The Permian (Changhsingian) Umm Irna Formation (Jordan) yields important plant fossils that have influenced our understanding of the early evolution of plants that later dominated the vegetation of the Mesozoic. The formation was deposited in a fluvial regime characterised by the co-occurrence of stacked small-scale braided channels and of larger, higher-sinuosity channels with deposition on point bars, all intercalated into a heterogeneous background sedimentation of finer-grained floodplain deposits, including stacked crevasse-splay deposits, interspersed abandoned-channel and oxbow-lake fills, and lenses of peaty back-swamp deposits. Here, we document the remarkable compositional variability expressed in 124 palynomorph assemblages that were recovered from different argillaceous deposits in the Umm Irna Formation within a ∼2 km2 outcrop area around a major meandering channel. Our results reveal lateral and vertical variation within individual argillaceous units, as well as lateral variation between different argillaceous units in the same sedimentary complex at similar stratigraphic levels. When combined with plant macrofossil data, the results show that variability in the palynological content may be influenced mainly by two factors. The first is connectedness to the major channel; the drainage system carried a mixture of palynomorphs from across the floodplain but also from a large hinterland of tributaries draining higher parts of the floodplain and uplands beyond to the south and east. The second is the influence of local vegetation communities on the floodplain which would likely be more marked in less connected parts of the floodplain. These findings have implications for the palynological study of alluvial sequences of the Palaeozoic and Mesozoic, and for understanding the ability to identify mudstone units via palynological fingerprinting. Such individual units are of great importance as baffles to fluid flow in reservoirs that might be used for oil and gas, low-temperature geothermal energy storage, carbon capture and storage, and hydrogen storage in alluvial successions.
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