Nomenclatural Combinations Research Articles

Таксономический обзор видов секции Candidae рода Stachys (Lamiaceae)

A taxonomic review of the Stachys L. section Сandidae R. Bhattacharjee is given. The keys for determination of the species and vareties are made. The section comprises 5 species cllassified into 2 subsections: Сandidae and a new monotypic subsection Stenophyllae Krestovsk. Two new nomenclatural combinations in the rank of varieties: Stachys iva Griseb. var. macedonica (Micevski) Krestovsk. and S. spreitzenhoferi Heldr. var. virella (D. Perss.) Krestovsk. are published, the latter in a new rank. The presented information includes main synonyms, type citations, ecology and distribution data. S. candida Bory et Chaub., S. chrysantha Boiss. et Heldr., S. spreitzenhoferi Heldr. are endemics of the southern Greece, S. iva Griseb. is endemic of the northern Greece and Republic of Macedonia, S. saxicola Coss. et Balansa — endemic of Marocco and Algeria.

“Senecio ucranicus” Besser (1822) versus Senecio ucranicus Hodálová (1999) и Jacobaea borysthenica (Asteraceae): номенклатурное уточнение

It is demonstrated that the name “Senecio ucranicus” has never been validated by Besser (1822) because he did not accept that name; moreover, this binomial is non-existing because the word “ucranicus” was not intended as a species epithet. The unnamed species No. 1082 discussed and compared to some other taxa by Besser in his Enumeratio… is identifiable as the currently recognized species Jacobaea borysthenica(Senecio borysthenicus). Consequently, the name S. ucranicus Hodálová (1999) proposed for a Carpathian species of Senecio s. str. is legitimate, and the name S. hercynicus Herborg subsp. ucranicus should be considered a nomenclatural combination with the authorship “(Hodálová) Greuter”. Synonymy of Jacobaea grandidentata is briefly discussed and updated.

First Record of Chrysococcus rufescens G.A. Klebs f. tripora J.W.G. Lund (Chrysophyta) in Ukraine

Mass development of Chrysococcus rufescens G.A. Klebs f. tripora J.W.G. Lund (Chrysophyta) new for the algal flora of Ukraine was observed in studies of phytoplankton of the ponds of the Goloseyevo National Natural Park. The peculiarities of the morphological structure and quantitative indices of the development of this species were investigated. Its original figures, microphotographs, and description, and also characteristics of the environmental conditions, are given in the paper. A new nomenclatural combination Chrysococcus lundianus Lilitskaya et Klochenko comb. nov. was put forward.

Новая комбинация в роде Thymus (Lamiaceae) флоры Балканского полуострова

A new nomenclatural combination for an endemic of the Balkan Peninsula is published: Thymus urumovii (Velen.) Vasjukov.

A new generic system for the pantropical Caesalpinia group (Leguminosae).

The Caesalpinia group is a large pantropical clade of ca. 205 species in subfamily Caesalpinioideae (Leguminosae) in which generic delimitation has been in a state of considerable flux. Here we present new phylogenetic analyses based on five plastid and one nuclear ribosomal marker, with dense taxon sampling including 172 (84%) of the species and representatives of all previously described genera in the Caesalpinia group. These analyses show that the current classification of the Caesalpinia group into 21 genera needs to be revised. Several genera (Poincianella, Erythrostemon, Cenostigma and Caesalpinia sensu Lewis, 2005) are non-monophyletic and several previously unclassified Asian species segregate into clades that merit recognition at generic rank. In addition, the near-completeness of our taxon sampling identifies three species that do not belong in any of the main clades and these are recognised as new monospecific genera. A new generic classification of the Caesalpinia group is presented including a key for the identification of genera, full generic descriptions, illustrations (drawings and photo plates of all genera), and (for most genera) the nomenclatural transfer of species to their correct genus. We recognise 26 genera, with reinstatement of two previously described genera (Biancaea Tod., Denisophytum R. Vig.), re-delimitation and expansion of several others (Moullava, Cenostigma, Libidibia and Erythrostemon), contraction of Caesalpinia s.s. and description of four new ones (Gelrebia, Paubrasilia, Hererolandia and Hultholia), and make 75 new nomenclatural combinations in this new generic system.

Los coniopterígidos de la Península Ibérica e Islas Baleares (Insecta: Neuropterida, Neuroptera: Coniopterygidae)

Se revisan las especies de coniopterígidos presentes en la fauna de la Península Ibérica e Islas Baleares donde, hasta la fecha, están representadas por 39 especies pertenecientes a 8 géneros. Partiendo de los datos generales conocidos, damos una diagnosis de la familia y citamos algunos elementos sobre su historial paleontológico y su distribución, y algunos datos sobre su morfología, su biología, sus estadios juveniles y comportamiento, y se comenta una breve introducción sobre el historial en el conocimiento de la familia, y en particular en la Península Ibérica y Baleares. Se propone revalidar a Coniopteryx (Holoconiopteryx) tullgreni Tjeder, 1930 sp. reval. como especie válida, se describe Helicoconis bachi n. sp. como nueva especie, y se mantiene, se discute y se argumenta nuestro criterio sobre la validez de Coniopteryx parthenia (Nav.s & Marcet, 1910) (sensu Meinander, 1972a) frente a Coniopteryx hoelzeli (sensu Günter, 1993). Se incluye una lista de las especies pertenecientes a la fauna de la Península Ibérica y Baleares, una clave de identificación de los imagos de las subfamilias, géneros y especies citadas, y una clave de sus larvas conocidas a nivel genérico. De cada una de estas especies se recopilan y se anotan todas las referencias bibliográficas existentes relativas a las especies de la zona estudiada, tanto con la denominación actualmente aceptada, como de sus sinonimias y/o cualquier otra combinación nomenclatural bajo las cuales hayan sido citadas. También de cada una de las especies se anotan los datos conocidos y los ahora aportados (1.800 nuevos ejemplares de 32 especies) sobre su distribución general, aspectos de su morfología externa y/o genital, as. como de su variabilidad, estadios juveniles, biolog.a, fenolog.a y distribuci.n altitudinal y geogr.fica en la Península y archipi.lago balear, en base a un total de 12.215 ejemplares estudiados.

Nomenclatural Adjustments in Phacelia sect. Glandulosae (Hydrophyllaceae, Boraginales)

Abstract. Three taxa are recognized at the rank of species based on molecular phylogenetic studies in Phacelia sect. Glandulosae (Hydrophyllaceae, Boraginales). Results of those studies did not support a monophyletic Phacelia crenulata Torr. ex S. Watson, because 3 varieties of that species were supported as distinct lineages based on molecular (nuclear and chloroplast) and morphological characters. Typification and status is clarified for P. corrugata A. Nelson and P. orbicularis Rydb. A new nomenclatural combination for P. crenulata var. angustifolia N.D. Atwood is established to elevate that variety to species rank, as Phacelia angustifolia (N.D. Atwood) Walden, comb. et stat. nov.

A new combination in Angraecoides (Angraecinae, Orchidaceae): a forgotten name

A new nomenclatural combination for the African species Angraecum moandense to the recently proposed genus Angraecoides is provided. Angraecum chevalieri is confirmed as a synonym of this species.

What is Ceratozamia mexicana (Zamiaceae)?

Explorations in central Veracruz on the Transvolcanic Mexican Belt and regions explored by 19th century botanists we found various populations of Ceratozamia whose taxonomic status is not yet completely clear. Especially two populations of C. mexicana with distinct morphology; one population at El Esquilón considered as C. mexicana by Chamberlain and other authors, and another population at El Mirador that Stevenson found a voucher at the herbarium in Paris with oblanceolate leaflets collected by Ghiesbrecht and designated it as Brongniart’s holotype. The plants from El Mirador are comparable to the holotype of C. mexicana , but those at El Esquilón do not. Stevenson later also found a voucher at Kew by Thiselton-Dyer as C. mexicana var. tenuis from Mexico with linear-lanceolate leaflets that is similar to the plants at the El Esquilón population. An analysis of 11 leaflet anatomical characters gave a significant discriminant pattern (Wilk’s lambda 0.00015 P < 0.01). The squared Mahalanobis distances were also significant ( P < 0.05) and the first canonic variable showed that 93.4 % of the variation was due to the anatomical characters. A new nomenclatural combination is proposed and Thiselton-Dyer’s voucher assigned as the lectotype of Ceratozamia tenuis . Epitypes were also designated for the two species in support of the anatomical analysis as well as a description for both species given.

Organellar phylogenomics of an emerging model system: Sphagnum (peatmoss).

Sphagnum-dominated peatlands contain approx. 30 % of the terrestrial carbon pool in the form of partially decomposed plant material (peat), and, as a consequence, Sphagnum is currently a focus of studies on biogeochemistry and control of global climate. Sphagnum species differ in ecologically important traits that scale up to impact ecosystem function, and sequencing of the genome from selected Sphagnum species is currently underway. As an emerging model system, these resources for Sphagnum will facilitate linking nucleotide variation to plant functional traits, and through those traits to ecosystem processes. A solid phylogenetic framework for Sphagnum is crucial to comparative analyses of species-specific traits, but relationships among major clades within Sphagnum have been recalcitrant to resolution because the genus underwent a rapid radiation. Herein a well-supported hypothesis for phylogenetic relationships among major clades within Sphagnum based on organellar genome sequences (plastid, mitochondrial) is provided. We obtained nucleotide sequences (273 753 nucleotides in total) from the two organellar genomes from 38 species (including three outgroups). Phylogenetic analyses were conducted using a variety of methods applied to nucleotide and amino acid sequences. The Sphagnum phylogeny was rooted with sequences from the related Sphagnopsida genera, Eosphagnum and Flatbergium Phylogenetic analyses of the data converge on the following subgeneric relationships: (Rigida (((Subsecunda) (Cuspidata)) ((Sphagnum) (Acutifolia))). All relationships were strongly supported. Species in the two major clades (i.e. Subsecunda + Cuspidata and Sphagnum + Acutifolia), which include >90 % of all Sphagnum species, differ in ecological niches and these differences correlate with other functional traits that impact biogeochemical cycling. Mitochondrial intron presence/absence are variable among species and genera of the Sphagnopsida. Two new nomenclatural combinations are made, in the genera Eosphagnum and Flatbergium Newly resolved relationships now permit phylogenetic analyses of morphological, biochemical and ecological traits among Sphagnum species. The results clarify long-standing disagreements about subgeneric relationships and intrageneric classification.

Los crisópidos de la Península Ibérica y Baleares (Insecta, Neuropterida, Neuroptera: Chrysopidae)

The present contribution reviews the green lacewing fauna of the Iberian Peninsula and Balearic Islands where, to date, 50 species belonging to 13 genera are known to occur. After a diagnosis of the family Chrysopidae, its paleontological history and distribution, as well as some general data on its morphology and biology, juvenile stages and behavior are presented. This general information is followed by a brief history of studies on the family, and particularly those on the Iberian Peninsula and Balearic Islands. Some corrections are reported for earlier publications dates that have undergone taxonomical changes. Thus, Rexa almerai (Navas, 1919) n. comb. is proposed as junior synonym of Rexa lordina Navas, 1920 n. syn.; and is included in the list of chrysopid species belonging to the fauna of the Iberian Peninsula and Balearic Islands. A key for identifying imagoes of the subfamilies, genera, and species recorded from the region and a generic level key for the larvae are presented. For these species all existing bibliographic references since 1972, are compiled and annotated, both the currently accepted names, their synonyms and/or any other nomenclatural combinations under which they have been recorded. New information is presented on the external and/or genital morphology of most species, and detailed information is compiled on the presently known general distribution of each species with new data (299 new specimens of 25 species). Moreover, the variability, juvenile stages, biology, phenology, and altitudinal and geographic distribution of species from the Peninsula and Balearic Archipelago are assessed on the basis of 20.667 studied specimens.

A taxonomic study of the genus Hellenocarum H.Wolff (Umbelliferae-Apioideae)based on morphology, fruit anatomy, and molecular data

Phylogenetic relationships among the species of Hellenocarum and its close allies (Umbelliferae-Apioideae) were investigated using nuclear (ITS, ETS) and plastid (psbA-trnH intergenic spacer) DNA sequences. The results obtained were supplemented with an examination of morphology from herbarium and field-collected materials, as well as details of fruit anatomy obtained from light and scanning electron microscopy. According to the molecular data, Hellenocarum is not monophyletic. In most molecular analyses, the genus comprises 2 disparate lineages, with each lineage supported by distinct morphological characters (e.g., structure of underground organs, shape of the umbel, petal color). The first lineage includes Hellenocarum multiflorum (the type species) and H. Strictum and corresponds to Hellenocarum sensu stricto. The second lineage includes Hellenocarum amplifolium and H. Pisidicum and is recognized herein as the new genus Neomuretia Kljuykov, Degtjareva and Zakharova. Carum depressum is united with Hellenocarum sensu stricto. Three new nomenclatural combinations are proposed: Hellenocarum depressum (Hartvig and Kit Tan) Kljuykov and Zakharova; Neomuretia amplifolia (Boiss. And Hausskn.) Kljuykov, Degtjareva and Zakharova; and Neomuretia pisidica (Kit Tan) Kljuykov, Degtjareva and Zakharova.

Generic delimitations, biogeography and evolution in the tribe Coleeae (Bignoniaceae), endemic to Madagascar and the smaller islands of the western Indian Ocean

This study presents the most complete generic phylogenetic framework to date for the tribe Coleeae (Bignoniaceae), which is endemic to Madagascar and the other smaller islands in the western part of the Indian Ocean. The study is based on plastid and nuclear DNA regions and includes 47 species representing the five currently recognized genera (including all the species occurring in the western Indian Ocean region). Bayesian and maximum likelihood analyses supported (i) the monophyly of the tribe, (ii) the monophyly of Phylloctenium, Phyllarthron and Rhodocolea and (iii) the paraphyly of Colea due to the inclusion of species of Ophiocolea. The latter genus was also recovered paraphyletic due to the inclusion of two species of Colea (C. decora and C. labatii). The taxonomic implications of the mutual paraphyly of these two genera are discussed in light of morphological evidence, and it is concluded that the two genera should be merged, and the necessary new nomenclatural combinations are provided. The phylogenetic framework shows Phylloctenium, which is endemic to Madagascar and restricted to dry ecosystems, as basal and sister to the rest of the tribe, suggesting Madagascar to be the centre of origin of this clade. The remaining genera are diversified mostly in humid ecosystems, with evidence of multiple dispersals to the neighboring islands, including at least two to the Comoros, one to Mauritius and one to the Seychelles. Finally, we hypothesize that the ecological success of this tribe might have been triggered by a shift of fruit-dispersal mode from wind to lemur.

The genus Tremella (Tremellales, Basidiomycota) in Russia with description of two new species and proposal of one nomenclatural combination

Twenty-two non-lichenicolous species of Tremella are reported from Russia. Detailed descriptions, photographs, illustrations and a dichotomous key for the identification of taxa are provided. Two species, Tremella subalpina and T. griseola, are described as new. One new combination is proposed, and an epitype for Tremella yokohamensis is designated. Molecular phylogenetic reconstructions are presented based on nrITS and nrLSU regions in Bayesian and likelihood analyses. Phylogenetic data were strongly correlated with morphological data and were useful to delimit closely related species.

New combinations in Neotropical Thelypteridaceae.

288 new combinations of Neotropical Thelypteridaceae taxa are proposed in order to recognize monophyletic genera, based on the results of the most recent molecular phylogeny of the family, as well as the morphological uniformity of characters for each genus. The new nomenclatural combinations correspond to 186 Amauropelta taxa, 77 species of Goniopteris, and 25 Steiropteris taxa. A key to all native Neotropical genera of the family is also presented.

AlyBase: database of names, chromosome numbers, and ploidy levels of Alysseae (Brassicaceae), with a new generic concept of the tribe

Database of published chromosome numbers and ploidy-level estimates of the tribe Alysseae is presented, together with the revised generic concept and the list of accepted names, to reflect the most recent taxonomic and phylogenetic studies in Alysseae. It is available on-line at www.alysseae.sav.sk . The tribe encompasses 24 genera and 277 species. Chromosome numbers and/or ploidy levels are known for 171 out of 297 recognized taxa. Of these, 95 (55.6 %) taxa are diploids, 43 (25.1 %) are polyploids, and 33 (19.3 %) involve both diploids and polyploids. The most common base chromosome number in the tribe is x = 8 and less frequent is x = 7. The highest variation in base chromosome numbers (x = 7, 8, 11, 15) is found in the genus Hormathophylla. A key to all genera and descriptions of the two new genera Cuprella and Resetnikia are presented. Many new nomenclatural combinations, mainly in the re-established Odontarrhena (77), are proposed.

Can a rainbow consist of a single colour? A new comprehensive generic arrangement of the ‘Iris sensu latissimo’ clade (Iridaceae), congruent with morphology and molecular data

When treated in a broad sense, Iris (the ‘rainbow-flowers’) is one of the most diverse and well-known genera in the Asparagales. However, recent conventional phylogenetic and three-taxon statement re-treatments of the molecular data (cpDNA) for the irises (‘Iris sensu latissimo’ clade) showed that the obtained patterns of relationships appeared to be fully congruent to the narrow taxonomical arrangement of Iris (sensu stricto) to include only the bearded irises. Given this, we propose a new taxonomic arrangement of the rainbow-flowers with at least 25 previously recognised infrageneric taxa here accepted at the generic rank, of which 19 have already been treated as separate genera by different authors in the past. Morphological, phytochemical, karyological, distributional and molecular data are discussed which support the newly proposed system. Five genera and one section are described as new, 1 genus is amended, and 114 new nomenclatural combinations (2 genera, 3 sections, 1 series, 86 species, 14 varieties and 8 nothospecies) are established to accommodate the accepted names to the new generic system. Morphological descriptions and nomenclatural types are reported for each accepted genus, with types of four genera being designated here for the first time. Tentative distribution maps of relevant taxa, and illustrations of the most reliable morphological characters are included for the accepted genera. Principal synonyms and publication details are shown, and accepted infrageneric taxa are only referred to when previous information is not available. A key is also reported for genera identification. Our proposal mostly accords with the traditional distinction of groups currently in use by horticulturists, and it is favoured against an alternative treatment of an expanded Iris which renders a highly heterogeneous genus.

New combinations for East European species of Sabulina (Caryophyllaceae)

Recent molecular studies (Greenberg & Donoghue 2011, Dillenberger & Kadereit 2014) clearly demonstrated that the genus Minuartia Linnaeus (1753: 89) (Caryophyllaceae Juss.) is highly polyphyletic and the taxa previously placed in that genus (e.g., McNeill 1962, Halliday 1993) represent several distinct phylogenetic lineages less related to each other than to some other readily recognized Caryophyllaceae genera. As a consequence, many taxa, which were earlier treated in Minuartia, should be now placed in several distinct genera. Dillenberger & Kadereit (2014) proposed to recognize within the former Minuartia s.l. several genera which are supported by both morphological and molecular characters. Three genera, Mcneillia Dillenberger & Kadereit (2014: 83), Minuartiella Dillenberger & Kadereit (2014: 84), and Pseudocherleria Dillenberger & Kadereit (2014: 84), were described as new ones. Resurrection and re-circumscription was proposed for previously neglected genera Cherleria Linnaeus (1753: 425), Facchinia Reichenbach (1841: 63), Mononeuria Reichenbach (1841: 118), Rhodalsine J. Gay (1845: 25), Sabulina Reichenbach (1832: 24), and Triplateia Bartling (1830: 305), as well as re-circumscription of Minuartia sensu stricto and Eremogone Fenzl (1833: 13). A key to the new proposed classification and a nomenclatural synopsis of the currently recognized species were provided as well (Dillenberger & Kadereit 2014). Accordingly other authors already validated additional nomenclatural combinations in Eremogone (Rabeler & Wagner 2015), Mcneillia (Bartolucci et al. 2015), and Sabulina (Iamonico 2014).

Recent inputs to the Lebanese orchid flora and proposal of a national checklist for Orchidaceae family

Since the publication of the last Lebanese and Syrian classical flora, the only complete work on Lebanese flora is an illustrated book based on the same taxonomy and nomenclature with a few exceptions and some additions. Specifically on the Lebanese orchids, one book has illustrated orchids in the field with an unusual taxonomic point of view. Several Euro-Mediterranean monographs propose a conflicting taxonomy that is not always useful in identifying living specimens. Several years of exploration throughout Lebanon allowed the authors to build considerable knowledge of the orchid flora and its taxonomy. Other works made on adjacent areas completed the investigations and provided some useful indications to build taxonomic understanding and compile a preliminary checklist of Orchidaceae. Historical nomenclature was reanalysed from a modern point of view; recent critical appellations were confronted with biogeography and integrative evolutionary taxonomy, and some poorly known taxa are highlighted and their taxonomy is reinvestigated. A total of 51 species and subspecies are listed for Lebanon. Two new chorological records for Lebanon (Epipactis helleborine “subsp. praecox” nom. provis.; Ophrys alasiatica) and four new nomenclatural combinations (Androrchis × ehdenica, Epipactis helleborine subsp. turcica, Ophrys episcopalis subsp. libanotica, Serapias vomeracea subsp. levantina) are presented. At least 14 taxa are regionally endemic, seven of which are limited to two border countries, the other seven being too poorly known for conclusions about their biogeography.

New nomenclature combinations in the green alder species complex (Betulaceae).

The name Alnus viridis (Chaix) DC., based on Betula viridis Chaix (1785), has traditionally been attributed to green alders although it is based on a later basionym. Alnus alnobetula (Ehrh.) K. Koch based on Betula alnobetula Ehrh. (1783) is the correct name for green alders. In light of the increasing use and recognition of the name Alnus alnobetula (Ehrh.) K. Koch in the literature. I herein propose new nomenclatural combinations to account for the Japanese and Chinese subspecies respectively: Alnus alnobetula subsp. maximowiczii (Callier ex C.K. Schneid.) J. Chery and Alnus alnobetula subsp. mandschurica (Callier ex C.K. Schneid.) J. Chery. Recent phylogenetic analyses place these two taxa in the green alder species complex, suggesting that they should be treated as infraspecific taxa under the polymorphic Alnus alnobetula.