Race differences in perceptions of discrimination are well documented, but questions remain about contextual- and individual-difference moderators of when White and Black Americans see racial bias. We examined how temporal framing (focusing on past decades or not), race, political party, and domain of discrimination influence discrimination perceptions. Temporal framing did not moderate perceptions for White or Black participants (Study 1). Perceived anti-White and anti-Black discrimination converged over time (from the 1950s to the present), but especially so among White participants (Studies 1 and 2). Domain of discrimination moderated perceptions, with White respondents perceiving the steepest rise in anti-White discrimination and the steepest decline in anti-Black discrimination in the education and employment domains (Study 2). Across both studies, only White Republicans reported that White Americans face more discrimination than Black Americans. This research extends the literature on racial and political divides in discrimination perceptions, and highlights variability in perceptions across discrimination domains.