Oak Sites Research Articles

Reconstruction of a regional drought index in southern Sweden since AD 1750

We used a network of eight pedunculate oak ( Quercus robur L.) sites ( ntrees = 70) and one Scots pine ( Pinus sylvestris L.) site ( ntrees = 53) to develop drought-sensitive master chronologies for the two areas in southern Scandinavia: a SW-area centred on 57°N 12.7°E and a NE-area centred on 58.8°N 18.2°E. The ratio of actual to equilibrium evapotranspiration (AET/EET) was used as a measure of drought during the growing season defined as the period with average daily temperatures above 9°C. Instrumental data were used to parameterize the relationship between tree-ring data and the drought index (DI) over 1922—2000 for the SW area and over 1922—1995 for the NE area. The DI reconstructions explained 29.7% (SW area) and 43.7% (NE area) of the variance in the observed DI index in the calibration period, and were extended back to AD 1770 for the SW area and to AD 1750 for the NE area. Reconstructed drought dynamics suggested strong decadal- and century-scale temporal variability and limited regional synchronicity over 1770—2000. Large variations in DI were observed in both regions in the second half of the 1700s. Dry conditions were synchronously reconstructed in both sub-regions during 1781—1784, 1853—1855, and, to a lesser degree, during 1974—1978. Over the 1945—1975 period the SW area exhibited a trend towards drier growing seasons, whereas no such trend could be identified for the NE area. Analysis of correlation maps indicated that regional DI dynamics reflected two different climate regimes, associated with the Kattegat area (SW reconstruction) and southeastern Swedish coast of the Baltic sea (NE reconstruction).

Reprint of: Lessons from long-term studies of harvest methods in southwestern ponderosa pine–Gambel oak forests on the Fort Valley Experimental Forest, Arizona, U.S.A.

The Fort Valley Experiment Station (now Fort Valley Experimental Forest) has contributed many long-term studies to forest research. This paper focuses on a “Methods of Cutting” study initiated in 1913 on the Coulter Ranch Unit of Fort Valley and how that long-term study yielded important ecological and management lessons. We quantified the historical and contemporary forest patterns at this ponderosa pine–Gambel oak site, which was harvested using three different harvesting systems in 1913 (seed tree, group selection, and light selection) and was partially excluded from livestock browsing in 1919. Using nine historically stem-mapped permanent plots for the following three stand structural scenarios: 1913 pre-harvest (modeled), post-harvest (actual), and 2003–2006 (actual) conditions, we examined the short- and long-term consequences of harvest and livestock grazing land-use and stand dynamics. We assessed changes in spatial pattern under each harvesting system and in each structural scenario, and lastly, we examined spatial and temporal tree recruitment patterns as observed in the contemporary (2003–2006) conditions. The seed tree harvests effectively converted the spatial patterns from aggregated to random and left few trees, while the group selection and light selection had varying effects, but consistently exaggerated the spatial patchiness of the stand. By 2003–2006, all plots were aggregated at all scales and were one large patch of predominately small trees. Sites that were harvested, but excluded from livestock browsing had 40% more trees in 2003–2006. Contemporary recruitment patterns were significantly aggregated under all harvesting systems, but were most strongly aggregated if the site received a group selection or light selection cut. For group and light selection, pine seedlings initially established in stump patches created by harvesting and then proceeded to fill-in the remaining area, with recruitment rarely found under the residual pine or oak trees. Long-term data sets, such as these established by the Fort Valley Experimental Forest in 1913, are essential for quantifying the impact of historical land-use practices on contemporary forest composition and structure. Ignoring land-use legacies may lead to the misinterpretation of stand dynamics and development, and therefore should be explicitly quantified and incorporated into future management and restoration activities.

Private Net Benefits from Afforesting Marginal Cropland and Shrubland with Cork Oaks in Spain

Abstract We estimate the private net benefits from afforesting marginal shrubland and cropland with cork oaks at two Spanish cork oak sites, Aljibe and Gavarres, in different time horizon frames. The analysis integrates market and nonmarket (landowner amenity) private capital incomes, including government afforestation grants. We apply cost-benefit analysis techniques for estimating the investment present value in the periods analyzed, taking residual values of cork oak woodland into account. The results show that the profitability rate of cork oak afforestation strongly depends on government subsidies. The value of private amenities, at least in Aljibe, seems to be positively affected by the forest area's share of the total estate surface. Thus, an increase in the afforested area would enhance the private amenity annual income with respect to maintaining the original property land's treeless uses. A higher level of private amenities attached to the forest area may imply a reduction in government grants for encouraging cork oak afforestation.

Towards a Trauma Network: An Interview with Professor Keith Porter

Arriving at Selly Oak rail station in Birmingham it is hard to avoid the new Queen Elizabeth Hospital Birmingham (QEHB), which dominates the Birmingham skyline and will form a prime hub of the new trauma network for the West Midlands. However, on the day I visit Keith Porter, Professor of Clinical Traumatology with University Hospitals Birmingham, the famous old Selly Oak site is in its last few days before the move to this new state-of-the-art centre.

Lessons from long-term studies of harvest methods in southwestern ponderosa pine–Gambel oak forests on the Fort Valley Experimental Forest, Arizona, U.S.A.

The Fort Valley Experiment Station (now Fort Valley Experimental Forest) has contributed many long-term studies to forest research. This paper focuses on a “Methods of Cutting” study initiated in 1913 on the Coulter Ranch Unit of Fort Valley and how that long-term study yielded important ecological and management lessons. We quantified the historical and contemporary forest patterns at this ponderosa pine–Gambel oak site, which was harvested using three different harvesting systems in 1913 (seed tree, group selection, and light selection) and was partially excluded from livestock browsing in 1919. Using nine historically stem-mapped permanent plots for the following three stand structural scenarios: 1913 pre-harvest (modeled), post-harvest (actual), and 2003–2006 (actual) conditions, we examined the short- and long-term consequences of harvest and livestock grazing land-use and stand dynamics. We assessed changes in spatial pattern under each harvesting system and in each structural scenario, and lastly, we examined spatial and temporal tree recruitment patterns as observed in the contemporary (2003–2006) conditions. The seed tree harvests effectively converted the spatial patterns from aggregated to random and left few trees, while the group selection and light selection had varying effects, but consistently exaggerated the spatial patchiness of the stand. By 2003–2006, all plots were aggregated at all scales and were one large patch of predominately small trees. Sites that were harvested, but excluded from livestock browsing had 40% more trees in 2003–2006. Contemporary recruitment patterns were significantly aggregated under all harvesting systems, but were most strongly aggregated if the site received a group selection or light selection cut. For group and light selection, pine seedlings initially established in stump patches created by harvesting and then proceeded to fill-in the remaining area, with recruitment rarely found under the residual pine or oak trees. Long-term data sets, such as these established by the Fort Valley Experimental Forest in 1913, are essential for quantifying the impact of historical land-use practices on contemporary forest composition and structure. Ignoring land-use legacies may lead to the misinterpretation of stand dynamics and development, and therefore should be explicitly quantified and incorporated into future management and restoration activities.

Process-based simulation of seasonality and drought stress in monoterpene emission models

Abstract. Canopy emissions of volatile hydrocarbons such as isoprene and monoterpenes play an important role in air chemistry. They depend on various environmental conditions, are highly species-specific and are expected to be affected by global change. In order to estimate future emissions of these isoprenoids, differently complex models are available. However, seasonal dynamics driven by phenology, enzymatic activity, or drought stress strongly modify annual ecosystem emissions. Although these impacts depend themselves on environmental conditions, they have yet received little attention in mechanistic modelling. In this paper we propose the application of a mechanistic method for considering the seasonal dynamics of emission potential using the "Seasonal Isoprenoid synthase Model" (Lehning et al., 2001). We test this approach with three different models (GUENTHER, Guenther et al., 1993; NIINEMETS, Niinemets et al., 2002a; BIM2, Grote et al., 2006) that are developed for simulating light-dependent monoterpene emission. We also suggest specific drought stress representations for each model. Additionally, the proposed model developments are compared with the approach realized in the MEGAN (Guenther et al., 2006) emission model. Models are applied to a Mediterranean Holm oak (Quercus ilex) site with measured weather data. The simulation results demonstrate that the consideration of a dynamic emission potential has a strong effect on annual monoterpene emission estimates. The investigated models, however, show different sensitivities to the procedure for determining this seasonality impact. Considering a drought impact reduced the differences between the applied models and decreased emissions at the investigation site by approximately 33% on average over a 10 year period. Although this overall reduction was similar in all models, the sensitivity to weather conditions in specific years was different. We conclude that the proposed implementations of drought stress and internal seasonality strongly reduce estimated emissions and indicate the measurements that are needed to further evaluate the models.

Degradability of soils under oak and pine in Central Spain

The purpose of this research was to study the influence of the vegetation on the soil C pool of forests of pines (Pinus sylvestris) and oaks (Quercus pyrenaica), located in Central-Western Spain. Horizons from selected soils located in these forests were sampled, and the soil organic C (SOC) was determined. In addition, in vitro incubation experiments were carried out, under controlled conditions, to monitor the stability of SOC against the microbial activity. Soil humus fractions were isolated following a classical procedure of chemical fractionation using alkaline solutions, before and after the incubation experiment. A deeper O horizon was found under the pine forest than under oak one; however, higher SOC content was found in the oak site than that under pine one. During the in vitro mineralization process, a lower CO2 production by the soil sample from pine forest was observed, in relation to that emitted by the oak soil. In addition, a lower humification degree was estimated for the soil humus under pines than for that under oaks. In conclusion, replacement of oaks by pines produced a decrease in SOC accumulation and a lower quality of humus in the forest soils.

Viability of Northern Pin and White Oak Reserve Trees in Wisconsin Scrub Oak Sites

Abstract Retention of reserve trees can serve multiple objectives when harvests are conducted on scrub oak sites. The objective of the study was to determine the growth and vigor response of northern pin and white oak held as reserve trees from harvests occurring over a 15-year time period onscrub oak sites in Central Wisconsin. Reserve trees were evaluated for volume growth, epicormic branching, live crown ratio, and tree vigor. White oak maintained or improved in vigor and exhibited increased rates of volume growth with no measured mortality. Northern pin oak, although exhibitingincreased volume growth, tended to decline in vigor and experienced an overall annual mortality rate of 5.5%. For managers that are considering multiple objectives, white oak can provide a long-lived vigorous reserve tree while northern pin oak may survive but will experience decline and higherrates of mortality, which may contribute to a long-term goal of snag retention as an element of wildlife habitat.

Northern Pin Oak Stump Sprouting Frequency on Scrub Oak Sites of Central Wisconsin

Abstract Stump sprouts are an important component of the regeneration for northern pin oak on scrub oak sites in Central Wisconsin, but there exists little quantitative information about their actual contribution. This study examines the occurrence of sprouting from northern pin oak stumps remaining after overstory removal in four stands in Central Wisconsin. On average, 85% of all stumps sprouted. Sprouting frequency diminished significantly (P < 0.0001) as stump diameter increased, but even the largest stumps sprouted at rates of 71% or better. With stump sprouting rates this high, coppice regeneration could fill much of the regeneration need for similar scrub oak sites.

Tree influence on soil biological activity: What can be inferred from the optical examination of humus profiles?

Humus forms may vary in different forest stands, but the local influence of trees upon soil microbial and faunal activities is still imperfectly known. Optical methods could help to discern processes of litter transformation and formation of organo-mineral assemblages, allowing a better diagnostic of tree influences upon humus-soil development. The microstratification of humus was studied under a beech ( Fagus crenata), a mixed oak forest ( Quercus crispula and Quercus serrata), and a cedar ( Cryptomeria japonica) plantation. The three sites are located in Kyoto (Japan), and share similar environmental conditions. Litter decomposition rates and soil fauna were also investigated. At the beech site, which had the thickest O horizon, the main process was the gradual fragmentation of litter. This process, together with shallow root and weak fungal development, gave rise to a stable sandwich-like structure in the O horizon. In contrast, the oak site showed a two-step transformation of litter. Initially, litter decomposition was triggered by the activity of white rot fungi, and the discarded litter decayed much more slowly thereafter. The cedar site exhibited a sharp vertical delineation between upper thick Oe horizon developed since plantation time and a relict A horizon. The optical method thus demonstrated differences in soil biological activities and litter transformation patterns under the three sites.

A contribution to the sessile oak site and beech site defining in central Serbia

The study was carried out in sessile oak forests and beech forests in the region central Serbia. The stands are classified as pure stands with the percentage of other species up to 10% per tree number, mixed forests of sessile oak with other species, and mixed forests of beech with other species, whose percentage does not exceed 50%. Altogether 257 stands were monitored - 202 beech stands and 55 sessile oak stands. By the applied method of defining the local heat potential (Lujic, 1960), modified by Ratknic et al. (2001) and Krstic (2004, 2008), which represents possibility of soil heating without vegetation, were determined. In this way, a scale of 162 possible combinations of local heat potential was obtained, which explains more precisely the dependence of beech stands and sessile oak stands on the topographic conditions. By applying the weighted values of the thermal co-ordinates of aspect and slope (E) for each altitudinal belt of 100 m, it was concluded that pure stands have the widest ecological range. Pure beech stands occur at the sites with 34 combinations of thermal co-ordinates E.V=4.6 to 8.12. Pure sessile oak stands occur at the sites with 12 combinations of thermal co-ordinates E.V=5.10 to 8.11. The percentage of mixed beech stands with other broadleaf species is the highest at the sites with the co-ordinate V=10-11 (at the altitudes between 700 and 900 m) is about 60 %. Mixed stands of sessile oak and beech are located on the terrains with combinations of thermal co-ordinates E.V=7.9 to 8.12. By using the local heat potential of a region, it can be identified which sites, i.e. which combinations of exposure, slope and altitude belong to the particular tree species. Consequently, a more reliable selection of tree species can be done for the bio-reclamation of barrens and other deforested terrains.

Complex climate controls on 20th century oak growth in Central-West Germany

We analyze interannual to multi-decadal growth variations of 555 oak trees from Central-West Germany. A network of 13 pedunculate oak (Quercus robur L.) and 33 sessile oak (Quercus petraea (Matt.) Liebl.) site chronologies is compared with gridded temperature, precipitation, cloud-cover, vapor pressure and drought (i.e., Palmer Drought Severity Index, PDSI) fluctuations. A hierarchic cluster analysis identifies three groups for each oak species differentiated by ecologic settings. When high precipitation is primarily a characteristic for one Q. robur and one Q. petraea cluster, the other clusters are more differentiated by prevailing temperature conditions. Correlation analysis with precipitation and vapor pressure reveals statistically significant (P < or = 0.05) correlations for June (r = 0.51) and annual (r = 0.43) means. Growth of both species at dry sites correlates strongly with PDSI (r = 0.39, P < or = 0.05), and weakly with temperature and cloud-cover. In natural stands, Q. robur responds more strongly to water depletion than Q. petraea. Twenty-one-year moving correlations show positive significant growth response to both PDSI and precipitation throughout the 20th century, except for the 1940s - an anomalously warm decade during which all oak sites are characterized by an increased growth and an enhanced association with vapor pressure and temperature. We suggest that the wider oak rings that are exhibited during this period may be indicative of a nonlinear or threshold-induced growth response to drought and vapor pressure, and run counter to the general response of oak to drought and precipitation that normally would result in suppressed growth in a warmer and drier environment. As the wide rings are formed during the severe drought period of the 20th century, a complex model seems to be required to fully explain the widespread oak growth. Our results indicate uncertainty in estimates of future growth trends of Central European oak forests in a warming and drying world.

Methodologically induced differences in oak site classifications in a homogeneous tree-ring network

Spatiotemporal variations in tree growth are induced by varying environmental conditions. Different methods like variants of the principal component analysis and the hierarchical cluster analysis are commonly applied in dendroecology to separate subsets of growth patterns within large tree-ring datasets. To seek for methodological differences in classification techniques and their specific characteristics, we compared three standard methods using a homogeneous oak ( Quercus spp.) network from temperate forests in Central-West Germany. Classifications of the original dataset consisting of 46 oak ring-width sites, carried out with the varimax rotated principle component analysis, Ward's method and the average linkage method, reveal differences in the classification of approximately 20% of the sites. Analyses with modified datasets are calculated to evaluate effects of dataset extension, different time periods and different tree-ring detrendings. The application of the principal component analysis generally leads to the most stable site classifications, whereas the most sensitive response to changes in the dataset is obtained by Ward's method. The average linkage method separates single sites in the classification and thus emphasises outliers within the tree-ring network.

Comparison of growth, nutrition and soil properties of pure stands of Quercus castaneifolia and mixed with Zelkova carpinifolia in the Hyrcanian forests of Iran

The continued existence of oakwoods in the Hyrcanian forests of Iran is threatened by lack of natural regeneration which is limited by several factors. Hence, in order to promote expansion and rehabilitation of the oak forests, a program of oak seedling planting may be required. Quercus castaneifolia (as target species) and Zelkova carpinifolia (as native component species) were planted in five proportions (100 Q, 70 Q:30 Z, 60 Q:40 Z, 50 Q:50 Z, 40 Q:60 Z) in the Noor region (North Iran). After 11 years, the effects of the species on the growth of the trees, nutrient concentrations in the live and senescent leaves and on soil properties were assessed. The results showed the survival, total height and clean bole of the individual Quercus trees were positively affected by the presence of Zelkova. Percent retranslocation of the nutrients in Quercus trees was: K > N > P. Leaf-litter fall production ranged from 4.89 to 6.27 Mg ha −1 year −1. P and Mg fluxes, P retranslocation, P concentration in fully expanded leaves and N, P, K and Mg concentrations in the senescent leaves of Quercus trees were higher in some of the mixed plantations than in the monocultures of the Quercus trees. The obtained results indicated that the mixing with Siberian elm increased the productivity and sustainability of the oak sites. Within the framework of this experiment, it appeared that production was maximized when these two species were grown together in the proportion of 70% Quercus and 30% Zelkova.

Parameterization of the Stomatal Component of the DO3SE Model for Mediterranean Evergreen Broadleaf Species

An ozone (O3) deposition model (DO3SE) is currently used in Europe to define the areas where O3 concentrations lead to absorbed O3 doses that exceed the flux-based critical levels above which phytotoxic effects would be likely recorded. This mapping exercise relies mostly on the accurate estimation of O3 flux through plant stomata. However, the present parameterization of the modulation of stomatal conductance (gs) behavior by different environmental variables needs further adjustment if O3 phytotoxicity is to be assessed accurately at regional or continental scales. A new parameterization of the model is proposed for Holm oak (Quercus ilex), a tree species that has been selected as a surrogate for all Mediterranean evergreen broadleaf species. This parameterization was based on a literature review, and was calibrated and validated using experimentally measured data of gs and several atmospheric and soil parameters recorded at three sites of the Iberian Peninsula experiencing long summer drought, and very cold and dry winter air (El Pardo and Miraflores) or milder conditions (Tietar). A fairly good agreement was found between modeled and measured data (R2 = 0.64) at Tietar. However, a reasonable performance (R2 = 0.47–0.62) of the model was only achieved at the most continental sites when gs and soil moisture deficit relationships were considered. The influence of root depth on gs estimation is discussed and recommendations are made to build up separate parameterizations for continental and marine-influenced Holm oak sites in the future.

Soil CO2Efflux in a Mixed Pine-Oak Forest in Valsaín (Central Spain)

Soil-surface CO2 efflux and its spatial and temporal variation were investigated in a southern Mediterranean, mixed pine-oak forest ecosystem on the northern slopes of the Sierra de Guadarrama in Spain from February 2006 to July 2006. Measurements of soil CO2 efflux, soil temperatures, and moisture were conducted in nine 1963-m2 sampling plots distributed in a gradient around the ecotone between Pinus sylvestris L. and Quercus pyrenaica Lam. forest stands. Total soil organic matter, Walkey-Black C, particulate organic matter, organic matter fraction below 53 μm, total soil nitrogen content, total soil organic carbon content, and pH were also measured under three representative mature oak, pine, and mixed pine-oak forest stands. Soil respiration showed a typical seasonal pattern with minimums in winter and summer, and maximums in spring, more pronounced in oak and oak-pine stands. Soil respiration values were highest in pine stands during winter and in oak stands during spring and summer.Soil respiration was highly correlated with soil temperatures in oak and pine-oak stands when soil moisture was above a drought threshold of 15%. Below this threshold value, soil moisture was a good predictor of soil respiration in pine stands. Greater soil organic matter, particulate organic matter, Walkey-Black C, total organic C, and total N content in pine compared to oak sites potentially contributed to the greater total soil CO2 efflux in these stands during the winter. Furthermore, opposing trends in the organic matter fraction below 53 μm and soil respiration between plots suggest that in oak stands, the C forms are less affected by possible changes in use. The effects of soil properties on soil respiration were masked by differences in soil temperature and moisture during the rest of the year. Understanding the spatial and temporal variation even within small geographic areas is essential to assess C budgets at ecosystem level accurately. Thus, this study bears important implications for the study of large-scale ecosystem dynamics, particularly in response to climatic change.

Effects of Land Use on Soil Respiration

We examined constraints on soil CO2 respiration in natural oak woodlands, and adjacent vineyards that were converted approximately 30 yr ago from oak woodlands, in the Oakville Region of Napa Valley, California. All sites were located on the same soil type, a Bale (variant) gravelly loam (fine-loamy, mixed, superactive, thermic Cumulic Ultic Haploxeroll) and dominated by C3 vegetation. Seasonal soil CO2 efflux was greatest at the oak woodland sites, although during the summer drought the rates of soil CO2 efflux measured from oak sites were generally similar to those measured from the vineyards. Soil profile CO2 concentrations at the oak woodland sites were lower below 15 cm despite higher CO2 efflux rates. Soil gas diffusion coefficients for oak sites were larger than for vineyard sites, and this indicated that the apparent discrepancy in soil profile carbon dioxide concentration ([CO2]) may be caused by a diffusion limitation. Soil profile [CO2] and delta13C values showed substantial temporal changes over the course of a year. Vineyard soil CO2 was more depleted in 13CO2 below 25 cm in the soil profile during the active growing season as indicated by more negative delta13C ratios. This result indicated that different C sources were being oxidized in vineyard soils. Annual C losses were less from vineyard soils (7.02 +/- 0.58 Mg C ha(-1) yr(-1)) as compared to oak soils (15.67 +/- 1.44 Mg C ha(-1) yr(-1)), and both were comparable to losses reported in previous investigations.

Four-year comparison of water contents beneath a grass ley and a deciduous oak wood overlying Triassic sandstone in lowland England

Differences in the seasonal water dynamics of a sand soil overlying Triassic sandstone have been investigated to a depth of 9 m beneath both a grass ley and a pendunculate oak woodland within Clipstone Forest, Nottinghamshire, England. Fortnightly measurements with a neutron-probe over a four-year period allowed a comparison of the soil–rock water content to depths rarely reported. In spring, the rate of decrease of water content of the uppermost 2 m of soil was much greater under grass than oak woodland. In contrast, the rate of decrease under oak was greater after leafing out in May, while the rate of rewetting in early autumn was lower for this land-use until senescence and leaf-fall in late autumn. In the uppermost 2 m of soil – a depth that includes all plant roots – the soil moisture minima were between 47 and 58 mm lower under oak than grass in each year of monitoring. As a result of both these drier conditions and the comparatively late leaf fall of this species, penetration of the winter-season wetting front to 2 m was delayed by between one and three months at the oak site relative to grass in years of near-average rainfall. Rewetting at 9 m lags by 10–12 months, compared to the surface, giving an average penetration rate for the wetting front to this depth of 25–30 mm day −1 , with little observable differences between the land-uses. Preferential flow is evident under both sites, affecting the profile to 3 m in all years and to at least 6 m following winters experiencing exceptional rainfall.

Results of the centralisation of adult emergency oral and maxillofacial surgical services at the University Hospital, Birmingham

In 2001, the oral and maxillofacial surgical (OMFS) services for adults in Birmingham were centralised at the Selly Oak site of the University Hospital and the OMFS department was given access to the emergency operating theatre every day. We examined the effects of this on the emergency workload during the 6 months after centralisation and compared it with the emergency workload at the University Hospital during the 6 months before centralisation. The number of adult patients who attended the University Hospital as emergencies increased from 242 in the 6 months before centralisation to 545 in the subsequent 6 months (an increase of 127%). Of the latter 164 (30%) were admitted and operated on, 138 (84%) within 24 h of admission. Despite the large increase in the number of patients, 102 (74%) were operated on during normal working hours. After centralisation, all operations were done on multidisciplinary emergency lists compared with only 55% before centralisation. Centralisation did not reduce the access of patients to the emergency service and conformed with the guidelines of the National Confidential Enquiry into Perioperative Deaths (NCEPOD).

Stand development on reforested bottomlands in the Mississippi Alluvial Valley

Reforestation of bottomland hardwood sites in the southeastern United States has markedly increased in recent years due, in part, to financial incentives provided by conservation programs. Currently >250,000 ha of marginal farmland have been returned to hardwood forests. I observed establishment of trees and shrubs on 205 reforested bottomlands: 133 sites were planted primarily with oak species (Quercus spp.), 60 sites were planted with pulpwood producing species (Populus deltoides, Liquidambar styraciflua, or Platanus occidentalis), and 12 sites were not planted (i.e., passive regeneration). Although oak sites were planted with more species, sites planted with pulpwood species were more rapidly colonized by additional species. The density of naturally colonizing species exceeded that of planted species but density of invaders decreased rapidly with distance from forest edge. Trees were shorter in height on sites planted with oaks than on sites planted with pulpwood species but within a site, planted trees attained greater heights than did colonizing species. Thus, planted trees dominated the canopy of reforested sites as they matured. Planted species acted in concert with natural invasion to influence the current condition of woody vegetation on reforested sites. Cluster analysis of species importance values distinguished three woody vegetation conditions: (1) Populus deltoides stands (2) oak stands with little natural invasion by other tree species, and (3) stands dominated by planted or naturally invading species other than oaks. Increased diversity on reforested sites would likely result from (a) greater diversity of planted species, particularly when sites are far from existing forest edges and (b) thinning of planted trees as they attain closed canopies.