Mixedwood Stands Research Articles

Long term aboveground litterfall production in boreal jack pine (Pinus banksiana) and black spruce (Picea mariana) stands along the Boreal Forest Transect Case Study in western central Canada

Boreal Forest Transect Case Study with a natural climate gradient resulting from latitudinal variation provides a spatial proxy for expected climate change, and can help to understand its impact on different ecosystem processes and how the forest species are adapting to these changes. We investigated year to year variation in aboveground litter production for 2001–2010 in 13 different mature jack pine (JP) and black spruce (BS) stands in relationship to climatic variables. Simple linear and step-wise regression analysis showed that tree species interacted differently with climate and stand variables affecting litterfall. Annual total and woody litterfall varied from 57 to 164 g·m−2.y−1 in JP and 18 to 63 g·m−2.y−1in BS. Needle litterfall accounted for about 70% of the total litter production in JP and BS and about 80% in mixed wood stands, and explained significant variation in total litter production. On regional scale, needle litter production was strongly influenced by growing degree days (R2 = 0.96). Woody litterfall decreased with winter precipitation (R2 = 0.95) in JP, and increased with productivity (R2 = 0.88) in BS. Regression models developed here are useful tools in carbon budget models.

Effects of aerial strip spraying on mixedwood stand structure and tree growth

Herbicide application in strips offers a cost-effective way to accelerate spruce growth in young mixedwood stands, while also retaining the mixedwood character of the stand. In June of 2006 data were collected to evaluate the effects of aerial strip spraying on subsequent growth of treated stands. These blocks had been planted in 1991 and treated in 2000 with Triclopyr ester herbicide applied in strips. Treated and untreated strips averaged 5.8 m and 8.4 m in width, respectively, with 38% of the block area being effectively treated. In 2006 aspen density, DBH, and height were significantly lower in the treated strips compared to either untreated strips or completely untreated blocks. Spruce growth was significantly larger in treated strips with diameter growth responses beginning in the first and height growth responses in the second growing season following treatment. Yield estimates from the Mixedwood Growth Model (MGM) indicate that the strip spray treatments result in the highest spruce yields. However, when understory protection (harvesting aspen at age 70 and spruce at 120) is applied, the untreated blocks provided higher total yield and net present value.

Individual-level analysis of damage to residual trees after single-tree selection harvesting in northern Japanese mixedwood stands

Quantifying tree damage and mortality caused by single-tree selection harvesting is critical to understanding postharvest forest dynamics and management. In this study, we quantified the effects of tree size and species and the distance from residual trees to felled trees and skid trails on damage to residual trees and mortality in mixed coniferous–broadleaved stands of Hokkaido, northern Japan. Among the 4,961 trees that we studied, 373 (7.5 %) were damaged, and 148 of these trees (3.0 %) died during or immediately after logging. Hierarchical Bayesian modeling showed that the risk of damage to residual trees increased with increasing size of the felled trees and with increasing proximity to felled trees and skid trails. Smaller residual trees had the greatest risk of damage. Species differed in their susceptibility to damage; Abies sachalinensis (Fr. Schm.) Masters and Picea jezoensis (Sieb. et Zucc.) Carr. were the most susceptible species in our sample plots. Smaller damaged trees had the highest risk of mortality. The damaged trees that did not die at the time of logging had a higher risk of postharvest mortality than undamaged trees. Our results indicate that, to minimize logging-induced damage and mortality: (1) the spatial arrangement of skid trails should be optimized and fixed, (2) the risk of skidding damage should be considered before marking the trees to be harvested and choosing the felling direction, and (3) logging operators should be instructed to avoid damaging small trees, especially those of the species that are most susceptible to damage.

Growth Response of Northern White-Cedar (Thuja occidentalis) to Natural Disturbances and Partial Cuts in Mixedwood Stands of Quebec, Canada

Northern white-cedar (Thuja occidentalis) is a species of high commercial and ecological value, the abundance of which has been declining since the middle of the 19th century. Very little information regarding its silviculture in mixedwood stands is currently available, even though a significant portion of wood resources comes from these stands. The present study is a retrospective analysis of white-cedar growth in partially harvested mixedwood stands of western Quebec, Canada. Eight stands distributed across two regions were analyzed. Dendrochronological approaches examined long-term diameter growth for sample white-cedar trees and stems of associated species. These approaches were used to reconstruct stand characteristics at the time of harvesting, together with local harvesting intensity. The study demonstrated white-cedar’s capacity to maintain good growth for long periods of time and at large tree sizes. Accession to the upper canopy positions occurs through repeated episodes of suppression/release, most of which seem to be associated with spruce budworm (Choristoneura fumiferana) outbreaks. White-cedar response to partial harvesting varies with tree size, residual basal area and species composition. Growth response was generally stronger for small trees, even though large trees still maintained the best diameter growth. Growth of white-cedar was negatively affected by an increase in softwood proportion in basal area. Growth responses to harvesting could be sustained for a period of 20 years.

Silvicultural Rehabilitation of Cutover Mixedwood Stands

We investigated rehabilitation of mixedwood stands degraded by exploitative cutting on the Penobscot Experimental Forest in Maine. Three precommercial rehabilitation treatments were applied: control (no rehabilitation), moderate rehabilitation (crop tree release [CTR]), and intensive rehabilitation (CTR, timber stand improvement [TSI], and red spruce fill planting). Crop trees (primarily red maple, paper birch, spruce, aspen, and eastern hemlock) were selected and released based on their potential for improved growth and value, spacing, and species composition. Rehabilitation reduced sapling basal area, density, and hardwood abundance and increased crop tree diameter increment. Fill planting increased spruce stocking, but many planted seedlings were browsed. Long-term projections suggested that future stand value will repay costs of moderate rehabilitation (CTR); intensive rehabilitation (CTR-TSI-planting) as applied in this study requires greater investment than can be repaid through quality and growth improvements of low-value hardwoods and softwoods. Although simulations suggested no difference in future stand value between treated and untreated stands, improvements in composition, growth, and quality after rehabilitation will facilitate later commercial thinning and shelterwood regeneration in stands which otherwise have few management options.

Development and activity of early saproxylic fungal communities in harvested and unmanaged boreal mixedwood stands

Limited scientific information is currently available regarding saproxylic fungal communities in the boreal forest of North America. We aimed to characterize the community development, richness and activity of saproxylic fungi on fresh wood in harvested and unmanaged boreal mixedwood stands of northwestern Quebec (Canada). Fresh wood blocks (n = 480) of balsam fir (Abies balsamea (L.) Mill.) and trembling aspen (Populus tremuloides Michx.) were placed on the forest floor in a range of stand conditions (n = 24). Blocks were harvested every 6 months for up to 30 months and characterized for species composition and richness (PCR–DGGE, DNA sequencing), respiration, wood density and lignin and cellulose content. Colonization by a wide range of functional groups proceeded rapidly under different stand conditions. We detected a total of 35 different fungal operational taxonomic units, with the highest species richness at the wood block level being observed within the first 12 months. No differences in community composition were found between wood host species or among stand conditions. However, the variability in fungal communities among blocks (β diversity) was lower on trembling aspen wood compared with balsam fir and decreased over time on trembling aspen wood. Also, fungal activity (respiration and wood decomposition) increased on trembling aspen wood blocks and species richness decreased on balsam fir wood over time in partial-cut sites. The overlap in tree composition among stands, the high volume of logs and the recent management history of these stands may have contributed to the similarity of the saproxylic fungal community among stand types and disturbances.

Influence of partial harvesting and site factors on the abundance and composition of natural regeneration in the Acadian Forest of Maine, USA

Understanding the factors regulating the composition and abundance of natural regeneration in forest ecosystems is critical to sustainable management worldwide. Using a long-term silvicultural experiment in Maine, we partitioned the variation in natural regeneration and examined the contribution of overstory and understory vegetation (biotic factors), substrate and soil attributes (abiotic factors), and silvicultural treatment. We hypothesized that silvicultural treatment (single-tree selection with 5, 10, and 20-year cutting cycles and diameter-limit cutting) would be less influential than abiotic and biotic factors in structuring regeneration pattern. Overall, treatment and site (biotic and abiotic) factors accounted for 26% of the variation in regeneration species composition, highlighting stochasticity of the regeneration process. In support of our hypothesis, regeneration pattern that could be explained was primarily associated with local site factors; overstory and understory vegetation uniquely accounted for 16% of the variation in species composition. Substrate and soil attributes along with silvicultural treatment accounted for 4% of variation, and uniquely accounted for 2% and 4%, respectively. In support of earlier reports, shifts toward hardwood dominance were apparent at early stages of cohort development and were associated with high harvest intensities. Our findings suggested that overstory and understory vegetation exhibit greater control over regeneration patterns than substrate and soil attributes, or silvicultural treatment, in partially harvested Acadian Forest mixedwood stands, and also indicated the potential for composition control through selective overstory removal.

Ecosystem Responses to Partial Harvesting in Eastern Boreal Mixedwood Stands

Partial harvesting has been proposed as a key aspect to implementing ecosystem management in the Canadian boreal forest. We report on a replicated experiment located in boreal mixedwoods of Northwestern Quebec. In the winter of 2000–2001, two partial harvesting treatments, one using a dispersed pattern, and a second, which created a (400 m2) gap pattern, were applied to a 90-year-old aspen-dominated mixed stand. The design also included a clear cut and a control. Over the course of the following eight years, live tree, coarse woody debris, regeneration and ground beetles were inventoried at variable intervals. Our results indicate that all harvesting treatments created conditions favorable to balsam fir (Abies balsamea) sapling growth and trembling aspen (Populus tremuloides) sapling recruitment. However, balsam fir and trembling aspen regeneration and ground beetles response to gap cuts were closer to patterns observed in clear cuts than in dispersed harvesting. The underlying reasons for these differing patterns can be linked to factors associated with the contrasting light regimes created by the two partial harvesting treatments. The study confirms that partially harvesting is an ecologically sound approach in boreal mixedwoods and could contribute to maintaining the distribution of stand ages at the landscape level.

Stand age structural dynamics of conifer, mixedwood, and hardwood stands in the boreal forest of central Canada

To study the effects of stand development and overstory composition on stand age structure, we sampled 32 stands representing conifer, mixedwood, and hardwood stand types, ranging in ages from 72 to 201 years on upland mesic sites in northwestern Ontario. We defined the stages of stand development as: stem exclusion/canopy transition, canopy transition, canopy transition/gap dynamics, and gap dynamics. Stand age structure of conifer stands changed from bimodal, bimodal, reverse-J, and bimodal, respectively, through the stages of stand development. Mixedwood and hardwood stands revealed similar trends, with the exception of missing the canopy transition/gap dynamic stage in mixedwoods. Canopy transition/gap dynamic stage in hardwoods showed a weaker reverse-J distribution than their conifer counterparts. The results suggest that forest management activities such as partial and selection harvesting and seed-tree systems may diversify standard landscape-level age structures and benefit wildlife, hasten the onset of old-growth, and create desired stand age structures. We also recommend that the determination of old-growth using the following criteria in the boreal forest: 1) canopy breakdown of pioneering cohort is complete and stand is dominated by later successional tree species, and 2) stand age structure is bimodal, with dominating canopy trees that fall within a relatively narrow range of age and height classes and a significant amount of understory regeneration.

Distribution of white spruce lateral fine roots as affected by the presence of trembling aspen: root mapping using simple sequence repeat DNA profiling

Mixedwood forests of white spruce ( Picea glauca (Moench) Voss) and trembling aspen ( Populus tremuloides Michx.) may possess ecological advantages over monospecific white spruce stands. Belowground competition may be reduced through vertically stratified roots; facilitation of growth may occur in upper soil layers through nutrient-rich trembling aspen litterfall. These effects may incentivize white spruce to preferentially exploit upper soil layers in mixedwood stands, resulting in wider root systems. This research contrasted white spruce fine root (diameter <2 mm) distributions in organic layers of white spruce and mixedwood stands. Research occurred at the Fallingsnow Ecosystem Project site in northwestern Ontario. Eighteen plots represented mixedwood and pure white spruce stands. Trees were mapped; foliage and root samples were collected. Roots were separated by species and scanned to determine length. Simple sequence repeat DNA profiles were determined for all white spruce trees and for 45 white spruce root fragments per plot. Root and tree DNA profiles were matched; corresponding distances were calculated. Most (80%) root fragments were within 3.2 m of tree stems. Root prevalence decreased rapidly with distance. Organic layer pH was significantly less acidic in mixedwood plots, but only in one block. A subtle significant trend towards wider root distributions occurred in mixedwood stands.

Nest-site selection by old boreal forest cavity excavators as a basis for structural retention guidelines in spatially-aggregated harvests

Cavity excavators (woodpeckers, chickadees, nuthatches) are vulnerable to loss of old forest associated with traditional harvest practices. In the boreal plains of Canada, old forest excavators are retained in the short term in small (<100ha) and large (1000sha) harvests with residual trees and patches. Information on habitat selection can be used to guide “what, where, and how much” structure to retain and ensure excavators, and the secondary species that use their cavities, are conserved in harvests in the short term. We examined multi-scale nest-site use and selection by excavators in intact old upland forest and spatially-aggregated harvests with structural retention and developed guidelines for the types and spatial arrangement of trees, patches, and stands retained during forest planning (i.e. planned residuals) and harvesting (i.e. operator residuals). In both intact and harvest landscapes, live large (>35cm) and tall (>25m) trembling aspen (Populus tremuloides) with multiple (∼20) conks of a heartrot fungus (Phellinus tremulae) were the preferred cavity trees for yellow-bellied sapsucker (Sphyrapicus varius), hairy woodpecker (Picoides villosus), and pileated woodpecker (Dryocopus pileatus). Short (∼10m), broken-top aspen snags of medium (>20cm) and large (>35cm) diameter were preferred by black-capped chickadees (Poecile atricapillus) and northern flickers (Colaptes auratus), respectively. In the harvests, sapsuckers, hairy woodpeckers, and flickers selected cavity trees in clumps with other medium- and large-diameter aspen with conks. Black-capped chickadees selected trees with no live large conifers within ∼10m. Sapsuckers selected old aspen and mixedwood stands in intact landscapes. In harvest landscapes, sapsuckers selected trees in or near planned patches greater than 5ha and composed of mature and old aspen and mixedwood stands. Sapsuckers tended to nest in areas with ∼50% residual forest cover whereas flickers selected open areas of cutblocks away from planned patches. We recommend harvest operators retain several live, large (>35cmdbh) aspen with numerous (∼20) fungal conks in clumps that are adjacent to or near (<60m) large (>5ha) patches. Planned patches should have one to two thirds of their area derived from mature and old aspen and mixedwood stands and should be at least 5ha, with most greater than 15ha. These prescriptions will facilitate retention of the cavity-using assemblage of old upland forest in aggregated harvests in the short term. However, regional strategies for old forest conservation may be necessary to conserve cavity users in managed forest landscapes over the long term.

Evidence that soil fertility controls the mixing of jack pine with trembling aspen

Jack pine and trembling aspen are boreal tree species that are found growing either in naturally regenerated mono-specific stands, or in mixed-wood stands. We conducted a field survey and a manipulative field study to test the productivity–diversity hypothesis, which predicts that mixed-wood stands are more likely to occur on fertile soils, or following fertilization. We surveyed 44 mixed-wood stands and found 43 of these occurring on fertile clay deposits, and only one occurring on a nutrient poor till deposit. By contrast, the area surveyed comprised 45% clay and 55% till deposits. In a second study, we conducted a five year fertilization and brushing trial in a recently burned area dominated by jack pine saplings with patches of regenerating trembling aspen. Fertilization without brushing improved the growth and recruitment of aspen stems, but had no effect on jack pine growth and recruitment. Fertilization + brushing increased the growth of jack pine. Brushing the aspen, with or without fertilization, resulted in higher recruitment of jack pine. We conclude that soil fertility controls the mixing of jack pine with trembling aspen, that fertilization increases the likelihood of encroachment of aspen into areas formerly dominated by jack pine, and that brushing along with fertilization is necessary to promote jack pine growth.

Root Distributions of Planted Boreal Mixedwood Species on Reclaimed Saline–Sodic Overburden

Alberta’s oil sands are located in the boreal forest where surface mining requires reconstruction of these landscapes using waste saline and sodic overburden (SSOB) piles. The impact of these SSOB materials, however, on root development of planted boreal species is unknown. The objective of this study was to examine the effect of SSOB material on the root distributions of planted boreal species. Root distributions for planted mixedwood stands were measured using soil cores and compared with soil physical and chemical properties on three reclaimed sites. Soil pH ranged from 6.1 to 7.5 across all three reclaimed sites. Sodium adsorption ratio ranged from <30 in the SSOB at the youngest site to <4 at the oldest site while soil electrical conductivity ranged from <12 and <4 dS m−1 in the SSOB at the youngest and oldest site, respectively. Root length distributions were concentrated in the upper 30 cm of the soil profile and ranged from 0.96 to 7.99 cm cm−3. The roots were observed in the SSOB and accounted for 1.3% to 2.2% of the total root length in the profile. The root length density was also negatively correlated with Na and EC at all sites. The root distributions on these young reclaimed sites were similar to those from undisturbed boreal forest stands overlying saline soils, suggesting that root distributions on these reclaimed sites appear to be unaffected by the SSOB; however, further monitoring will be required as the stand matures to determine future impacts of the SSOB on forest productivity.

Influence of Populus tremuloides density on air and soil temperature

Density manipulation of overstory aspen provides increased light levels to promote growth of understory spruce, while it may also impact the air and soil temperature. We monitored temperature for three growing seasons in young mixedwood stands of variable density at two boreal locations in Alberta, Canada. Results indicate that partial aspen cover provides frost protection that may be location specific. Mean air temperature was similar or increased slightly across aspen densities during the growing season, while mean soil temperature significantly increased with decreasing overstory cover. The amount of accumulated air and soil heat over the growing season was inversely related to aspen density. The differential impact of density on temperature contributes to variation in growth relationships and may explain regional differences in competition effects.

Factors affecting northern white-cedar (Thuja occidentalis) seedling establishment and early growth in mixedwood stands

Regeneration of northern white-cedar ( Thuja occidentalis L.) is often deficient after harvesting in mixedwood stands growing on mesic sites even where browsing pressure is low. We compared the effectiveness of silviculture treatments on early regeneration of white-cedar after single-tree selection cutting (25% of basal area removed), shelterwood seed cut (50% of basal area removed), and group selection cutting (gaps of 625 m2) in three yellow birch ( Betula alleghaniensis Britt.) – softwood uneven-aged stands in Quebec, Canada. Three years after harvesting, the combination of factors that maximized abundance of white-cedar seedlings was single-tree selection cutting with artificial seeding on exposed mineral seedbeds (68.8% of plots with the presence of white-cedar). Early growth of planted white-cedar seedlings (40 cm tall) and biomass production were proportional to light availability, i.e., best under group selection cutting (mean height increment = 14.8 cm/year, mean root collar diameter increment = 3.0 mm/year). Browsing pressure has regional impacts depending on herbivore occupancy of the area. Our study was conducted near the northern limit of white-tailed deer ( Odocoileus virginianus (Zimmerman, 1780)), and deer were locally absent, while snowshoe hare ( Lepus americanus Erxleben, 1777) did not have a consistent effect on seedling abundance and early survival, limiting height growth only during the first year following planting.

Structural changes and potential vertebrate responses following simulated partial harvesting of boreal mixedwood stands

Partial harvesting, where different numbers and arrangements of live trees are retained in forest stands, has been proposed for maintaining late-successional structure and associated vertebrate species within managed boreal forests. Using the stand dynamics model SORTIE-ND, we examined 80-year patterns of structural change in response to different intensities (30–70% basal area removal) and spatial patterns (22–273 m 2 mean patch size) of harvesting. We also applied habitat models for seven late-successional vertebrates to the structural conditions present after harvesting to assess potential species responses. Partial harvesting increased understory and downed woody debris (DWD) cover and decreased overstory structure for the first 25 years after harvest, in comparison to unharvested stands, with this effect subsequently reversing as harvest-induced regeneration reached the canopy. Although harvesting enhanced long-term structural development in this regard, large trees, large snags, and large DWD all remained below unharvested levels throughout the simulation period. Harvesting also produced transient increases in early-decay DWD and ground exposure. Most changes in structural attributes increased in proportion to harvest intensity, but structural differences among harvest patterns were generally small. Dispersed harvesting induced somewhat less pronounced decreases in vertical structure, and produced more post-harvest slash, than aggregated harvesting. All seven vertebrate species decreased in abundance as harvest intensity increased from 30 to 70%. In comparison to their pre-harvest abundances in old stands, vertebrates associated with DWD (redback salamander, marten, red-backed vole) showed neutral or positive responses at one or more harvest intensities, whereas those associated with large trees and snags (brown creeper, flying squirrel) consistently exhibited substantial adverse impacts.

Factors affecting species richness of tree regeneration in mixed-wood stands of central Maine

Question: Two questions about within-stand spatial variability are addressed in this paper. How does species richness of tree regeneration respond to small-scale ecological gradients, and what effect does natural Abies balsamea abundance have on the species richness of other tree regeneration? Location: A long-term, gap-silviculture experiment, Acadian mixed-wood forest, Maine, USA. Methods: Eight stands treated with and without gap harvesting were sampled to capture sub-stand heterogeneity of understorey tree regeneration concurrently with patterning of local stand conditions. Spatial and non-spatial models were developed to test the relationships between two response variables [species richness of small (height ≥0.1 m, but <0.75 m) and large (height ≥0.75 m, but <1.4 m) regeneration] and five explanatory variables (depth to water table, percentage canopy transmittance, A. balsamea regeneration density, and overstorey basal area and species richness). Results: Despite high unexplained variance for all models, consistent associations among variables were found. Negative associations were found between: (1) the species richness of small regeneration and A. balsamea regeneration density and (2) the species richness of large regeneration and overstorey basal area. Positive associations were found between: (1) the species richness of small regeneration and both overstorey basal area and species richness and (2) the species richness of small and large regeneration and canopy transmittance. Conclusions: Promoting tree species diversity in Acadian mixed-wood stands may not be achievable through the use of gap-harvesting alone if the density of understorey Abies balsamea is not reduced either naturally or through silvicultural intervention.

Managing understory light conditions in boreal mixedwoods through variation in the intensity and spatial pattern of harvest: A modelling approach

In the context of partial harvesting, adequately managing post-harvest light conditions are essential to obtain a desired composition of tree species regeneration. The objective of this study was to determine how varying the intensity and spatial pattern of harvest would affect understory light conditions in boreal mixedwood stands of northwestern Quebec using the spatially explicit SORTIE-ND light model. The model was evaluated based on comparisons of observed and predicted light levels in both mapped and un-mapped plots. In mapped plots, reasonably accurate predictions of the overall variation in light levels were obtained, but predictions tended to lack spatial precision. In un-mapped plots, SORTIE-ND accurately predicted stand-level mean GLI (Gap Light Index) under a range of harvest intensities. The model was then used to simulate nine silvicultural treatments based on combinations of three intensities of overstory removal (30%, 45% and 60% of basal area) and three harvest patterns (uniform, narrow strips, large gaps). Simulations showed that increasing overstory removal had less impact on light conditions with uniform harvests, and a more marked effect with more aggregated harvest patterns. Whatever the harvest intensity, uniform cuts almost never created high light conditions (GLI > 50%). Gap cuts, on the other hand, resulted in up to 40% of microsites receiving GLI > 50%. Our results suggest that either a 30% strip or gap cut or a 45–60% uniform partial harvest could be used to accelerate the transition from an aspen dominated composition to a mixedwood stand because both types of cut generate the greatest proportion of moderately low light levels (e.g., 15–40% GLI). These light levels tend to favour an accelerated growth response among shade-tolerant conifers, while preventing excessive recruitment of shade-intolerant species. A better understanding of how spatial patterns of harvest interact with tree removal intensity to affect understory light conditions can provide opportunities for designing silvicultural prescriptions that are tailored to species’ traits and better suited to meet a variety of management objectives.

Effects of Postfire Salvage Logging on Deadwood-Associated Beetles

In Canada and the United States pressure to recoup financial costs of wildfire by harvesting burned timber is increasing, despite insufficient understanding of the ecological consequences of postfire salvage logging. We compared the species richness and composition of deadwood-associated beetle assemblages among undisturbed, recently burned, logged, and salvage-logged, boreal, mixed-wood stands. Species richness was lowest in salvage-logged stands, largely due to a negative effect of harvesting on the occurrence of wood- and bark-boring species. In comparison with undisturbed stands, the combination of wildfire and logging in salvage-logged stands had a greater effect on species composition than either disturbance alone. Strong differences in species composition among stand treatments were linked to differences in quantity and quality (e.g., decay stage) of coarse woody debris. We found that the effects of wildfire and logging on deadwood-associated beetles were synergistic, such that the effects of postfire salvage logging could not be predicted reliably on the basis of data on either disturbance alone. Thus, increases in salvage logging of burned forests may have serious negative consequences for deadwood-associated beetles and their ecological functions in early postfire successional forests.

Effects of fire severity on early survival and growth of planted jack pine, black spruce and white spruce

Two boreal mixedwood stands burned by the 1999 Black River wildfire in southeastern Manitoba, Canada were selected to study the effect of fire severity on early survival and growth of planted jack pine (Pinus banksiana), black spruce (Picea mariana) and white spruce (Picea glauca) seedlings. In each stand, three fire severity classes (scorched, lightly burned, and severely burned) were identified based on the degree of forest floor consumption. Fire severity was not a significant factor on mortality. No mortality difference was found among species, except for year 5 when jack pine had significantly higher mortality than both black spruce and white spruce. Jack pine and black spruce had their highest mortality in year 4, while white spruce had its highest mortality in year 1. Under natural competition, seedling growth increased with increasing fire severity. When competition was removed, fire severity did not affect seedling growth. Regardless of fire severity and competition, jack pine had better diameter and height growth than black spruce, which, in turn, grew slightly taller than white spruce. Planted seedlings faced less intense vegetation competition on severely burned plots compared to scorched or lightly burned plots. Regardless of fire severity and species, competition increased with time since planting. Our study results indicate that planting immediately after a wildfire is a viable option to establish conifer components on burned boreal mixedwood stands. Key words: fire severity, plantation, regeneration, Pinus banksiana, Picea mariana, Picea glauca