Northern Continents Research Articles

The Biogeographic South-North Divide of Polygonatum (Asparagaceae Tribe Polygonateae) within Eastern Asia and Its Recent Dispersals in the Northern Hemisphere.

Eastern Asia (EA) is a key region for the diversification of flowering plants in the Northern Hemisphere, but few studies have focused on the biogeographic history within EA in the context of the other northern continents. Polygonatum is an important medicinal genus widely distributed in the Northern Hemisphere with its highest species richness in EA, and it represents an excellent model for studying the evolution of biogeographic patterns in this region. Divergence time estimation was used to examine the biogeographic history of Polygonatum based on nuclear ITS and four plastid sequences (rbcL, matK, psbA–trnH and trnC–petN) from 30 Polygonatum species and 35 outgroup taxa. The ancestral area of Polygonatum and subsequent dispersal routes were inferred using Bayes-Lagrange. Polygonatum was estimated to have originated in southern EA during the middle Miocene (14.34–13.57 Ma) with subsequent south-to-north expansion in the late Miocene. Multiple intercontinental dispersal events were inferred between EA and Europe or North America, and all of them have occurred recently in the late Miocene to Pliocene. The separation of Polygonatum into the south and north lineages and their subsequent diversifications in the late Miocene supports the existence of a biogeographic divide between the northern and southern parts of EA that also coincides with the retreat and redevelopment of the arid zone in EA in the Neogene. Our results demonstrate the complexity of biogeographic history of Polygonatum in the Northern Hemisphere including early vicariance followed by frequent and recent dispersals in the Neogene.

Response of the Pacific inter-tropical convergence zone to global cooling and initiation of Antarctic glaciation across the Eocene Oligocene Transition.

Approximately 34 million years ago across the Eocene–Oligocene transition (EOT), Earth’s climate tipped from a largely unglaciated state into one that sustained large ice sheets on Antarctica. Antarctic glaciation is attributed to a threshold response to slow decline in atmospheric CO2 but our understanding of the feedback processes triggered and of climate change on the other contents is limited. Here we present new geochemical records of terrigenous dust accumulating on the sea floor across the EOT from a site in the central equatorial Pacific. We report a change in dust chemistry from an Asian affinity to a Central-South American provenance that occurs geologically synchronously with the initiation of stepwise global cooling, glaciation of Antarctica and aridification on the northern continents. We infer that the inter-tropical convergence zone of intense precipitation extended to our site during late Eocene, at least four degrees latitude further south than today, but that it migrated northwards in step with global cooling and initiation of Antarctic glaciation. Our findings point to an atmospheric teleconnection between extratropical cooling and rainfall climate in the tropics and the mid-latitude belt of the westerlies operating across the most pivotal transition in climate state of the Cenozoic Era.

Paleoproterozoic evolution of the Guiana Shield in Suriname: A revised model

AbstractThe Proterozoic basement of Suriname consists of a greenstone–tonalite–trondhjemite–granodiorite belt in the northeast of the country, two high-grade belts in the northwest and southwest, respectively, and a large granitoid–felsic volcanic terrain in the central part of the country, punctuated by numerous gabbroic intrusions. The basement is overlain by the subhorizontal Proterozoic Roraima sandstone formation and transected by two Proterozoic and one Jurassic dolerite dyke swarms. Late Proterozoic mylonitisation affected large parts of the basement. Almost 50 new U–Pb and Pb–Pb zircon ages and geochemical data have been obtained in Suriname, and much new data are also available from the neighbouring countries. This has led to a considerable revision of the geological evolution of the basement. The main orogenic event is the Trans-Amazonian Orogeny, resulting from southwards subduction and later collision between the Guiana Shield and the West African Craton. The first phase, between 2.18 and 2.09 Ga, shows ocean floor magmatism, volcanic arc development, sedimentation, metamorphism, anatexis and plutonism in the Marowijne Greenstone Belt and the adjacent older granites and gneisses. The second phase encompasses the evolution of the Bakhuis Granulite Belt and Coeroeni Gneiss Belt through rift-type basin formation, volcanism, sedimentation and, between 2.07 and 2.05 Ga, high-grade metamorphism. The third phase, between 1.99 and 1.95 Ga, is characterised by renewed high-grade metamorphism in the Bakhuis and Coeroeni belts along an anticlockwise cooling path, and ignimbritic volcanism and extensive and varied intrusive magmatism in the western half of the country. An alternative scenario is also discussed, implying an origin of the Coeroeni Gneiss Belt as an active continental margin, recording northwards subduction and finally collision between a magmatic arc in the south and an older northern continent. The Grenvillian collision between Laurentia and Amazonia around 1.2–1.0 Ga caused widespread mylonitisation and mica age resetting in the basement.

The palaeobiology of high latitude birds from the early Eocene greenhouse of Ellesmere Island, Arctic Canada.

Fossils attributable to the extinct waterfowl clade Presbyornithidae and the large flightless Gastornithidae from the early Eocene (~52–53 Ma) of Ellesmere Island, in northernmost Canada are the oldest Cenozoic avian fossils from the Arctic. Except for its slightly larger size, the Arctic presbyornithid humerus is not distinguishable from fossils of Presbyornis pervetus from the western United States, and the Gastornis phalanx is within the known size range of mid-latitude individuals. The occurrence of Presbyornis above the Arctic Circle in the Eocene could be the result of annual migration like that of its living duck and geese relatives, or it may have been a year-round resident similar to some Eocene mammals on Ellesmere and some extant species of sea ducks. Gastornis, along with some of the mammalian and reptilian members of the Eocene Arctic fauna, likely over-wintered in the Arctic. Despite the milder (above freezing) Eocene climate on Ellesmere Island, prolonged periods of darkness occurred during the winter. Presence of these extinct birds at both mid and high latitudes on the northern continents provides evidence that future increases in climatic warming (closer to Eocene levels) could lead to the establishment of new migratory or resident populations within the Arctic Circle.

Church, Humboldt, and the Politics of Display

This essay takes as its starting point the spectacle of Heart of the Andes, a painting of South America by Frederic Edwin Church (American, 1826–1900), surmounted in an 1864 exhibition by portraits of the first three United States presidents. The author interprets the painting’s presentation in this context as a possibly unconscious imperial act of an expanding United States. Church’s South American travels, inspired by the writings of Alexander von Humboldt (German, 1769–1859), assimilate for the United States the Western Hemisphere’s southern continent in grand style landscapes that became entertaining reports on a world Humboldt had traveled intimately. Humboldt’s revelations in volumes of scientific inquiry, inspired observation, and disciplined romantic passion also left their mark on the northern continent and its own explorations. The essay debates other interpretations of Church’s and Humboldt’s aims and achievements against a carefully documented background, thus revealing the not-so-hidden ambitions that eventually led the United States to its corporate and political power in the southern continent.

First Digital Cranial Endocasts of late Oligocene Notohippidae (Notoungulata): Implications for Endemic South American Ungulates Brain Evolution

Notohippidae were middle-sized toxodonts recorded from the Eocene to the early Miocene. We provide the first description of the cranial endocasts of Rhynchippus equinus and Eurygenium latirostris based on three-dimensional reconstructions extracted from high-resolution X-ray computed tomography imagery. The endocasts of R. equinus and E. latirostris indicate that they were similar in size, proportions of the encephalic components, and neocortical design. The endocranial morphology of these notohippids is very close to that of toxodontids Adinotherium and Nesodon and differs markedly from both other Toxodontia and Typotheria. Notohippids, together with toxodontids, show the most complex neocortical surface among notoungulates. On the other hand, the neuromorphology of notohippids is in contrast to the neocortical morphological pattern described for Tertiary euungulates (Perissodactyla and Artiodactyla) from the Northern Hemisphere and litoptern Protetotheriidae, South American native euungulates. The relative brain size of R. equinus and E. latirostris compared with coeval holarctic euungulates from the late Oligocene are significantly below Perissodactyla and near the values obtained for Artiodactyla. Regarding the location of some functional neocortical areas, the expansion of the frontal lobe in Notohippidae may reflect the acquisition of heightened tactile sensitivity in the front of the snout, as recorded in the somatic sensory cortex of living euungulates. The bulging temporal lobe may reflect expansion of the auditory cortex, likely related to the marked enlargement of the middle ear chamber. Both neuromorphological and quantitative data suggest that during the late Paleogene, notohippids developed as complex and encephalized brains as those of the coeval Artiodactyla of northern continents.

The Southern Glacial Maximum 65,000 years ago and its Unfinished Termination

Glacial maxima and their terminations provide key insights into inter-hemispheric climate dynamics and the coupling of atmosphere, surface and deep ocean, hydrology, and cryosphere, which is fundamental for evaluating the robustness of earth's climate in view of ongoing climate change. The Last Glacial Maximum (LGM, ∼26–19 ka ago) is widely seen as the global cold peak during the last glacial cycle, and its transition to the Holocene interglacial, dubbed 'Termination 1 (T1)', as the most dramatic climate reorganization during this interval. Climate records show that over the last 800 ka, ice ages peaked and terminated on average every 100 ka (‘100 ka world’). However, the mechanisms pacing glacial–interglacial transitions remain controversial and in particular the hemispheric manifestations and underlying orbital to regional driving forces of glacial maxima and subsequent terminations remain poorly understood.Here we show evidence for a full glacial maximum in the Southern Hemisphere 65.1 ± 2.7 ka ago and its ‘Unfinished Termination’. Our 10Be chronology combined with a model simulation demonstrates that New Zealand's glaciers reached their maximum position of the last glacial cycle during Marine Isotope Stage-4 (MIS-4). Southern ocean and greenhouse gas records indicate coeval peak glacial conditions, making the case for the Southern Glacial Maximum about halfway through the last glacial cycle and only 15 ka after the last warm period (MIS-5a). We present the hypothesis that subsequently, driven by boreal summer insolation forcing, a termination began but remained unfinished, possibly because the northern ice sheets were only moderately large and could not supply enough meltwater to the North Atlantic through Heinrich Stadial 6 to drive a full termination. Yet the Unfinished Termination left behind substantial ice on the northern continents (about 50% of the full LGM ice volume) and after another 45 ka of cooling and ice sheet growth the earth was at inter-hemispheric Last Glacial Maximum configuration, when similar orbital forcing hit maximum-size northern ice sheets and ushered in T1 and thus the ongoing interglacial. This argument highlights the critical role of full glacial conditions in both hemispheres for terminations and implies that the Southern Hemisphere climate could transition from interglacial to full glacial conditions in about 15,000 years, while the Northern Hemisphere and its continental ice-sheets required half a glacial cycle.

Projected change in East Asian summer monsoon by dynamic downscaling: Moisture budget analysis

The summer monsoon considerably affects water resource and natural hazards including flood and drought in East Asia, one of the world’s most densely populated area. In this study, we investigate future changes in summer precipitation over East Asia induced by global warming through dynamical downscaling with the Weather Research and Forecast model. We have selected a global model from the Coupled Model Intercomparison Project Phase 5 based on an objective evaluation for East Asian summer monsoon and applied its climate change under Representative Concentration Pathway 4.5 scenario to a pseudo global warming method. Unlike the previous studies that focused on a qualitative description of projected precipitation changes over East Asia, this study tried to identify the physical causes of the precipitation changes by analyzing a local moisture budget. Projected changes in precipitation over the eastern foothills area of Tibetan Plateau including Sichuan Basin and Yangtze River displayed a contrasting pattern: a decrease in its northern area and an increase in its southern area. A local moisture budget analysis indicated the precipitation increase over the southern area can be mainly attributed to an increase in horizontal wind convergence and surface evaporation. On the other hand, the precipitation decrease over the northern area can be largely explained by horizontal advection of dry air from the northern continent and by divergent wind flow. Regional changes in future precipitation in East Asia are likely to be attributed to different mechanisms which can be better resolved by regional dynamical downscaling.

Accumulation of organic carbon in chernozems (Mollisols) under shelterbelts in Russia and the United States

Shelterbelts that were created in place of meadow and meadow-steppe landscapes of the forest-steppe zone of northern continents serve as areas of carbon accumulation and participate in the formation of soil organic matter. In the Great Plains of the United States (in North Dakota, South Dakota, and Nebraska) and on the Central Russian Upland (Belgorod, Voronezh, and Kursk oblasts), a general tendency toward an increase in the Corg pool in the topsoil (0–30 cm) from the marginal parts of the shelterbelts toward their central parts by about 3.5–10.0 t per each 10 m has been identified. In 55 years of the existence of shelterbelts on chernozems in the European part of Russia, the mean annual rate of the organic carbon accumulation in the upper meter has been varying within 0.7–1.5 t/ha. In 19 years of the existence of a shelterbelt in the area of Huron (South Dakota), the mean annual rate of the organic carbon accumulation in the 1-m-thick layer of the Bonilla soil series (Haplustolls) has reached 1.9 t/ha.

Last glacial maximum constraints on the Earth System model HadGEM2-ES

We investigate the response of the atmospheric and land surface components of the CMIP5/AR5 Earth System model HadGEM2-ES to pre-industrial (PI: AD 1860) and last glacial maximum (LGM: 21 kyr) boundary conditions. HadGEM2-ES comprises atmosphere, ocean and sea-ice components which are interactively coupled to representations of the carbon cycle, aerosols including mineral dust and tropospheric chemistry. In this study, we focus on the atmosphere-only model HadGEM2-A coupled to terrestrial carbon cycle and aerosol models. This configuration is forced with monthly sea surface temperature and sea-ice fields from equivalent coupled simulations with an older version of the Hadley Centre model, HadCM3. HadGEM2-A simulates extreme cooling over northern continents and nearly complete die back of vegetation in Asia, giving a poor representation of the LGM environment compared with reconstructions of surface temperatures and biome distributions. The model also performs significantly worse for the LGM in comparison with its precursor AR4 model HadCM3M2. Detailed analysis shows that the major factor behind the vegetation die off in HadGEM2-A is a subtle change to the temperature dependence of leaf mortality within the phenology model of HadGEM2. This impacts on both snow-vegetation albedo and vegetation dynamics. A new set of parameters is tested for both the pre-industrial and LGM, showing much improved coverage of vegetation in both time periods, including an improved representation of the needle-leaf forest coverage in Siberia for the pre-industrial. The new parameters and the resulting changes in global vegetation distribution strongly impact the simulated loading of mineral dust, an important aerosol for the LGM. The climate response in an abrupt 4× pre-industrial CO2 simulation is also analysed and shows modest regional impacts on surface temperatures across the Boreal zone.

High-level systematics of placental mammals: Current status of the problem

According to molecular data, the modern clades of placental mammals can be grouped into four supraordinal taxa (Xenarthra, Afrotheria, Euarchontoglires, and Laurasiatheria) whose phylogenetic interrelationships have been interpreted inconsistently. Among these taxa, the group Afrotheria has no morphological support. Within this group, there are orders that are clearly related to “condylarthrans” (“afroungulates”) and to insectivores (“afrosoricids”). The radiation of placental mammals (the crown clade Placentalia) started before the K/T boundary on the northern continents. Laurasiatheria are likely to be the oldest clade of placentals: the diversification of Lipotyphla and Euungulata started ∼80 Mya (Campanian) and ∼70 Mya (Maastrichtian), respectively. It is shown that the fossil record is the only reliable method to test the phylogenetic hypotheses based on the material of the molecular and morphological studies of recent taxa.

A rare diving beetle from Baltic amber:Hydrotrupes prometheusnew species reveals former widespread distribution of the genus (Coleoptera, Dytiscidae)

A new species in the extant genus,HydrotrupesSharp,H. prometheusn. sp., (Coleoptera, Dytiscidae) is described from Baltic amber (Eocene) based on examination of a single female specimen. This description represents one of a limited number of diving beetle taxa described from fossiliferous amber and is currently the oldest known fossil assigned to Agabinae. Based on this specimenHydrotrupesis at least 40 million years old. The occurrence ofHydrotrupesduring the Eocene suggests that the current disjunct distribution of livingHydrotrupesspecies occurring in western North America and eastern China is a relict of former widespread distribution in the northern continents. Considering the age of this fossil and its similarity to living members of the genusHydrotrupes, the conservation of morphology in this lineage of diving beetles is notable. Key morphological characters of the new species are illustrated, and the significance of this discovery for understanding Agabinae evolution and the biogeography of this previously hypothesized trans-Beringian lineage is discussed.

The earliest colubroid-dominated snake fauna from Africa: perspectives from the Late Oligocene Nsungwe Formation of southwestern Tanzania.

The extant snake fauna has its roots in faunal upheaval occurring across the Paleogene - Neogene transition. On northern continents, this turnover is well established by the late early Miocene. However, this transition is poorly documented on southern landmasses, particularly on continental Africa, where no late Paleogene terrestrial snake assemblages are documented south of the equator. Here we describe a newly discovered snake fauna from the Late Oligocene Nsungwe Formation in the Rukwa Rift Basin of Tanzania. The fauna is small but diverse with eight identifiable morphotypes, comprised of three booids and five colubroids. This fauna includes Rukwanyoka holmani gen. et sp. nov., the oldest boid known from mainland Africa. It also provides the oldest fossil evidence for the African colubroid clade Elapidae. Colubroids dominate the fauna, comprising more than 75% of the recovered material. This is likely tied to local aridification and/or seasonality and mirrors the pattern of overturn in later snake faunas inhabiting the emerging grassland environments of Europe and North America. The early emergence of colubroid dominance in the Rukwa Rift Basin relative to northern continents suggests that the pattern of overturn that resulted in extant faunas happened in a more complex fashion on continental Africa than was previously realized, with African colubroids becoming at least locally important in the late Paleogene, either ahead of or as a consequence of the invasion of colubrids. The early occurrence of elapid snakes in the latest Oligocene of Africa suggests the clade rapidly spread from Asia to Africa, or arose in Africa, before invading Europe.

Structure of the North Kara Shelf from results of seismostratigraphic analysis

The sedimentary cover section of the North Kara Shelf had been subdivided based on extensive seismic data, and seismic correlation was carried out. The predominant role of Upper Riphean-Middle Paleozoic rocks has been corroborated. A series of relatively deepwater basins filled with primarily terrigenous fly-schoid rocks up to 7–9 km in thickness existed in the Late Riphean-Vendian at the place of the shelf. In the Cambrian, isolated basins merged into a wide and shallow-water basin as a result of the Baikalian reactivation in southeast Severnaya Zemlya and probably in Taimyr. After the pre-Ordovician hiatus, a vast sedimentation basin with a regressive section was formed on the shelf from Ordovician to Late Devonian. Shallow-water marine and near-shore carbonate and carbonate-terrigenous sequences accumulated in this basin and gave way to continental and less frequent near-shore, marine, variegated, and red beds in the Devonian. The thickness of the Ordovician-Devonian sequence reaches 6 km. Since the Mid-Carboniferous, block emergence and deep erosion of Ordovician-Devonian complexes have occurred in the north Kara shelf as a result of Hercynian events in northern Taimyr, Severnaya Zemlya, and in the southern Kara Sea. No Permian-Triassic rifts existed on the North Kara Shelf. At that time, the shelf was an area of erosion. The thickness of the Middle Carboniferous-Cretaceous sequence was insignificant and gradually increased toward Barents Sea troughs. The newly formed Svyataya Anna and Voronin troughs arose due to opening of the Eurasia Basin of the Arctic Ocean. The terrane concept has been subjected to criticism. The available data show that a large epi-Grenvillian continental block existed, and the North Kara region was part of it. Collision of the northern continent with the Paleosiberian Platform in the Late Paleozoic resulted in the formation of the Hercynian fold arc to the south of the North Kara Shelf, and an inverted orogenic arch arose at the place of shelf basin. The individual geological features that distinguish the North Kara Shelf from the Barents Sea troughs and the South Kara Syneclise are emphasized. The ancient pre-Riphean basement, a system of Late Riphean-Vendian relatively deepwater troughs and basins, Hercynian tectonic inversion, deep erosion of the most uplifted part of the arch, and significant block motions are the distinguishing features of the North Kara Shelf.

Multi-year observations of the spatial and vertical distribution of aerosols and the genesis of abnormal variations in aerosol loading over the Arabian Sea during Asian summer monsoon season

Intense aerosol plumes engulf a vast region of the Arabian Sea during the Asian summer monsoon season (ASM: June–September). The largest value of aerosol optical depth (AOD) at 550nm in this region generally occurs in July when the mean AOD attains its annual peak value of >0.8. However, the AOD over this region is abnormally large during the ASM in some years, especially during the June–July period. Long term satellite observations using MODIS reveal that the largest AOD during the 11-year period of March 2000–February 2011 occurred in June 2008 with a regional mean AOD of 1.1, which was ~97% larger than that of the corresponding long term mean value in June and ~49% larger than that in July. The availability of CALIPSO data since June 2006 provides a unique opportunity to quantify the role of continental aerosols transported from the West Asian desert regions in the genesis of the above abnormality over the Arabian Sea. We examine the spatial and vertical distributions of aerosols over the Arabian Sea and adjoining continents using multi-year data from MODIS and CALIPSO and explore the genesis of the above abnormal enhancement in AOD. The observed anomalies in AOD are substantially larger than that can be attributed to changes in wind-generated sea salt aerosols. The CALIPSO observations show that the anomalous enhancement in aerosol loading over the Arabian Sea during June 2008 was primarily caused by an enhancement in aerosol abundance in the altitude range of ~1–4km with a distinctly large volume depolarization ratio of >0.25, clearly indicating the dominance of highly non-spherical mineral dust. Although the aerosol loading over the Arabian Sea during the ASM is observed to be mainly caused by the mineral dust transported from the West Asian Deserts at northwest of the Arabian Sea, the abnormal enhancement in the observed AOD during June 2008 was primarily caused by a distinct increase in dust storms over the northern continents and subsequent transport into the Arabian Sea.

Molecular phylogenetic reconstructions identify East Asia as the cradle for the evolution of the cosmopolitan genus Myotis (Mammalia, Chiroptera)

Sequences of the mitochondrial cytochrome b (1140bp) and nuclear Rag 2 (1148bp) genes were used to assess the evolutionary history of the cosmopolitan bat genus Myotis, based on a worldwide sampling of over 88 named species plus 7 species with uncertain nomenclature. Phylogenetic reconstructions of this comprehensive taxon sampling show that most radiation of species occurred independently within each biogeographic region. Our molecular study supports an early divergence of species from the New World, where all Nearctic and Neotropical species plus a lineage from the Palaearctic constitute a monophyletic clade, sister to the remaining Old World taxa. The major Old World clade includes all remaining Eurasian taxa, most Oriental species, one Oceanian, and all Ethiopian species. Another lineage, including M. latirostris from Taiwan, appears at the base of these two major biogeographic clades and, because it bears nyctalodont molars, could be considered as a distinct genus. However, this molar configuration is also found in crown-group species, indicating that these dental characters are variable in the genus Myotis and may confound interpretation of the fossil record. Molecular datings suggest an origin of all recent Myotis in the early Miocene (about 21MYA with 95% highest posterior density interval 23–20MYA). This period was characterized by a global climatic cooling that reduced the availability of tropical habitats and favoured the development of more temperate vegetation. This sharp climatic change might have triggered the evolution of Myotis in the Northern continents, because Myotis ancestors seem to have been well adapted and successful in such temperate habitats. Ancestral area reconstructions based on the molecular phylogeny suggest that the eastern portion of the Asian continent was an important center of origin for the early diversification of all Myotis lineages, and involved relatively few subsequent transcontinental range expansions.

Cold winter extremes in northern continents linked to Arctic sea ice loss

The satellite record since 1979 shows downward trends in Arctic sea ice extent in all months, which are smallest in winter and largest in September. Previous studies have linked changes in winter atmospheric circulation, anomalously cold extremes and large snowfalls in mid-latitudes to rapid decline of Arctic sea ice in the preceding autumn. Using observational analyses, we show that the winter atmospheric circulation change and cold extremes are also associated with winter sea ice reduction through an apparently distinct mechanism from those related to autumn sea ice loss. Associated with winter sea ice reduction, a high-pressure anomaly prevails over the subarctic, which in part results from fewer cyclones owing to a weakened gradient in sea surface temperature and lower baroclinicity over sparse sea ice. The results suggest that the winter atmospheric circulation at high northern latitudes associated with Arctic sea ice loss, especially in the winter, favors the occurrence of cold winter extremes at middle latitudes of the northern continents.

Late Paleocene–early Eocene carbon isotope stratigraphy from a near-terrestrial tropical section and antiquity of Indian mammals

Late Paleocene to early Eocene (~56 to 51 Ma) interval is characterized by five distinct transient warming (hyperthermal) events (Paleocene–Eocene thermal maximum (PETM), H1/ETM2/ELMO, H2, I1 and I2) in a super greenhouse globe associated with negative carbon isotope excursions (CIEs). Although well-documented marine records exist at different latitudes, terrestrial PETM sections are rare. In particular, almost no terrestrial records of either the PETM or early Eocene hyperthermals (EEHs) are yet available from the tropics. Further, evolution of modern order of mammals near the PETM has been recorded in many northern continents; however, the response of mammals in the tropics to these warming events is unknown. A tropical terrestrial record of these hyperthermal/CIE events, encompassing the earliest modern order mammal bearing horizon from India, can therefore be vital in understanding climatic and biotic evolution during the earliest Cenozoic time. Here, for the first time, we report high resolution carbon isotope (δ 13C) stratigraphy, nannofossil, and Sr isotope ratio of marine fossil carbonate from the Cambay Shale Formation of Western India. The record shows complete preservation of all the above CIE events, including the PETM, hitherto unknown from the equatorial terrestrial records. δ 13C chemostratigraphy further suggests that at least the present early Eocene mammal-bearing horizon, recently discovered at Vastan, does not support the ‘out of India’ hypothesis of earliest appearance of modern mammals and subsequent dispersal to the Holarctic continents.

Analysis of IASI tropospheric O<sub>3</sub> data over the Arctic during POLARCAT campaigns in 2008

Abstract. Ozone data retrieved in the Arctic region from infrared radiance spectra recorded by the Infrared Atmospheric Sounding Interferometer (IASI) on board the MetOp-A European satellite are presented. They are compared with in situ and lidar observations obtained during a series of aircraft measurement campaigns as part of the International Polar Year POLARCAT activities in spring and summer 2008. Different air masses were sampled during the campaigns including clean air, polluted plumes originating from anthropogenic sources, forest fire plumes from the three northern continents, and stratospheric-influenced air masses. The comparison between IASI O3 [0–8 km], [0–12 km] partial columns and profiles with collocated aircraft observations is achieved by taking into account the different sensitivity and geometry of the sounding instruments. A detailed analysis is provided and the agreement is discussed in terms of vertical sensitivity and surface properties at the location of the observations. Overall, IASI O3 profiles are found to be in relatively good agreement with smoothed in situ and lidar profiles in the free troposphere with differences of less than 40% (25% over sea for both seasons) and 10%, respectively. The correlation between IASI O3 retrieved partial columns and the smoothed aircraft partial columns is good with DC-8 in situ data in spring over North America (r = 0.68), and over Greenland with ATR-42 lidar measurements in summer (r = 0.67). Correlations with other data are less significant highlighting the difficulty of IASI to capture precisely the O3 variability in the Arctic upper troposphere and lower stratosphere (UTLS). This is particularly noted in comparison with the [0–12 km] partial columns. The IASI [0–8 km] partial columns display a low negative bias (by less than 26% over snow) compared to columns derived from in situ measurements. Despite the relatively high biases of the IASI retrievals in the Arctic UTLS, our analysis shows that IASI can be used to identify, using O3 / CO ratios, stratospheric intrusions.

Influence of Asian continental outflow on the regional background ozone level in northern South China Sea

Asian continental outflow is known to have very pronounced seasonality, which in turn can alter the trace gas contents of the Northern Hemisphere. In this study, field measurements were conducted during the spring of 2010, Dongsha, a small island situated between Taiwan and the Philippines, served as a remote site for monitoring surface ozone. Ozone was used as an effective indicator to distinguish between continental and marine air masses. Our measurements suggested that strong northeasterly winds arising from the winter Asian monsoon may have transported polluted air masses from the northern continent to locations as far south as Dongsha (latitude 20.70°N), as indicated by elevated ozone levels of approximately 60 ppbv. In contrast, during periodic calm periods when the monsoon subsided low ozone levels of about 30 ppb were detected, which is typical for marine air masses. This outflow of polluted air masses from the Asian continent and Taiwan facilitated by the winter monsoon was also successfully simulated using the Pennsylvania State University/National Center for Atmospheric Research Mesoscale Model, version 5 (PSU/NCAR MM5) and the Taiwan Air Quality Model (TAQM).