lifetime of a bird, as all three accipiters hunt rather elusive prey (primarily birds), and the gradual improvement of hunting techniques seems especially plausible for such prey. Though we have no evidence for or against it, there could be a preferential mating system in which adults with the darkest red eyes could have the greatest success in gaining mates. The disadvantages inherent in red eye coloration might prevent any tendency toward early assumption of this color in inexperienced birds, and the relatively greater danger to eyes of females (the sex that normally feeds chicks) could explain the relatively slow transition to red eyes in this sex. Though one might suggest that age recognition could be achieved without employing the color red, it is questionable whether a preferential mating system based on differences in eye coloration could be SLIStained if the preferred color did not also have disadvantages to its bearer at least at some age in life. Otherwise, selection would presumably lead to a uniform population with preferred eye coloration. The above hypothesis is not without difficulties. It is relevant to note that iris color changes of Marsh Hawks (Circus cyangus), as intensively studied by Hamerstrom ( Inland Bird Banding News 40:43, 1968 ) and Balfour (Bird Study 17:47, 1970), also occur faster in males than in females. In this species the transition from an initial brown coloration to a final yellow coloration takes approximately 1 year for males and up to 6 or 7 years for females. It seems unlikely that the hypothesis developed above could be directly applied to Marsh Hawks, though pecking preferences of Marsh Hawk chicks remain to be explored. A second difficulty concerns the role of an experience factor in delaying red eye coloration in male accipiters. As an order of magnitude guess, the chances of a male losing his mate during the breeding cycle and carrying on alone in feeding chicks might be on the order of one in a hundred breeding attempts.
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