central Iran and yielded a reasonably well-preserved rich planktonic foraminiferal assemblage. This formation consists of 154 m of fossiliferous marl and limestone. The lower and upper bounding surfaces of the Farrokhi Formation are unconformable with the underlying Haftoman Formation and the overlying Chupanan Formation. In this research, 46 planktonic foraminiferal species belonging to 19 genera were identified and resulted in recognition of the following three biozones (from base to top); the Globotruncana aegyptiaca Interval Zone (IZ), the Gansserina gansseri IZ, and the Contusotruncana contusa IZ. This biozonal framework places deposition of the Farrokhi Formation during the late Campanian to early Maastrichtian. Biostratigraphic comparison between the samples for this research and data from three other studied sections within the Farrokhi Formation indicates that the basal and upper boundaries of the formation are time-transgressive and migrate chronologically from north to southeast. Since the first occurrences of Pseudoguembelina palpebra and Rugoglobigerina pennyi mark the latest Campanian worldwide, we approximate the Campanian/Maastrichtian boundary based on the FOs of Trinitella scotti and Rugoglobigerina hexacamerata, and identify the basal Maastrichtian by the last occurrence of Laeviella bollii.

Retrieving planktonic foraminifera from lithified rocks, examples from the Eocene limestones and marls (External Dinarides, Croatia)

Paleoecologic (paleoclimatologic) and biostratigraphic studies of pelagic and deep-water deposits rely on the identification of planktonic foraminifera. Here we report and compare the results of planktonic foraminiferal assemblages from the Middle Eocene indurated limestones and marls collected in the External Dinarides extracted with acetic acid of different concentrations (50%, 60%, 70% and 80%) and different reaction (exposure) times. The deposits originated within the Dinaric foreland basin, have been assigned to the so-called Transitional beds and Flysch, and are characterized by different ratio of carbonate content and degree of lithification. The aim of this paper is to compare the efficiency of the laboratory procedures for obtaining isolated specimens and to evaluate the impact of preparation procedure on the quality of tests (complete test vs. secondary dissolution effects). For each acetic concentration we assessed:(1)the effectiveness of the treatment in terms of the time required for successful extraction of planktonic foraminifera, and(2)the degree of dissolution by analyses of dissolution proxies, including the weight percentage of sieved residues after disaggregation and preservation features of the tests. Our results indicate that accurate taxonomic analysis of carbonate rocks requires the use of 60% acetic acid for a shorter reaction time, and hydrogen peroxide methods for marls.

Planktonic foraminiferal biostratigraphy and palaeobiogeography of the Albian–Turonian succession from the Sulaiman Range, Pakistan

Detailed taxonomy and biostratigraphy of the limestone, shale‐rich Cretaceous Goru Formation within the western Sulaiman Range refer to diverse and well‐preserved planktonic foraminifera. The age‐diagnostic planktonic foraminifera encountered are; Biticinella breggiensis (middle Albian), Rotalipora ticinensis (late Albian), Rotalipora appenninica (latest Albian), Rotalipora brotzeni (early Cenomanian), Rotalipora cushmani (late Cenomanian), Whiteinella archeocretacea (latest Cenomanian–Earliest Turonian), Helvetoglobotruncana helvetica (early to middle Turonian), and Marginotruncana sigali (late middle to late Turonian). Based on the recovered planktonic foraminifera a middle Albian–late Turonian age has been assigned to the studied formation. The Rotalipora reicheli (middle Cenomanian) Zone was not identified as the First Appearance Datum of the nominal taxa present within the Rotalipora cushmani Zone. The planktonic foraminiferal assemblage characteristics suggest that the bioprovince of the Cretaceous Goru Formation belongs to the subtropical, warm water environment belonging to the Cretaceous Tethyan Realm and the palaeobiogeographic reconstruction suggests that the Neotethys was well‐connected during the Albian–Turonian time interval.