The study was conducted among a sample of 113 Israeli fathers, and aimed to examine the variables that explain paternal involvement in care of children between the ages of one and six. Based on Bronfenbrenner’s ecological approach, the study examined the contribution of variables representing three ecological systems of the fathers: the ontogenic system – the fathers' background variables, and their childhood experiences with paternal involvement; the microsystem – children’s characteristics (age and temperament); and mothers' characteristics (education, hours of work outside of the home, and fathers' assessments of maternal gatekeeping); and the macrosystem – type of community of residence (rural or urban). Maternal gatekeeping most significantly explained paternal involvement in most of the domains of childcare that were examined. Fathers living in urban communities who assessed their wives' maternal gatekeeping as high tended to be more involved in the domain of physical care than their counterparts living in rural communities. The child’s temperament explained paternal involvement in the domains of showing love and playing with the children, whereas the fathers' childhood experiences with paternal involvement were related to their involvement in the domain of physical care. The implications of the findings for families and communities are discussed.