Mixed Conifer-broadleaf Forests Research Articles

Vertical stratification and effects of crown damage on maximum tree height in mixed conifer–broadleaf forests of Yakushima Island, southern Japan

We investigated vertical stratification and effects of crown damage on maximum tree height in two mixed conifer–broadleaf forests in Yakushima Island, southern Japan. In both research plots, the conifer trees dominated the upper canopy while the broadleaved trees dominated the middle to lower canopy. Most broadleaved trees were shorter than the median crown-base height (H CB) of the conifer trees. Estimates of the maximum height (H max) of the conifer trees were greater than those of the broadleaved trees. Crown damage had significant negative effects on maximum height of the conifer trees. Crown damage was observed for 72.8–88.7% of the conifer trees, and severe types of damage such as stem breakage and top die-back were the most predominant. The H max of the damaged conifer trees was 16–17% shorter than that of the intact trees and as much as 16–28% shorter than the potential maximum height estimated from the diameter–height relationship of the tallest intact trees. We inferred that crown disturbance is an important factor determining the maximum height of the canopy of the two mixed forests. Our results suggested that vertical stratification between conifer and broadleaved trees may be an important mechanism contributing to their coexistence and additive basal area of mixed forests on Yakushima Island.

Photosynthetic characteristics of Fagus sylvatica and Quercus robur established for stand conversion from Picea abies

Efforts in Europe to convert Norway spruce (Picea abies) plantations to broadleaf or mixed broadleaf-conifer forests could be bolstered by an increased understanding of how artificial regeneration acclimates and functions under a range of Norway spruce stand conditions. We studied foliage characteristics and leaf-level photosynthesis on 7-year-old European beech (Fagus sylvatica) and pedunculate oak (Quercus robur) regeneration established in open patches and shelterwoods of a partially harvested Norway spruce plantation in southwestern Sweden. Both species exhibited morphological plasticity at the leaf level by developing leaf blades in patches with an average mass per unit area (LMA) 54% greater than of those in shelterwoods, and at the plant level by maintaining a leaf area ratio (LAR) in shelterwoods that was 78% greater than in patches. However, we observed interspecific differences in photosynthetic capacity relative to spruce canopy openness. Photosynthetic capacity (A1600, net photosynthesis at a photosynthetic photon flux density of 1600μmolphotonsm−2s−1) of beech in respect to the canopy gradient was best related to leaf mass, and declined substantially with increasing canopy openness primarily because leaf nitrogen (N) in this species decreased about 0.9mgg−1 with each 10% rise in canopy openness. In contrast, A1600 of oak showed a weak response to mass-based N, and furthermore the percentage of N remained constant in oak leaf tissues across the canopy gradient. Therefore, oak photosynthetic capacity along the canopy gradient was best related to leaf area, and increased as the spruce canopy thinned primarily because LMA rose 8.6gm−2 for each 10% increase in canopy openness. These findings support the premise that spruce stand structure regulates photosynthetic capacity of beech through processes that determine N status of this species; leaf N (mass basis) was greatest under relatively closed spruce canopies where leaves apparently acclimate by enhancing light harvesting mechanisms. Spruce stand structure regulates photosynthetic capacity of oak through processes that control LMA; LMA was greatest under open spruce canopies of high light availability where leaves apparently acclimate by enhancing CO2 fixation mechanisms.

Infiltration characteristics of water in forest soils in the Simian mountains, Chongqing City, southwestern China

Spearman rank-correlation analysis and grey relational grade analysis were used to study infiltration characteristics of water in different forest soils in the Simian mountains, Chongqing City. The results indicate that the soil bulk density, contents of coarse sand, and porosity of macropores were significantly correlated with saturated hydraulic conductivity. Porosity of macropores and contents of coarse sand were positively correlated with soil saturated hydraulic conductivity and soil bulk density negatively. Based on the initial infiltration rate, the stable infiltration rate, time required for infiltration to reach a stable state, and cumulative infiltration, all of which are crucial parameters determining soil infiltration capacity, the results of grey relational grade analysis showed that the grey relational grades of the different forest soils were listed from high to low as broad-leaved forest (0.8031) > Phyllostachys pubescens forest (0.7869) > mixed conifer-broadleaf forest (0.4454) > coniferous forest (0.4039). Broadleaf forest had the best ability to be infiltrated among the four soils studied. The square roots of the coefficients of determination obtained from fitting the Horton infiltration equation, simulated in our study of forest soils, were higher than 0.950.We conclude that soils of broad-leaved forests were the best suited for infiltration processes of forestry in the Simian mountains.

Gap phase regeneration recruitment of mixed conifer-broadleaf forests in Wolong Nature Reserve

Wolong Nature Reserve is the largest reserve for protecting the endangered wild giant panda. Due to historical factors, even in many sections of the core protection area, the forests have been seriously destroyed and natural forests are poorly restored. However, the relative importance of the determinants for recruitment of communities under disturbance is rarely explored. In our study the endogenous and exterior factors in a forest gap that affect the conifer-broad-leaved mixed forest regeneration were investigated near Wuyipeng, one of the observation stations at Wolong, to explore which determinant had the greatest effect on gap regeneration and to discover the recruitment of seedling establishment in forest gaps. With a linear sampling method, environmental factors, gap characteristics and recruitment of new individuals were measured and examined in every forest gap along three sampling lines. Data of environmental factors in the gaps were collected for a Pearson correlation analysis in order to explore the disturbance and preprocessed characteristics of the gaps, using principal component analysis in SPSS. Correlation analysis was applied to further explore the relationship between changes in the gaps and the response of the regenerating seedlings. The results show that a range of natural and human disturbances affected the pattern and characteristics of the forest gaps in this area. The richness in the composition of the seedlings was higher than that of gap makers, but the order of dominance of the composition was different between seedlings and gap makers. The success of dominant species in establishing themselves was affected by different environmental factors. For instance, the establishment of Betula spp. was correlated significantly with topographic factors, while that of Abies faxoniana was affected by soil characteristics and that of Rhododendron spp. correlated significantly with topographic factors and characteristics of gap makers. Moreover, all the biodiversity indices of regenerating seedlings were significantly correlated with environmental principle components mostly reflecting soil properties. From this we can infer that soil characteristics are the most important factors affecting the regeneration of dominant species and seedling diversity under gap disturbances.

Evaluation of a mechanistic approach to mixedwood modelling

Mixed conifer–broadleaf forests (mixedwoods), covering more than a third of the productive forest landbase in BC, are highly valuable both as sources of fibre and as areas rich in biodiversity. In recognition of the multiple benefits of this forest type, management paradigms have transitioned from a focus on promoting conifer plantations in mixedwood areas to the management of intimate mixtures. The exceptionally dynamic growth properties and species interactions in mixedwood forests present a challenge for projecting the growth and development of different types of mixedwoods and their response to different silviculture systems. Here we evaluate the ability of a mechanistic forest growth model (FORECAST) to project patterns of stand growth and dynamics in two mixedwood forest types subjected to different silvicultural treatments. Model output is compared against field measurements from long-term silviculture trials in the Sub Boreal Spruce (SBS)—18 years, and Interior Cedar Hemlock (ICH)—10 years, biogeoclimatic zones in British Columbia, Canada. FORECAST was able to reproduce patterns of growth response in both mixedwood forest types with reasonable accuracy. An analysis of the simulated relative impact of light and nutrient competition on growth dynamics and treatment response is provided. Results suggest that competition for both light and nutrients are important factors in the dynamics of these mixedwood forest types and that long-term response data and modelling are required to adequately assess the rotation-length effects of treatments on stand development. The analysis described herein provides a level of confidence for the use of the model as a decision-support tool in these ecosystem types, but more validation work should be conducted across a range of different mixedwood forest types and management interventions as long-term datasets become available. Key words: FORECAST, mixedwood management, model testing, process-based model, resource competition

Effects of reforestation on the hydrological function of a small watershed in the Three Gorges Reservoir Area

For vegetation communities with hydrological function in the Three Gorges Reservoir Area, the storm event distributed hydrological model Precipitation-Runoff Modeling System (PRMS)-Storm was built based on modular modeling system developed by the US Geological Survey, and was employed to study the effects of forests on peak flows in the Xiangshuixi forest watershed in the Three Gorges Reservoir Area. The results showed that: 1) this simulation study suggested that PRMS-Storm can meet the second level national flood prediction standards of China for simulating storm events of small forest watersheds, and can issue flood forecasting; 2) hydrological functions of different vegetation communities were evaluated, and three simulation scenarios were arranged: mixed conifer-broadleaf forests (scenario 1), broad-leaved forests (scenario 2), and general forests arrangement (scenario 3); 3) the well-arranged forest scenarios can reduce over 20% of surface runoff, result in an increase of over 16% in subsurface flow, and decrease peak flow by 20.8%, 9.6%, and 18.9%, respectively. The reduction of peak flow rates was significant when rainfall peak was higher than 0.8 mm/min, especially for short-term rainfall events. In general, we found that scenarios 1 and 3 were preferable for reducing peak flow rates and volumes in the reforestation practices in the study region, and scenario 1 was better than scenario 3, so the mixed conifer-broadleaf forests had the best hydrological function.

Host effects on ectomycorrhizal fungal communities: insight from eight host species in mixed conifer–broadleaf forests

To advance our understanding of host effects on the community structure of ectomycorrhizal fungi (EMF), EMF communities were compared among different host species, genera and families in two mixed conifer-broadleaf forests in Japan. Using molecular identification methods we examined EMF root tips of eight coexisting species belonging to six genera (three families): Abies and Tsuga (Pinaceae), Betula and Carpinus (Betulaceae) and Fagus and Quercus (Fagaceae). In total, 205 EMF species were detected, and the total richness was estimated to exceed 300 species using major estimators. Of the 55 EMF species occurring three or more times, eight showed significantly biased host preference. A Mantel test showed a significantly negative correlation between EMF community similarity and host taxonomic distance. Detrended correspondence analysis separated EMF communities mainly by host taxonomic and successional status. Thus, EMF communities are similar on hosts with similar taxonomic and successional status. A significant proportion of EMF exhibited host specificity, which may contribute to the extremely diverse EMF community in conifer-broadleaf forests.

Effects of selection cutting on the longicorn and scolytid beetle assemblages in an old-growth boreal mixed conifer-broadleaf forest in Hokkaido

Effects of selection cutting on the longicorn and scolytid beetle assemblages in an old-growth boreal mixed conifer-broadleaf forest in Hokkaido

A simple and efficient method to estimate microsite light availability under a forest canopy

This study presents a new, simple, and efficient method for estimating microsite light availability in the understory of a forest. The percentage of above-canopy photosynthetic photon flux density (%PPFD) transmitted above 16 microsites in the understory of a mixed conifer–broadleaf forest was measured every minute between 07:00 and 19:00 for both a completely overcast and a cloudless day. Instantaneous measures of %PPFD were also taken at different times on 3 overcast days. The instantaneous measures of %PPFD were strongly and directly related (P < 0.001) among themselves and with mean daily %PPFD values. These results demonstrate the usefulness of using an instantaneous measure of %PPFD taken under overcast sky conditions for estimating the mean daily %PPFD at any microsite under a forest canopy.