Grouse Moor Management Research Articles

Illegal killing associated with gamebird management accounts for up to three-quarters of annual mortality in Hen Harriers Circus cyaneus

Predators are frequently victims of wildlife crime due to conflicts with human interests. Where predators are protected, killing may occur covertly and novel methods, including satellite tracking, are often required to assess population consequences. Wildlife crime persists in the British uplands, where raptors are illegally killed on moorland managed for Red Grouse Lagopus lagopus scotica shooting. To understand impacts on one such species, the Hen Harrier Circus cyaneus, we analysed data from 148 individuals tracked across Britain between 2014 and 2021. Using remotely sensed land-use data and continuous-time survival methods, we quantified survival rates, contributions of natural causes and illegal killing to mortality, and spatial and temporal associations between mortality and land managed for grouse shooting. Annual survival was low, especially among first-year birds (males: 14 %; females: 30 %), with illegal killing accounting for 27–43 % and 75 % of mortality in first-year and subadult (1-2 years) harriers respectively. Illegal killing is likely attributable to grouse moor management because i) a 10 % increase in grouse moor use resulted in a 43 % increase in mortality risk; ii) a strong overlap existed between mortality and grouse moor extent in 20 km squares, identifying hotspots of illegal killing in northern England and northeast Scotland; iii) death due to natural causes showed different spatial and temporal patterns; and iv) timing of mortality peaked around the shooting season and during breeding territory establishment. Governments have failed to reduce illegal killing of Hen Harriers and other raptors in Britain and our results emphasise that further legislative reform is needed to tackle this enduring criminality.

Alternative future land use options in the British uplands

A commonly promoted view is that in the absence of driven grouse moor management in the UK uplands, land use on moorland would default to agriculture or afforestation. This forum article questions the orthodoxy of this thinking, discusses alternative scenarios for land use in the uplands and considers the potential effects of these on moorland‐breeding birds.

Long-term changes in the abundance and breeding success of raptors and ravens in periods of varying management of a Scottish grouse moor

Management of heather moorland for driven Red Grouse (Lagopus lagopus scotica) shooting in the British uplands may benefit some raptors by reducing predation risk, especially when breeding, and by increasing food availability. We describe changes in abundance and breeding success of four raptor species and Common Raven (Corvus corax) during a 27-year study on a grouse moor in southwest Scotland in relation to whether or not the moor was managed by gamekeepers. Ground-nesting raptors, Hen Harrier (Circus cyaneus) and Merlin (Falco columbarius), increased during periods of grouse moor management and had a higher proportion of successful nesting attempts. Predation was the main apparent cause of breeding failure. In contrast, grouse moor management did not influence either abundance or breeding success of tree- and crag-nesting species, i.e., Peregrine (Falco peregrinus), Common Buzzard (Buteo buteo), and Raven. Buzzard sightings increased during the study, in line with their national recovery, whereas Peregrine and Raven showed little change in abundance. The results of our study highlight that management for Red Grouse can benefit both Hen Harrier and Merlin. However, on a UK scale these benefits to Hen Harriers, but not Merlin, are outweighed by their illegal killing, caused by fears that their consumption of Red Grouse can undermine the economics of grouse moor management.

The influence of different aspects of grouse moorland management on nontarget bird assemblages.

Conflict between stakeholders with opposing interests can hamper biodiversity conservation. When conflicts become entrenched, evidence from applied ecology can reveal new ways forward for their management. In particular, where disagreement exists over the efficacy or ethics of management actions, research clarifying the uncertain impacts of management on wildlife can move debates forwards to conciliation.Here, we explore a case‐study of entrenched conflict where uncertainty exists over the impacts of multiple management actions: namely, moorlands managed for the shooting of red grouse (willow ptarmigan) Lagopus lagopus in the United Kingdom (UK). Debate over how UK moorlands should be managed is increasingly polarized. We evaluate, for the first time at a regional scale, the relative impacts of two major moorland management practices—predator control and heather burning—on nontarget bird species of conservation concern.Birds were surveyed on 18 estates across Northern England and Southeast Scotland. Sites ranged from intensively managed grouse moors to moorland sites with no management for grouse shooting. We hypothesised that both targeted predator control and burning regimes would enhance ground‐nesting wader numbers and, as a consequence of this, and of increased grouse numbers, nontarget avian predators should also be more abundant on heavily managed sites.There were positive associations between predator control and the abundance of the three most widespread species of ground‐nesting wader: strong effects for European golden plover Pluvialis apricaria and Eurasian curlew Numenius arquata and, less strongly, for common snipe Gallinago gallinago. These effects saturated at low levels of predator control. Evidence for effects of burning was much weaker. We found no evidence of enhanced numbers of nontarget predators on heavily managed sites.

Assessing soil compaction and micro-topography impacts of alternative heather cutting as compared to burning as part of grouse moor management on blanket bog.

BackgroundOver 25% of the UK land area is covered by uplands, the bulk of which are comprised of blanket bog. This not only contains most of the UK’s terrestrial carbon stocks, but also represents 15% of this globally rare habitat. About 30% of UK blanket bog is managed for red grouse by encouraging ling heather (Calluna vulgaris) with rotational burning, which has been linked to habitat degradation, with reduced carbon storage and negative impacts on water storage and quality. Alternative cutting is currently being pursued as a potential restoration management. However, the often used heavy cutting machinery could cause considerable compaction and damage to the peat surface. Two particular issues are (i) a potential increase in bulk density reducing water storage capacity (i.e., less pore volume and peat depth), and (ii) a possible reduction of the micro-topography due to cutting off the tops of hummocks (i.e., protruding clumps or tussocks of sedges).MethodsWe set up a fully replicated field experiment assessing cutting versus burn management impacts on peat physical and surface properties. Both managements reflected commonly used grouse moor management practice with cutting using heavy tractors fitted with load distributing double wheel and tracks (lowering ground pressure), whilst burning was done manually (setting heather areas alight with flame torches). We assessed management impacts on peat depth, bulk density and peat surface micro-topography which either included pre-management measurements or plot-level data for uncut plots. Total peat depth and bulk density in four 5 cm sections within the top 50 cm was assessed. Micro-topography was determined as the standard deviation of the height offsets measured over several plot transects in relation to the plot peat surface level at the start and end points of each transect.ResultsDespite an anticipated compaction from the heavy machinery used for cutting, the peat showed resilience and there was no lasting plot-level impact on either peat depth or bulk density. Notably, bulk density showed differences prior to, and thus unrelated to, management, and an overall increasing bulk density, even in uncut plots. However, cutting did reduce the plot micro-topography by about 2 cm, mostly due to removing the tops of hummocks, whereas burnt plots did not differ from uncut plots.DiscussionCutting is suggested as a suitable alternative to burning on grouse moors, although compaction issues might be site specific, depending on the nature of the peat, the machinery used and impacts at resting and turning points (which were not assessed). However, any observed bulk density differences could reflect natural changes in relation to changes in peat moisture, requiring adequate experimental comparisons. Moreover, where micro-topography is a priority, cutting equipment might need to consider the specific ground conditions, which could involve adjusting cutting height and the type of cutting machinery used.

Responses of breeding waders to restoration of grouse management on a moor in South-West Scotland

Worldwide, many wader species have recently exhibited steep declines in range and abundance. Low productivity, frequently associated with predation, is considered a major proximate driver of declines and often reflects underlying land use and habitat change. We hypothesised that restoration of grouse moor management, which includes control of predators and heather habitat management, would halt and reverse these declines. We monitored changes in the abundance of four upland-breeding wader species following restoration of grouse moor management at Langholm Moor in South-west Scotland from 2008 to 2017. Here, Curlew, Golden Plover and Lapwing had previously declined when management ceased, whereas Snipe had increased. During the 10-year study period, Curlew numbers increased on average by 10% per annum, Golden Plover numbers by 16% and Snipe numbers by 21%, whereas Lapwing numbers did not change. These local trends contrast with national and regional trends over the same period, which all showed declines of all these species. However, the population trends for Curlew and Snipe did not differ in relation to habitat management for Red Grouse, suggesting that population increases were primarily associated with predator control across the whole study area. Our results support the hypothesis that restoring predator control as part of grouse moor management can reverse declines of some wader species. At Langholm, full recovery to levels observed prior to management cessation may have been constrained by delayed recruitment within a time-restricted study period in combination with low initial numbers, availability and isolation of suitable habitat, and incomplete predator removal.

Spatial and temporal variation in mountain hare (Lepus timidus) abundance in relation to red grouse (Lagopus lagopus scotica) management in Scotland

Mountain hares (Lepus timidus) are regularly harvested on Scottish moorland managed for recreational hunting of red grouse (Lagopus lagopus scotica). There are concerns that large-scale culls of mountain hares on grouse moors are contributing to population declines of mountain hares in Scotland. We use hare data collected during red grouse counts in Scotland, to consider spatial and temporal variation in annual mountain hare indices of spring abundance in relation to different grouse management intensities, classified as either ‘Driven’ (driven grouse shooting), ‘Walked-up’ (walked-up shooting) or ‘Not-shot’ (where there was no grouse shooting interest). Trends in mountain hare abundance indices varied with region and grouse management intensity. Hare indices were higher and relatively stable on moors where driven grouse shooting was practised relative to lower indices and declines on moors where grouse were either walked-up or not shot. Abundance indices fluctuated over time in a quasi-cyclical manner, fluctuations being more pronounced where hares were more abundant, i.e. on driven grouse moors. Density-dependent processes such as resource competition and parasitism could explain greater fluctuations in areas where hare abundance indices were higher, whilst reductions in indices over time probably reflect sampling at decline phases of the cycle, particularly on driven moors, rather than being part of true long-term declines. Given our understanding of grouse moor management and mountain hare ecology, enhanced habitat management and control of generalist predators seem to be the most parsimonious explanation for higher abundances indices of mountain hare on driven grouse moors.

Peatland carbon stocks and burn history: Blanket bog peat core evidence highlights charcoal impacts on peat physical properties and long‐term carbon storage

Peatlands are globally important carbon stores, yet both natural and human impacts can influence peatland carbon accumulation. While changes in climate can alter peatland water tables leading to changes in peat decomposition, managed burning of vegetation has also been claimed to reduce peat accumulation. Particularly in the UK, blanket bog peatlands are rotationally burned to encourage heather re‐growth on grouse shooting estates. However, the evidence of burning impacts on peat carbon stocks is very limited and contradictory. We assessed peat carbon accumulation over the last few hundred years in peat cores from three UK blanket bog sites under rotational grouse moor burn management. High resolution (0.5 cm) peat core analysis included dating based on spheroidal carbonaceous particles, determining fire frequency based on macro‐charcoal counts and assessing peat properties such as carbon content and bulk density. All sites showed considerable net carbon accumulation during active grouse moor management periods. Averaged over the three sites, burns were more frequent, and carbon accumulation rates were also higher, over the period since 1950 than in the period 1700–1950. Carbon accumulation rates during the periods 1950–2015 and 1700–1850 were greater on the most frequently burnt site, which was linked to bulk density and carbon accumulation rates showing a positive relationship with charcoal abundance. Charcoal input from burning was identified as a potentially crucial component in explaining reported differences in burning impacts on peat carbon accumulation, as assessed by carbon fluxes or stocks. Both direct and indirect charcoal impacts on decomposition processes are discussed to be important factors, namely charcoal production converting otherwise decomposable carbon into an inert carbon pool, increasing peat bulk density, altering peat moisture and possibly negative impacts on soil microbial activity. This study highlights the value of peat core records in understanding management impacts on peat accumulation and carbon storage in peatlands.

Fighting talk: Organisational discourses of the conflict over raptors and grouse moor management in Scotland

Conflict is currently one of the greatest challenges facing wildlife conservation. Whilst conflicts may first appear to concern wildlife, they are often embedded within wider debates surrounding land use, land ownership, and the governance of natural resources. Disputes over the impacts or management of a species therefore become symbols for conflicts that are fundamentally between the divergent interests and values of the people involved. NGOs representing the interests of local stakeholders can become actors within the conflict, often utilising publicly available platforms such as websites and social media in an attempt to influence over others and gain a dominant foothold in the debate.Here, we examined discourses of organisations in relation to a contentious and high-profile case of conflict in Scotland, that occurs between interests of raptor conservation and grouse moor management. News articles sourced from the websites of six organisations – identified as key voices in the debate – were subjected to discourse analysis. 36 storylines were drawn from common phrases and statements within the text. Storylines demonstrated a clear divide in the discourse; organisations differed not only in their portrayal of central issues, but also in their representation of other actors. Discourses were strategic; organisations interpreted the situation in ways that either supported their own interests and agendas, or damaged the image of opposing parties. We argue that discursive contestation at this level could be damaging to mitigation efforts – widening barriers between stakeholders and risking already fragile relationships. This in turn reduces the likelihood of consensus and impacts on successful decision-making and policy implementation. We conclude that conflict managers should be aware of the contestation between high-profile actors, and the ramifications this may have for conflict mitigation processes. An understanding of what constitutes these discourses should therefore be used as a foundation to improve dialogue and collaborative management.

Unraveling past impacts of climate change and land management on historic peatland development using proxy-based reconstruction, monitoring data and process modeling.

Peatlands represent globally significant soil carbon stores that have been accumulating for millennia under water‐logged conditions. However, deepening water‐table depths (WTD) from climate change or human‐induced drainage could stimulate decomposition resulting in peatlands turning from carbon sinks to carbon sources. Contemporary WTD ranges of testate amoebae (TA) are commonly used to predict past WTD in peatlands using quantitative transfer function models. Here we present, for the first time, a study comparing TA‐based WTD reconstructions to instrumentally monitored WTD and hydrological model predictions using the MILLENNIA peatland model to examine past peatland responses to climate change and land management. Although there was very good agreement between monitored and modeled WTD, TA‐reconstructed water table was consistently deeper. Predictions from a larger European TA transfer function data set were wetter, but the overall directional fit to observed WTD was better for a TA transfer function based on data from northern England. We applied a regression‐based offset correction to the reconstructed WTD for the validation period (1931–2010). We then predicted WTD using available climate records as MILLENNIA model input and compared the offset‐corrected TA reconstruction to MILLENNIA WTD predictions over an extended period (1750–1931) with available climate reconstructions. Although the comparison revealed striking similarities in predicted overall WTD patterns, particularly for a recent drier period (1965–1995), there were clear periods when TA‐based WTD predictions underestimated (i.e. drier during 1830–1930) and overestimated (i.e. wetter during 1760–1830) past WTD compared to MILLENNIA model predictions. Importantly, simulated grouse moor management scenarios may explain the drier TA WTD predictions, resulting in considerable model predicted carbon losses and reduced methane emissions, mainly due to drainage. This study demonstrates the value of a site‐specific and combined data‐model validation step toward using TA‐derived moisture conditions to understand past climate‐driven peatland development and carbon budgets alongside modeling likely management impacts.

Lethal and sub‐lethal impacts of respiratory cryptosporidiosis on Red Grouse, a wild gamebird of economic importance

Respiratory cryptosporidiosis was first diagnosed in Red Grouse Lagopus lagopus scotica, the UK subspecies of Willow Ptarmigan, in 2010. In the next 3 years, respiratory infection by Cryptosporidium baileyi had manifested itself in Grouse on half the moors in northern England and 80% of moors in the North Pennine Hills. In this first account of the impact of respiratory cryptosporidiosis on the population dynamics of a wild bird we fitted 111 diseased and 67 healthy Grouse with radio‐transmitters at two North Pennine moors where disease prevalence averaged 8.1% and monitored their survival and fecundity between autumn 2013 and autumn 2015. Six‐month natural survival rates (excluding shooting) were 0.70 in healthy Grouse, but only 0.44 in diseased females, and 0.22 in diseased males. Some 39% of diseased birds died from their infection, whereas 28% of healthy birds were shot. A similar proportion of each group were killed by predators, either by Stoat Mustela erminea or raptors. Diseased females bred 8 days later than healthy females, but clutch size, egg volume and nesting success did not differ in relation to disease status. Productivity was 43% lower among pairs with a diseased member than in healthy pairs, but appeared impaired only if the female was diseased, not the male. Differences in productivity were related to chick survival rather than the proportion of pairs that reared broods, with chick survival being lower in the 10 days after hatching and again when chicks were 20–50 days old. This latter period was when respiratory infection among chicks was first noticed and the onset of infection may have been a contributing factor to higher mortality during this period. Described levels of respiratory infection reduced the number of birds available to shoot in August by 6%, which represented a mean annual loss of £0.9 million in revenue across managed grouse moors. Likely reductions in shoot economics could escalate should prevalence increase. This disease is a welfare concern and potentially a conservation concern, too, should infection cross to other bird species occupying the same moors.

Status of Golden Eagle Aquila chrysaetos in Britain in 2015

ABSTRACTCapsule: A complete survey of Golden Eagles Aquila chrysaetos in Britain in 2015 found that the population had increased by 15% since 2003 to 508 territorial pairs.Aims: The survey aimed to investigate the population size, distribution and breeding success of Golden Eagles in Britain, and to compare results with similar surveys since the early 1980s.Methods: Every home range was visited on a minimum of three occasions between January and August 2015. First, to look for eagles or signs of their presence (January–March), then to look for evidence of breeding or further checks for occupation (April–June) and finally to record productivity of nesting pairs (July–August).Results: The figure of 508 territorial pairs represents a 15% increase in the population from 442 pairs in 2003. The proportion of home ranges occupied was 70%. The largest increases in the proportion of occupied home ranges were in south-central Highlands (71%), northern moors and flows (38%) and northwest Highlands (29%), with modest increases of up to 10% in the other regions. Productivity was lower in 2015 than in 2003, and there was significant variation in breeding success between regions.Conclusion: The British Golden Eagle population has increased since 2003, although the species is absent from England and Wales. The population now meets the abundance target identified to define favourable conservation status in Scotland, and while home range occupancy has increased there is regional variation, with some regions falling below the target levels. A combination of increased annual monitoring and tagging of eagles, as well as the introduction of new legislation, may serve as effective deterrents against persecution of eagles thus facilitating this population increase. However, concerns remain over low levels of home range occupancy particularly in the east Highlands, but also the proportion of sub-adult pairs holding territory in that region and in the south-central Highlands. Persecution associated with grouse moor management has been highlighted as a particular population constraint in both of these areas.

Long-term trends in abundance and breeding success of red grouse and hen harriers in relation to changing management of a Scottish grouse moor

Large areas of heather moorland in the British uplands are managed for shooting red grouse Lagopus lagopus scotica. However, there has been a long-standing conflict between grouse moor management and the conservation of raptors, particularly the hen harrier Circus cyaneus. Langholm Moor, a grouse moor in southwest Scotland, has hosted studies aiming to resolve this conflict for 24 years. Between 1992 and 1997, whilst managed as a grouse moor, hen harrier numbers increased from two to 20 breeding females, and raptor predation removed large proportions of both adult grouse and chicks. As driven shooting was no longer viable, grouse moor management ceased in 1999, and was not restored until 2008. This paper considers how cessation and subsequent restoration of grouse moor management, which involved heather management and legal control of generalist predators, affected the abundance and breeding success of red grouse and hen harrier, as well as the abundance of their perceived key predators; red fox Vulpes vu...

Can acaricide‐impregnated leg bands fitted to female red grouse reduce sheep tick parasitization of chicks and increase chick survival?

In parts of northern England, North Wales and the Scottish Highlands, increasing numbers of sheep ticks Ixodes ricinus (Ixodida: Ixodidae), and the louping ill virus they can carry, are considered to be important factors that reduce red grouse Lagopus lagopus scotica productivity. The present study tested this hypothesis by fitting adult female grouse with leg bands impregnated with the acaricide cypermethrin to experimentally control ticks on their chicks on two managed grouse moors in northeast Scotland. The chicks of females fitted with acaricide leg bands showed reduced tick infestations and improved survival in one of the two study years, relative to chicks of control females. Acaricide leg bands constitute a potential management technique that may be adopted by grouse moor managers in circumstances of high tick infestations on grouse chicks.

Hen harriers on a Scottish grouse moor: multiple factors predict breeding density and productivity

Summary Predator density and productivity can be influenced by ecological and anthropogenic factors. Given the controversy over predators impacting on prey of economic and conservation importance, it is essential that these relationships are understood to provide a sound scientific basis for their conservation and management. We compared numbers and productivity of hen harriers Circus cyaneus, a protected specialist predator of conservation importance, in relation to a change in generalist predator management at Langholm, a moor managed for red grouse Lagopus lagopus scoticus shooting in southern Scotland between 1992 and 2007. During 1992–1999, the moor was managed for grouse and keepers legally controlled predators, thereafter keepering ceased. Following full protection being given to nesting harriers, their numbers increased from two breeding females in 1992 to 20 in 1997, when predation by harriers limited numbers of grouse available for shooting. After grouse management stopped in 1999, carrion crows Corvus corone and red foxes Vulpes vulpes increased and numbers of female harriers dropped to below five from 2002 onwards. Numbers of breeding harriers were negatively correlated with meadow pipit Anthus pratensis, crow, and July grouse abundance during the keepered period and positively with spring grouse abundance. Harrier clutch size was positively correlated with vole abundance. Harrier clutch survival and productivity were higher when the moor was keepered. Predation by foxes was the main cause of harrier breeding failure. Synthesis and applications. We consider this study to be the first that quantifies how control of generalist predators as part of grouse moor management can benefit harrier productivity. This adds to the importance of finding ways to ensure that grouse moors are managed for harriers, but are still economically viable. If techniques can be devised and put in place to reduce the impact of harriers on grouse, then the control of generalist predators may be viewed as a more acceptable component of conservation management for ground‐nesting birds.

Linking nest histories, remotely sensed land use data and wildlife crime records to explore the impact of grouse moor management on peregrine falcon populations

Wildlife crime can be difficult to quantify, and its true impact on populations can be underestimated if rates are under-recorded. The illegal killing of birds of prey is an important form of wildlife crime, which in the UK, is often associated with land managed for the recreational shooting of red grouse Lagopus lagopus scoticus. In the UK, increases in peregrine falcons Falco peregrinus following recovery from organo-chlorine pesticides have not been uniform, with slow growth and localised declines in some areas, including those managed for red grouse shooting. In this study, we combined 1081 peregrine nest histories across northern England between 1980 and 2006 with a remotely sensed map of grouse moor management, to test whether breeding performance was lower in areas with active management for grouse shooting. Productivity of pairs on grouse moors was 50% lower than pairs breeding on non-grouse moor habitat. However, clutch size and brood size of successful nests did not differ between habitat types, suggesting that food constraints were unlikely to explain this difference. Population models suggested source-sink dynamics, with populations on grouse moors unable to sustain themselves without immigration. Population data confirmed that growth rates were indeed lower on grouse moors than on non-grouse moor sites. Analysis of wildlife crime data confirmed that persecution of the species was more frequent on grouse moors than in other habitat types. This population will be more secure, and better able to function as a barometer of environmental health and climate change, if illegal persecution of the species ceases on areas of land managed for grouse shooting.

Hen harrier management: insights from demographic models fitted to population data

Summary1. The impact of hen harriers Circus cyaneus on red grouse Lagopus lagopus scoticus populations has received much attention. However, little has been done to model the population dynamics of the hen harrier alone. Such a model is needed to help inform the differing aims of conserving harriers and managing grouse moors, which serves as a reflection of human–wildlife conflicts around the globe.2. On Langholm estate in Scotland, intensive studies have resulted in harrier numbers being known without error. We fit a Bayesian population model to these data, using a super‐population model to permit inference in the presence of demographic and environmental stochasticity and in the absence of observation error.3. Hen harriers have a straightforward life history. After fledging, juveniles show little natal site fidelity, often dispersing long distances into breeding areas rich in their preferred prey, the field vole Microtus agrestis and meadow pipit Anthus pratensis. Therefore, any increase in a local population is largely because of recruitment into the area as opposed to fledging success. Once birds have settled in an area, harriers are generally site faithful, with year‐to‐year survival depending, in part, on the density of meadow pipits.4. Our model suggests that temporal patterns in harrier numbers on managed grouse moors, in the absence of illegal persecution, are influenced by vole numbers, whereas meadow pipit density appears to have a limited effect.5. Synthesis and applications. Our modelling approach is a useful way to infer population processes, and the effects of the environment on these processes, for populations censused without error. When used to predict future harrier numbers under alternate management scenarios, our model indicates that harrier numbers on Langholm estate, Scotland, could be reduced without any direct human intervention if the estate can be managed in a way that reduces vole populations. In contrast, there appears little to gain from managing meadow pipit densities. If these conclusions apply to other harrier populations, then management to reduce vole numbers, while maintaining grouse densities, may help alleviate the conflict between conservationists and managers of grouse moors.

The extent and impact of shooting on black grouse Tetrao tetrix in northern England

AbstractIn northern England, 95% of black grouse Tetrao tetrix leks occur on the fringes of managed grouse moors. Recognising the threatened conservation status of black grouse in northern England, most grouse moors have refrained from deliberate shooting for more than a decade. Despite this, black grouse are unintentionally shot in mistake for red grouse Lagopus lagopus scoticus. We assessed the circumstances and frequency of shooting in northern England and its potential impact on population recovery from three independent sources, annual shooting returns as part of the Game and Wildlife Conservation Trust's (GWCT) National Gamebag Census (NGC), incidents of shooting reported independently to the North Pennines Black Grouse Recovery Project (NPBGRP) and losses of radio‐tagged black grouse to shooting. Moors contributing to the NGC between 1998 and 2008 reported 110 black grouse shot from 13 of 23 moors, averaging 1.2% of the estimated post‐breeding population. From 2001 to 2008, 152 black grouse from 28 moors were reported shot independently to the NPBGRP, equivalent to an annual 1.0% of the post‐breeding population. The majority appeared unintentional (78%), with females (68%) more frequently shot. Out of 244 radio‐tagged black grouse on 15 estates, four adult females were shot (1.6%). The voluntary restraint from harvesting black grouse in northern England appears effective, with incidents of shooting infrequent. Continued effort to minimise shooting incidents, particularly on the fringe of the range to encourage settlement of dispersing females, may contribute to increasing numbers and range.

Exploring the relationships between wader declines and current land‐use in the British uplands

Capsule Declines of upland waders were associated with habitat cover, forest edge exposure, grouse moor management intensity and crow abundance. Aims To explore whether changes in the abundance of five wader species in the uplands correlate with the three key hypotheses proposed for their declines. Methods Using data from repeat upland bird surveys, we examined at two spatial scales (region and plot) if population changes correlate with vegetation cover, forest edge exposure, grouse moor management intensity or crow abundance. Results For Northern Lapwings Vanellus vanellus and Eurasian Curlews Numenius arquata, there was an association between declines and broad‐scale vegetation cover; declines being greatest on heather‐dominated plots. Exposure to forest edge was associated with declines of European Golden Plovers Pluvialis apricaria and Common Snipe Galinago galinago at the plot scale, and of Northern Lapwings at the regional scale. More intensive grouse moor management was associated with lower declines in Northern Lapwings at both the plot and regional scale, but with greater declines in European Golden Plovers at the plot scale. Northern Lapwings also declined most on plots with the highest Carrion Crow Corvus corone abundance. Conclusion These analyses represent the first attempt to assess empirically the balance of evidence for different broad‐scale land‐use and habitat‐related drivers of upland wader population declines, providing pointers to their respective roles in driving these declines.

Increases in humic dissolved organic carbon export from upland peat catchments: the role of temperature, declining sulphur deposition and changes in land management

Peat soils represent a highly significant carbon store and are usually assumed to serve as net sinks within carbon budgets for climate modelling. However, marked increases in dissolved organic carbon (DOC) concentrations in upland peat drainage waters over the past few decades are widespread in England. This suggests that the assumed sink status of some upland peats could be changing, although, as these observations are seldom quantified into actual carbon flux measure- ments, their overall importance is difficult to gauge. We investigated the fluvial export of the humic component of DOC (hDOC) from 3 South Pennine catchments and found that the actual carbon export via this pathway has doubled over the last 3 decades. Temperature increases to date appear poorly related to this change. Declining sulphur deposition was more strongly related (inversely), but was not a significant determinant in any of the catchments. The major driver for elevated hDOC export appears to be changes in local land management, namely the extensive use of burning for grouse moor management. Substitution of increased hDOC export into published carbon budget models for upland peat environments imply the study sites have shifted from carbon sinks to net sources over the examined period. This suggests that moorland burning, where undertaken on blan- ket peat, is incompatible with the ecosystem services these landscapes are presumed to provide and that ideas about the role of upland peat moorland in regional and national carbon cycling need reappraisal.