Everyday life for undocumented Americans often entails coping with the fear, stress, and anxiety of potential deportation (Fussell, 2011; Valenzuela & Erickson 2015). Yet, despite this troubling emotional state, undocumented Americans are increasingly taking to the streets, social media, and the halls of government demanding that their rights be upheld. This article contributes to understandings of how the political participation of undocumented Americans occurs in spite of the barriers this group faces. Through a comparative analysis of Catholic parishes in Los Angeles, California, and Albuquerque, New Mexico, the author found that a sense of belonging and expanded opportunities to participate in political and civic activities supported undocumented churchgoers as they set aside deportation fear to participate in local public life. This study builds on Verba et al.’s (1995) model of political participation which underscores the potential of churches to serve as political mobilizers. The author also argues that belonging and the provision of opportunities to participate both at and through the church should be incorporated into future models of political participation among undocumented Americans.