Eocene Research Articles

Transient micropaleontological turnover across a late Eocene (Priabonian) carbon and oxygen isotope shift on Blake Nose (NW Atlantic)

Abstract. The Gulf Stream, a western boundary current transporting warm water into the North Atlantic, plays a key role in climate regulation and oceanographic stability at a regional and global scale as part of the Atlantic Meridional Overturning Circulation (AMOC). Evidence suggests that an ancestral Gulf Stream has existed since the Mesozoic, and it has altered its course repeatedly over Cenozoic times. In this study, we focus on the upper Eocene (Priabonian, ca. 36 Ma) from Ocean Drilling Program Site 1053 on Blake Nose (subtropical North Atlantic). Bulk carbon and oxygen stable isotopes, as well as benthic foraminiferal and calcareous nannofossil assemblages, provide an integrated assessment of the palaeoceanographic changes impacting the area through the water column to the seafloor. Micropaleontological assemblages suggest changes in surface ocean stratification and nutrient supply to the seafloor coeval with a paired negative carbon and oxygen isotope excursion and the return to background conditions higher up in the study section. These transitory changes are compatible with the longitudinal displacement of the proto-Gulf Stream and its related eddies. Our results build on previous work and support the hypothesis that links palaeoceanographic changes in the Blake Nose area with shifts in the proto-Gulf Stream during the middle and late Eocene.

Tectonic Stress Analysis of Shivnath-Salena Area, Using Stress Response Structure

The geological mapping of the Shivnath-Salena area and structure analysis of the same area were conducted in the LesserHimalayan sequence, Far West Nepal, unveiling the characteristics of kinematics and the associated stress field in the region. Thearea comprises of the Salena Formation and Chachura Formation. The Salena Formation is sandwiched between the PachkoraThrust and Dudulakhan Thrust, and the Chachura Formation occurs between the North Dadeldhura Thrust and Pachkora Thrust.The Salena Formation is composed of light grey and white quartzite and black slates that are intensely deformed, with folds. The Malena Anticline and the Lamalek Syncline are the intraformational folds observed in the study area. Superimposed folding was also observed. The Chachura Formation comprises Paleocene to Late Eocene green sandstone, grey and purple shale, and approximately 1 m thick grey fossiliferous limestone. This study interprets stress-response structures to show the tectonic stress field in the Salena Formation. Twenty-seven fault slip data (slickenside) were collected in the field and used to determine the stress regime by applying the stress inversion technique. The direction of the maximum principal stress axes is interpreted as NE-SW. The average value of the stress index R' is about 0.30, indicating an extensional tectonic stress regime in the study area. As σ1 is vertical, R = R' in the study area suggests normal faulting in an extensional regime.

Mineral chemistry, petrography and crystallization conditions of the Middle Eocene Kazıkbeli Pluton (Eastern Pontides, NE Türkiye)

The Eastern Pontides host a diverse suite of plutonic rocks spanning a wide range of ages and compositions. Among these, the Middle Eocene Kazıkbeli pluton, located in the Kürtün district of Gümüşhane, stands out due to its distinctive petrological characteristics. This study aims to unravel the petrological implications of petrographic and mineral chemical data to constrain the physicaydınçakırochemical conditions (temperature, pressure, oxygen fugacity) under which the Kazıkbeli magma crystallized and was emplaced. By integrating mineral chemical data, we seek to quantify emplacement pressure, crystallization temperature, and oxygen fugacity. A comprehensive understanding of the genetic relationships and physicochemical properties of the Kazıkbeli pluton rocks, as determined through geological, petrographic and mineral chemistry, is crucial for elucidating the geological evolution of the Eastern Pontides. The Kazıkbeli Pluton exhibits a predominant NE-SW orientation and encompasses an area of roughly 46 km². Modal mineralogical analysis reveals a compositional spectrum ranging from gabbroic diorite to monzogranite, with granodiorite and tonalite being the most prominent rock types. Textural variations encompass fine- to medium-grained, porphyritic, poikilitic, and occasionally graphic textures. The primary mineral assemblage of the pluton comprises plagioclase, orthoclase, quartz, amphibole, biotite, and Fe-Ti oxides. Accessory minerals include zircon, apatite, sphene, and allanite. Plagioclases are labradorite to oligoclase (An26 to An66) in composition. K-feldspar minerals exhibited an orthoclase composition (Or80 to Or97). All amphiboles belong to the calcic amphibole field and exhibit a magnesio-hornblende (Mg#=0.63-0.73) composition. Biotites crystallized as solidified melt products with compositions between annite and phlogopite endmembers, plotting close to the magnesium-rich (Mg#=0.52-0.58) end of the phlogopite solid solution series. Calculated crystallization temperatures derived from amphibole and biotite data range from 712°C to 824°C. Pressure estimations calculated using amphibole-plagioclase, amphibole and biotite suggest a range of 0.04 to 2.06 kbar. Oxygen fugacity (ƒO2) values calculated using amphibole and biotite fall between -12.5 and -16.1. Amphibole-based water content estimations indicate a range of 3.7% to 5.7% for the pluton. Biotite compositions within the studied Kazıkbeli pluton rocks exhibit characteristics suggestive of a potential mantle origin. Geobarometric calculations based on mineral chemistry data with geological and petrographic features indicate the emplacement of the Kazıkbeli Pluton at relatively shallow depths within the crust (~1 to 8 km).

Detrital-zircon U-Pb and (U-Th)/He double-dating provenance signatures in the Jaca foreland basin: interplay of direct vs. recycled sources during Pyrenean orogenic growth

ABSTRACT The Eocene to Miocene clastic wedge of the south Pyrenean basin constitutes a reference model to understand the progressive evolution of sediment provenance and source-to-sink dynamics in a foreland basin. We present new detrital-zircon (DZ) U-Pb and U-Pb-He (ZHe) double-dating data from the Jaca basin and the Ebro basin, providing insights into the evolution of the sedimentary systems that record a major tectonic and drainage reorganization from the late Eocene to Miocene. Three distinct DZ U-Pb signatures have been identified: i) Variscan dominated, ii) mixed Cadomian–Variscan, iii) Cadomian dominated, and two DZ ZHe signatures: i) Pyrenean dominated, ii) pre-Pyrenean dominated. Coupling DZ U-Pb, ZHe, and petrographic data allows us to discriminate among distinct Pyrenean sources as well as to understand how DZ signatures are propagated in a source-to-sink system. Our results indicate that while the eastern Jaca basin was fed from eastern source areas located in the central and eastern Pyrenees, the western Jaca basin was fed from the Basque massifs and the Urbasa–Andía Sierra (Basque–Cantabrian Pyrenees).

Integrated record of the Late Lutetian Thermal Maximum at IODP site U1508, Tasman Sea: The deep-sea response

The Late Lutetian Thermal Maximum (LLTM) was a transient and brief global warming event recorded in the middle Eocene, at 41.52 Ma. The biotic response to the LLTM has been documented at only a few marine sites so far. Here, we present the first record of deep-sea benthic foraminiferal assemblage changes during the LLTM in the southwest Pacific at International Ocean Discovery Program Hole U1508C (1609 m water depth) in the Tasman Sea. The LLTM coincides with a negative excursion in bulk sediment δ13C (0.47‰) and benthic foraminifera δ13C (0.36‰), with changes in the relative abundance of benthic foraminiferal species and in the deep-water organic geochemistry. The decrease in diversity of the assemblages indicates environmental stress during the event, potentially linked to oxygen deficiency, as evidenced by the occurrence of dysoxic taxa (e.g. Lenticulina spp., Turrillina brevispira). Although calcareous taxa dominate, the presence of corrosion-resistant species and poorly preserved foraminiferal tests suggest slightly CaCO3-corrosive bottom waters, but no dissolution was evident. We suggest the shallowing of the thermocline and enhanced water column stratification at this site during the LLTM.

First Record of Middle Eocene Elephant Ancestors’ Footprints in the Gonjo Basin, East Tibet Plateau

First Record of Middle Eocene Elephant Ancestors’ Footprints in the Gonjo Basin, East Tibet Plateau

A new dichobunoid artiodactyl from the middle Eocene of Yunnan, China

Dichobunoids were the most basal artiodactyl group, widely distributed throughout the Eocene of Eurasia and North America, but were rarely reported from southern China. In this paper, we report the first dichobunoid artiodactyl, Shilinhyus chowi n. gen. n. sp., from the middle Eocene of Lunan Basin (Shilin County), China. Shilinhyus is characterized by its relatively large size, bunoselenodont tooth morphology with a highly developed p4 metaconid, and a mesially interrupted paracristid formed by a weak premetacristid and a strong preprotocristid. This morphology distinguishes Shilinhyus from all known dichobunoid artiodactyls. Within dichobunoids, Shilinhyus resembles Pakkokuhyus from the Eocene of Southeast Asia and “Gobiohyus” yuanchuensis from the Eocene of Shanxi, China, by the relatively large size, the absence of the paraconid, and the well-development of the hypolophid. However, Shilinhyus still differs from Pakkokuhyus and “Gobiohyus” yuanchuensis in the complete interruption of the paracristid and in the alignment of the entoconid and hypoconid as well as the protoconid and metaconiod. Shilinhyus, Pakkokuhyus, and “Gobiohyus” yuanchuensis might have derived from the Eocene Gobiohyus from northern China, constituting a distinct group. The discovery of Shilinhyus chowi enhances our understanding of the early evolution of dichobunoid artiodactyls, especially in southern China.

New pollen taxon Syncolpraedapollis angolensis nov. gen. sp. nov.: A noteworthy discovery reported in the preliminary investigation of the latest Eocene-latest Oligocene deposits in the Kwanza Basin, Angola

A palynostratigraphic study of the upper Cunga and lower Quifangondo deposits in the Cabo de São Brás section, Kwanza Basin, Angola, has revealed a new pollen named Syncolpraedapollis angolensis nov. gen. sp. nov. This finding was part of a wider survey in the upper Cunga and lower Quifangondo deposits of the Cabo de São Brás section, Kwanza Basin, Angola, covering the latest Eocene to the latest Oligocene.The novel pollen features unique characteristics, notably a 3-syncolporate structure with well-defined and distinctive pores. The pollen grain is adorned with a free but irregular reticulum with an irregular polygonal mesh. Syncolpraedapollis angolensis nov. gen. sp. nov. is sporadically but consistently observed within the latest Eocene-latest Oligocene interval, conspicuously absent in the underlying Eocene sediments (lower Cunga Formation) and occurring infrequently in the latest Oligocene sediments (lower Quifangondo Formation). Thus, it is plausible to infer a limited age range, likely restricted to the latest Eocene-latest Oligocene, as previous studies in the Kwanza Basin have not documented this pollen.

Tectonic Inversion and Deformation Differences in the Transition from Ionian Basin to Apulian Platform: The Example from Ionian Islands, Greece

The studied areas (the Ionian Islands: Paxoi, Lefkas, Kefalonia, and Zakynthos), are situated at the western ends of the Ionian Basin in contact with the Apulian Platform and named as Apulian Platform Margins. The proposed model is based on fieldwork, previously published data, and balanced geologic cross-sections. Late Jurassic to Early Eocene NNW–SSE extension, followed by Middle Eocene to Middle Miocene (NNW–SSE compression, characterizes the Ionian basin). The space availability, the distance of the Ionian Thrust from the Kefalonia transform fault and the altitude between the Apulian Platform and the Ionian Basin that was produced during the extensional regime were the main factors for the produced structures due to inversion tectonics. In Zakynthos Island, the space availability (far from the Kefalonia Transform Fault), and the reactivation of normal bounding faults formed an open geometry anticline (Vrachionas anticline) and a foreland basin (Kalamaki thrust foreland basin). In Kefalonia Island, the space from the Kefalonia Transform Fault was limited, and the tectonic inversion formed anticline geometries (Aenos Mountain), nappes (within the Aenos Mountain) and small foreland basins (Argostoli gulf), all within the margins. In Lefkas Island, the lack of space, very close to the Kefalonia Transform Fault, led to the movement of the Ionian Basin over the margins, attempting to overthrust the Apulian Platform. Because the obstacle between the basin and the platform was very large, the moving part of the Ionian Basin strongly deformed producing nappes and anticlines in the external part of the Ionian Basin, and a very narrow foreland basin (Ionian Thrust foreland basin).

Volcanic and orbitally forced carbon release during the Middle Eocene Climatic Optimum

The drivers of the Middle Eocene Climatic Optimum (MECO) remain enigmatic. Here we report a high-resolution terrestrial MECO record from the Bohai Bay Basin, eastern China. The record shows episodic Hg enrichments and a volcanogenic Δ199Hg signature during the MECO, and an abrupt chemical weathering enhancement and negative δ13Corg excursion (CIE) during the MECO peak warmth. A high-resolution age model constrains the MECO to ca. 40.59−40.18 Ma with the onset in the first ∼320 k.y. Peak warmth and the CIE initiated at 40.27 Ma, corresponding to 405 k.y. and 100 k.y. eccentricity maxima, and lasted for ∼90 k.y., reminiscent of the early Eocene hyperthermals. Our findings suggest that episodic volcanism contributed to gradual atmospheric pCO2 rise, leading to the long duration of the MECO onset. Orbital forcing at 40.27 Ma, on top of CO2 forcing, caused the peak warmth and a positive carbon cycle feedback. The regional increase in weathering is inconsistent with evidence for weakened global continental weathering, suggesting strong spatial heterogeneity in the weathering response to MECO warming.

Heavy Mineral and Zircon Age Constraints on Provenance of Cenozoic Sandstones in the Gulf of Mexico Subsurface

Combined heavy mineral analysis and detrital zircon geochronology have enabled us to track detritus supplied by the ancestral river systems draining the North American continent into the deep subsurface of the Gulf of Mexico, in both the coastal plain and the offshore deep water areas. During deposition of the Paleocene–Eocene Wilcox Group, sandstones in the western part of the area are interpreted as the products of the Rosita system derived via paleo-Rio Grande material, with a large component of sediment shed from the Western Cordillera. By contrast, samples from wells further east have high proportions of zircons derived from the Yavapai-Mazatzal Province and are attributed to the Rockdale system with sediment fed predominantly by the paleo-Colorado or paleo-Colorado-Brazos. There is evidence that sediment from the Rosita system occasionally extended into the central Gulf of Mexico, and, likewise, data indicate that the Rockdale system sporadically supplied sediment to the western part of the basin. During the Late Eocene of the central Gulf of Mexico (Yegua Formation) there was a distinct shift in provenance. The earlier Yegua sandstones have a large Grenville zircon component and are most likely to have had a paleo-Mississippi origin, whereas the later Yegua sandstones are dominated by zircons of Western Cordilleran origin, similar to Wilcox sandstones fed by the Rosita system via the paleo-Rio Grande. The switch from paleo-Mississippi to paleo-Rio Grande sourcing implies there was a major reorganisation of drainage patterns during the Late Eocene. Miocene sandstones in the deepwater Gulf of Mexico were principally sourced from the paleo-Mississippi, although the paleo-Red River is inferred to have contributed to the more westerly-located wells.

Astronomically forcing salinity variations in a marginal-marine environment, Bohai Bay Basin, NE China

Exploring the interdependence between watermass conditions and climate change in marginal marine regions is essential for gaining insights into contemporary environmental challenges arising from global warming. Salinity, a key parameter of watermass conditions in coastal areas, plays a crucial role in influencing nutrient availability, stratification, and the spatial-temporal distribution of redox conditions. Here, we focus on the cyclostratigraphic analysis of gamma-ray (GR) data obtained from Eocene strata in the Luo-69 drillcore from the Bohai Bay Basin (NE China). This analysis establishes a high-resolution astronomical time scale (ATS) for the lower third member of the Shahejie Formation (Es3l), spanning 39.04 Ma to 41.46 million years ago (Ma). Various proxies for watermass conditions, such as salinity, sedimentary structures, mineral content, total organic carbon (TOC), organic carbon isotopes and nitrogen isotopes (δ13Corg and δ15N), exhibit a strong correlation with 405-kyr long orbital eccentricity climate forcing. Notably, spectral analysis of salinity data (represented by the Sr/Ba proxy) reveals a clear astronomical control, with 405-kyr eccentricity cycles that align closely with variations in the GR data. Peaks in paleosalinity, as indicated by Sr/Ba (as well as B/Ga and S/TOC), correspond with maxima in 405-kyr long orbital eccentricity cycles in the GR data. As such, we infer that astronomical forcing regulated long-term changes in watermass conditions, likely through mediation of the East Asia monsoon system. Specifically, during eccentricity maxima, the basin would have experienced a warm and humid climate with pronounced seasonality and a robust summer monsoon. During such periods, increased continental runoff would lead to a rise in lake-levels, facilitating better connections with the ocean, and hence increased salinity. By contrast, during eccentricity minima, cooler and more arid conditions with weaker seasonality would result in lower lake levels which together with a global eustatic lowstand would result in as weakened connection with the open ocean restricting the ingress of seawater into the basin lowering salinity. Our dataset spans the Middle Eocene Climatic Optimum (MECO, ∼40.60–40.10 Ma), identified by elevated GR values and a conspicuous negative excursion in oxygen isotopes. This interval is characterized by a significant decline in salinity, likely attributed to enhanced freshwater runoff driven by an extremely warm and humid climate. The paleosalinity fluctuation recorded by the proxies B/Ga, Sr/Ba, and S/TOC in Bohai Bay Basin demonstrate the sensitivity of marginal-marine environment to subtle changes in climate driven by astronomical forcing. Our results provide new insights into how elemental salinity proxies can be utilized in paleoenvironmental reconstruction studies.

RAD-seq data for Engelhardia roxburghiana provide insights into the palaeogeography of Hainan Island and its relationship to mainland China since the late Eocene

Hainan Island in southern China is situated within the Indo-Burma biodiversity hotspot, and has attracted the attention of evolutionary biologists, geologists, and biogeographers. However, the palaeogeography of Hainan Island and its relationship with mainland China remains contested. In this paper, we report RAD-seq data for Engelhardia roxburghiana populations from Hainan Island and adjacent mainland areas to identify genetic diversity, structure, divergence time, and demographic dynamics over geologic time. Findings were assessed using climate model data to constrain suitable distribution areas. Results indicate that E. roxburghiana dispersed from the Chinese mainland to Hainan Island via a hypothetical land bridge during the late Eocene so that the drift of Hainan Island was impossible. We cast doubt on the hypothesis that Hainan Island was connected to Vietnam and Guangxi at this time. We emphasize that higher genetic diversity in the Yunnan-Guizhou Plateau and Chiang-nan regions is the result of mixed populations or the existence of refugia, and lower genetic diversity in the Indochina Peninsula is due to a historical bottleneck. From the late Eocene to Oligocene, the arid belt that dominated East Asia retreated, facilitating the expansion of E. roxburghiana from the Indochina Peninsula to southern China. The main diversification of E. roxburghiana occurred in the Miocene following the strengthening precipitation within the East Asian Summer Monsoon. Climatic oscillations during the Quaternary led to the contraction of E. roxburghiana in the Indochina Peninsula, with expansion after Last Glacial Period (LGP; 119 to 11.7 ka). Southern China has served as a refugium and continues to do so in the future. In summary, our study elucidates the evolutionary trajectory of E. roxburghiana through large-scale sampling, providing insights into the palaeogeography of Hainan Island and itsbiogeographic relationships with adjacent mainlands.

Unraveling the tectonic evolution of the Andean hinterland (Argentina and Chile, 30°S) using multi-sample thermal history models

The Andean hinterland of Chile and Argentina defines a region of voluminous magmatism, polyphase deformation, and high topography above a long-lived subduction zone. Construction of this high-elevation domain has been variably attributed to isostatic uplift during crustal thickening induced by internal hinterland shortening, underthrusting during growth of the external thrust belt, or lower crustal flow. Alternatively, uplift may be related to dynamic processes associated with lithospheric foundering or flat-slab subduction. This study integrates geo/thermochronological, structural, magmatic, and sedimentological datasets to reconstruct hinterland evolution and evaluate potential drivers of exhumation. Our thermal history modeling approach implements a new time-depth extension to HeFTy 2 software, which enables simultaneous inversion of multiple samples along a structural or topographic profile. This extension addresses transient effects such as isotherm deflection and the transition from geothermal to atmospheric gradients and permits changes in the relative position among samples (folding, tilting) within user-defined constraints. Single- and multi-sample modeling results based on published apatite (U-Th)/He and fission track data (AHe, AFT) along the western Andean hinterland confirm late Eocene (∼30–40 Ma) cooling below at least ∼120°C, whereas thermal histories derived from new AHe, AFT, and zircon He analyses from the eastern hinterland resolve Mesozoic cooling below ∼80–160°C, followed by protracted residence at ∼60–80°C and rapid exhumational cooling in the early Miocene. Multi-sample models further require ∼10° eastward tilting of eastern sample locations, compatible with hinterland uplift via underthrusting and development of a crustal-scale fault-bend fold that was geometrically and kinematically linked to shortening in the external (eastern) thrust belt. Results underscore the enhanced resolution of multi-sample thermal history models in testing structural hypotheses and deciphering the timing, magnitude, and mechanisms of exhumational cooling during changing geodynamic conditions.

Multiproxy faunal analysis of the middle – upper Eocene deposits in the Fayum area, Egypt: Insights into sequence stratigraphy, trophic conditions, and oxygenation

The Eocene epoch experienced significant fluctuations in climate, ranging from intense greenhouse warming to icehouse cooling, which profoundly impacted the global depositional systems. The middle – upper Eocene deposits in Garet Gehannam, Fayum area, Egypt, present an exciting opportunity to explore paleoenvironmental dynamics using a multiproxy dataset of calcareous nannofossils, benthic foraminifera, ostracods, and molluscan assemblages. Calcareous nannofossil biostratigraphy constrained the studied sequence to the NP17 and NP-19-20 zones. Furthermore, three 3rd-order depositional sequences, BAR.SQ.1, BAR.SQ.2, and PR.SQ.3 were identified. The BAR.SQ.1 sequence encompasses the Gehannam Formation, with a transgressive systems tract (TST) and a highstand systems tract (HST). The BAR.SQ.2 sequence corresponds to the lower Birket Qarun Formation and exhibits a TST/HST pattern. The PR.SQ.3 sequence, however, consists solely of a TST spanning the upper Birket Qarun and Qasr El Sagha formations. The main trend of our multiproxy-based sea level curve shows an overall stepped Tethyan sea level regression within the 2nd-order cycle. This regression is characterized by a transition from outer neritic to inner settings, interrupted by two minor 3rd-order transgressive pulses. This sea level trend was likely driven by global eustatic changes and regional tectonic uplift related to the final collisional phase between Africa/Arabia and Eurasia, which led to the closure of the Neotethys Ocean. Multiproxy faunal analysis reveals a pronounced upsection shift from low-oxic and mesotrophic conditions to well-ventilated, oligotrophic settings tracking long-term sea level fall. The lowering of the sea level may have expanded aerobic ventilation windows on the continental shelf while potentially reducing nutrient influx. Our multiproxy sequence stratigraphic and paleoenvironmental analysis provides insights into the complex interplay of tectonic, eustatic, and climatic influences on the region during the middle – late Eocene episode along the southern Tethyan margin.

Controls on the genesis of a giant sand injection complex; insights into the Paleogene evolution of the stress of northern and central California

Giant sand injection complexes and localized swarms of sandstone intrusions are common in Upper Cretaceous to Miocene sedimentary successions of central and northern California within a distance of less than 100 km from the Pacific margin of the North America plate. One of the best preserved and extensively exposed injection complexes is the late Eocene Tumey Giant Injection Complex. The emplacement of sand injectites was driven by overpressure generated by thermal diagenesis of biosiliceous and smectite-rich mudstone host-rocks. The orientation and size distribution of sandstone intrusions was controlled by stress in which σ 1 and σ 3 were horizontal and, respectively, parallel and perpendicular to the present trace of the San Andreas Fault, and σ 2 was vertical. A strike-slip tectonic regime is inferred. Our analysis documents margin-orthogonal extension and provides support for a late Eocene phase of increase of strain, and possibly active slip, along a syn-subduction strike-slip fault zone. Comparison with other injection complexes in the region indicates that the near-field maximum principal stress rotated through time, from normal to parallel with respect to the plate margin, probably in relation to variations of the relative motion vector of the converging plates.

New middle Eocene radiolarian species (Rhizaria, Polycystinea) from Blake Nose, subtropical western North Atlantic Ocean

Abstract Diverse and well-preserved radiolarian assemblages were recovered from the middle Eocene sedimentary sequences drilled at Ocean Drilling Program Site 1051 (Leg 171B; western subtropical Atlantic). In addition to biostratigraphically important species, several unknown morphotypes were observed in this material, leading to the description of three new spumellarian species and 18 new nassellarian species. Described herein are: Periphaena petrushevskayae n. sp. (Phacodiscidae), Stylodictya oligodonta n. sp. (Trematodiscidae), Excentrosphaerella delicata n. sp. (Heliodiscidae), Eucyrtidium granatum n. sp. (Eucyrtidiidae), Dictyoprora echidna n. sp., Spirocyrtis matsuokai n. sp. (Artostrobiidae), Elaphospyris cordiformis n. sp., Elaphospyris quadricornis n. sp. (Cephalospyrididae), Ceratocyrtis oconnori n. sp. (Lophophaenidae), Botryocella? alectrida n. sp., Pylobotrys? bineti n. sp. (Pylobotrydidae), Lychnocanium cheni n. sp., Lychnocanium cingulatum n. sp., Lychnocanium croizoni n. sp., Lychnocanium forficula n. sp. (Lithochytrididae), Apoplanius hyalinus n. sp., Apoplanius cryptodirus n. sp. (Lophocyrtiidae), Albatrossidium messiaeni n. sp., Phormocyrtis microtesta n. sp., Cryptocarpium? judoka n. sp. (Pterocorythidae), and Thyrsocyrtis kamikuri n. sp. (Theocotylidae). Biostratigraphic information is provided for each new species. In addition, we re-describe and illustrate the morphological variability of a remarkable Pterocyrtidium species formerly published by Bütschli (1882a). UUID: http://zoobank.org/a01f7f03-73b0-458a-af7b-b85dc4666cc2

Unparallel resilience of shallow-water tropical calcifiers (foraminifera and scleractinian reef corals) during the early Paleogene global warming intervals

Investigating the fossil record to provide evidence about the response of shallow-water tropical calcifiers to past warming events is crucial considering that they are severely threatened by current global warming. We thus focused our attention on scleractinian reef corals (SRC) and shallow-water foraminifera (SWF), both mainly symbiont-bearing organisms, analysing and comparing their diversity patterns during the Paleocene and Eocene from datasets of the Neothetyan circum-Mediterranean region. In particular, at both genus and species level, we analysed changes in diversity, together with origination and extinction rates.Despite some biases related to biostratigraphic resolution of SRC and SWF data, our results show that they reacted differently to the major warming events of the Early Paleogene. The K/Pg mass extinction caused the disappearance of almost all SWF, whereas several corals passed this crisis and persisted up to the end of Paleocene, when the Paleocene-Eocene Thermal Maximum (PETM) caused a relatively small decrease of coral diversity together with a global collapse of coral reefs. On the other hand, a rapid radiation of nummulitids and alveolinids occurred at species level. The impact of the Early Eocene Climatic Optimum (EECO) was apparently more severe for SWF than for SRC. After the Middle Eocene Climatic Optimum (MECO), the general cooling trend led to the fading of Eocene SWF genera and species, whereas SRC began their rapid diversification from Bartonian to Priabonian and culminating in the Chattian.Our results suggest that rapid warming events favoured speciation in SWF, whereas the slow cooling trend (e.g., after the MECO) favoured diversification of SRC. Our data also underline that SWF reacted differently to warming events as compared to deep-sea smaller benthic foraminifera and that the same events in the planktonic realm are not strictly coeval with those occurring in shallow-water environments.On a wider perspective, we observe that shallow-water calcifiers demonstrate a good degree of resilience to global temperature increases, even if undergoing to more or less marked reduction of biodiversity. We provide evidences for the recovery of past ecosystems from both short and long stressors.

Oldest record of a crown anomaluroid rodent from sub-Saharan Africa: a new genus and species from the early Oligocene Topernawi formation of northern Kenya

ABSTRACT Anomaluroid rodents (i.e. Anomaluridae + Zenkerellidae) have a long evolutionary history in Africa dating back to at least the middle Eocene, but Palaeogene fossil members of the anomaluroid crown group have never been found south of the Sahara. Here, we describe the oldest crown anomaluroid from sub-Saharan Africa, Eliwourus topernawiensis (new genus and species) from the early Oligocene Topernawi Formation of northern Kenya. Bayesian phylogenetic tip-dating analysis of a combined molecular-morphological dataset places ~29.5 Ma Eliwourus as an advanced stem anomalurid, and suggests that crown Anomaluridae originated ~28.2 Ma. The gliding behaviour seen in all crown anomalurids had, therefore, likely evolved before the close of the early Oligocene. Bayesian geocoordinates analysis places the origin of crown Anomaluroidea (~47.2 Ma) in northern Africa, and identifies a stem anomalurid dispersal into equatorial eastern Africa in the latest Eocene or earliest Oligocene. We present the first quantitative analysis of the pace of anomaluroid dental evolution and confirm a remarkably slow or bradytelic rate of phenotypic change, despite significant transformations in postcranial morphology related to the evolution of gliding behaviour during the same interval. The Topernawi area was evidently sufficiently forested during the early Oligocene to support both arboreal rodents and primates.

Facies architecture and depositional evolution of the Middle Eocene mass flow-dominated fan delta complex in the multifeeder areas: Elazığ Marine Basin (Eastern Türkiye)

Facies architecture and depositional evolution of the Middle Eocene mass flow-dominated fan delta complex in the multifeeder areas: Elazığ Marine Basin (Eastern Türkiye)