Melanie J. Coathup, BSc, PhD, Niall Kent, BSc, PhD, Robert Hill, BSc, MSc, PhD, DIC, Roberta De Godoy, DVM, MSc, PhD, Wen-Yu Quak, BSc, Lyris Onwordi, MBBS, MSc, Gordon W. Blunn, Bsc, PhD. University College London, Division of Surgery and Interventional Science, Institute of Orthopaedics and Musculoskeletal Science, Stanmore, Middlesex, United Kingdom, Barts and The London School of Medicine and Dentistry, Queen Mary University, London, United Kingdom.