Log Abundance Research Articles

Associations of Two Ecologically Significant Social Insect Taxa in the Litter of an Amazonian Rainforest: Is There a Relationship between Ant and Termite Species Richness?

In spite of the ecological dominance of Neotropical ants and termites, little is understood about how their interactions influence their species richness and distribution. We surveyed ground-dwelling termite and ant species in a primary rainforest in Ecuador and analyzed ecological correlates of diversity. Termite richness was positively correlated with ant richness and abundance of twig-nesting ants. We found no evidence of competition for twigs between termites and ants. No ecological factors were correlated with termite diversity although elevation and twig and log abundance influenced ant diversity. When ant richness was compared to the richness of termites employing different predator defenses, a positive correlation was found with soldierless termites, but not genera employing chemical or mechanical defense. Our results suggest that multiple ecological factors influence ant and termite diversity, and that ant predation on termites may have a greater effect than competition between ant and termites for nest sites and food sources.

Temporal distribution and spatial pattern of abundance of the Rift Valley fever and West Nile fever vectors in Barkedji, Senegal

The temporal distribution and spatial pattern of abundance of mosquito vectors of Rift Valley fever (RVf) and West Nile fever (WNf) were studied during the 2005 and 2006 rainy seasons at Barkedji, Senegal. Mosquitoes were collected every two weeks with CDC light traps with dry ice at 79 sites including temporary ponds, barren, shrubby savannah, wooded savannah, steppes, and villages at different distances (between 0 and 600 m) from the nearest pond. The temporal distributions of these vectors varied between 2005 and 2006 and were positively correlated with rainfall for Aedes (Aedimorphus) vexans Patton, with rainfall after a lag time of one month for Culex (Culex) poicilipes (Theobald) and Culex (Culex) neavei Theobald. All the vectors had their highest abundances and parity rates between September and November. The highest vector abundances were observed in the barren and temporary ponds. The distance of trap location to the nearest ponds was negatively correlated to the abundance of the vectors. Taking into account the linear regression equations, it was predicted that mosquitoes would not disperse and be collected by the light trap, up to 1,500 m to the nearest ponds. The implications of these findings in the epidemiology and control of RVF and WNF at Barkedji are discussed.

Binary-induced magnetic activity?

We present a first and detailed study of the bright and active K0IV-III star HD 123351. The star is found to be a single-lined spectroscopic binary with a period of 147.8919+-0.0003 days and a large eccentricity of e=0.8086+-0.0001. The rms of the orbital solution is just 47 m/s, making it the most precise orbit ever obtained for an active binary system. The rotation period is constrained from long-term photometry to be 58.32+-0.01 days. It shows that HD 123351 is a very asynchronous rotator, rotating five times slower than the expected pseudo-synchronous value. Two spotted regions persisted throughout the 12 years of our observations. Four years of Halpha, CaII H&K and HeI D3 monitoring identifies the same main periodicity as the photometry but dynamic spectra also indicate that there is an intermittent dependence on the orbital period, in particular for Ca ii H&K in 2008. Line-profile inversions of a pair of Zeeman sensitive/insensitive iron lines yield an average surface magnetic-flux density of 542+-72 G. The time series for 2008 is modulated by the stellar rotation as well as the orbital motion, such that the magnetic flux is generally weaker during times of periastron and that the chromospheric emissions vary in anti-phase with the magnetic flux. We also identify a broad and asymmetric lithium line profile and measure an abundance of log n(Li) = 1.70+-0.05. The star's position in the H-R diagram indicates a mass of 1.2+-0.1 Msun and an age of 6-7 Gyr. We interpret the anti-phase relation of the magnetic flux with the chromospheric emissions as evidence that there are two magnetic fields present at the same time, a localized surface magnetic field associated with spots and a global field that is oriented towards the (low-mass) secondary component.

The cool magnetic DAZ white dwarf NLTT 10480

We have identified a new, cool magnetic white dwarf in the New Luyten Two-Tenths (NLTT) catalogue. The high proper-motion star NLTT 10480 (mu=0.5"/year) shows weak Zeeman-split lines of calcium, as well as characteristic Halpha and beta Zeeman triplets. Using VLT X-shooter spectra we measured a surface-averaged magnetic field B_S ~ 0.5 MG. The relative intensity of the pi and sigma components of the calcium and hydrogen lines imply a high inclination (i > 60 deg). The optical-to-infrared V-J colour index and the CaI/CaII ionization balance indicate a temperature between 4900 and 5200 K, while the Balmer line profiles favour a higher temperature of 5400 K. The discrepancy is potentially resolved by increasing the metallicity to 0.03x solar, hence increasing the electron pressure. However, the measured calcium abundance and abundance upper limits for other elements (Na, Al, Si. and Fe) imply a low photospheric metallicity < 10^{-4} solar. Assuming diffusion steady-state, a calcium accretion rate of log(dM/dt)(g/s)=5.6+/-0.3 is required to sustain a calcium abundance of log(n(Ca)/n(H))=-10.30+/-0.30 in the white dwarf atmosphere. We examine the implications of this discovery for the incidence of planetary debris and weak magnetic fields in cool white dwarf stars.

The trait and host plant ecology of aphids and their distribution and abundance in the United Kingdom

ABSTRACTAim We characterized the annual populations of 170 aphid species by their log abundance, site occupancy and site continuity (i.e. the persistence of species in time) and used this information to make predictions about groups of species that displayed characteristic patterns. By doing so, we aimed to identify commonalities in functional traits (host‐alternation; mode of reproduction; life‐cycle plasticity; median body size) and host plant geographic range sizes that may indicate why some species are common and others not.Location The population dynamics of winged aphids at 27 locations in the United Kingdom were studied.Methods The annual numbers of aphids were studied using a dataset comprising over 11 million individuals across 509 site‐years. Traits and host plants were analysed using linear mixed effects models and nonlinear regression models.Results Linear mixed effects models showed that the fixed effects of host alternation and winter host plant area of occupancy were important in predicting log abundance, site occupancy and site continuity. Life‐cycle plasticity was also a significant effect, although not for log abundance. Relationships between site continuity, site occupancy and log abundance were strongly nonlinear. Site continuity always lagged site occupancy, indicating that species were less likely to retain previously occupied sites when abundances were low.Main conclusions Aphid traits are a better paradigm than taxonomic relatedness in explaining macroecological patterns. Host alternation induces an annual flux of migrants that engenders higher annual log abundances, consistent with the theory that species with high local densities tend to confer a much wider distribution than those with low densities. The abundance of aphids is monotonically related to the geographic range size of their winter host, suggestive of a strong bottom‐up effect (i.e. resource controlled). The areas of occupancy (AOO) of winter host plants constrain aphids to low abundances because a greater proportion of hosts are trees and shrubs which tend to have smaller AOO than herbs and grasses, the common summer host types.

Disentangling the effects of heterogeneity, stochastic dynamics and sampling in a community of aquatic insects

A dynamic and heterogeneous species abundance model generating the lognormal species abundance distribution is fitted to time series of species data from an assemblage of stoneflies and mayflies ( Plecoptera and Ephemeroptera) of an aquatic insect community collected over a period of 15 years. In each year except one, we analyze 5 parallel samples taken at the same time of the season giving information about the over-dispersion in the sampling relative to the Poisson distribution. Results are derived from a correlation analysis, where the correlation in the bivariate normal distribution of log abundance is used as measurement of similarity between communities. The analysis enables decomposition of the variance of the lognormal species abundance distribution into three components due to heterogeneity among species, stochastic dynamics driven by environmental noise, and over-dispersion in sampling, accounting for 62.9, 30.6 and 6.5% of the total variance, respectively. Corrected for sampling the heterogeneity and stochastic components accordingly account for 67.3 and 32.7% of the among species variance in log abundance. By using this method, it is possible to disentangle the effect of heterogeneity and stochastic dynamics by quantifying these components and correctly remove sampling effects on the observed species abundance distribution.

Spectral analysis of Kepler SPB andβ Cephei candidate stars

We determine the fundamental parameters of SPB and Beta Cep candidate stars observed by the Kepler satellite mission and estimate the expected types of non-radial pulsators by comparing newly obtained high-resolution spectra with synthetic spectra computed on a grid of stellar parameters assuming LTE and check for NLTE effects for the hottest stars. For comparison, we determine Teff independently from fitting the spectral energy distribution of the stars obtained from the available photometry. We determine Teff, log(g), micro-turbulent velocity, vsin(i), metallicity, and elemental abundance for 14 of the 16 candidate stars, two of the stars are spectroscopic binaries. No significant influence of NLTE effects on the results could be found. For hot stars, we find systematic deviations of the determined effective temperatures from those given in the Kepler Input Catalogue. The deviations are confirmed by the results obtained from ground-based photometry. Five stars show reduced metallicity, two stars are He-strong, one is He-weak, and one is Si-strong. Two of the stars could be Beta Cep/SPB hybrid pulsators, four SPB pulsators, and five more stars are located close to the borders of the SPB instability region.

Assessing the nature of the combined effects of copper and zinc on estuarine infaunal communities

Elevated levels of copper and zinc in sediment have been shown to adversely affect estuarine infauna. We investigated the additivity of the combined effects of copper and zinc on infaunal recolonisation through a manipulative field experiment in Orewa estuary, New Zealand, using defaunated sediment discs treated with these metals. The nature of their combined effects varied among infaunal taxa and the particular variables being examined. Additive effects were detected for species richness, for the mean log abundances of the polychaetes Prionospio sp. and Scoloplos cylindrifer and for the multivariate response of the community as a whole. Antagonistic effects were detected for the mean log abundances of total infauna and the polychaete Heteromastus sp. Characterising the potentially interactive nature of the combined effects of multiple heavy metals is essential in order to build predictive models of future environmental impacts of metal accumulation in estuarine sediments.

Chemical evolution of the Galactic bulge as traced by microlensed dwarf and subgiant stars

<i>Context. <i/> To study the evolution of Li in the Galaxy it is necessary to observe dwarf or subgiant stars. These are the only long-lived stars whose present-day atmospheric chemical composition reflects their natal Li abundances according to standard models of stellar evolution. Although Li has been extensively studied in the Galactic disk and halo, to date there has only been one uncertain detection of Li in an unevolved bulge star. <i>Aims. <i/> Our aim with this study is to provide the first clear detection of Li in the Galactic bulge, based on an analysis of a dwarf star that has largely retained its initial Li abundance. <i>Methods. <i/> We performed a detailed elemental abundance analysis of the bulge dwarf star MOA-2010-BLG-285S using a high-resolution and high signal-to-noise spectrum obtained with the UVES spectrograph at the VLT when the object was optically magnified during a gravitational microlensing event (visual magnification <i>A<i/>~550 during observation). The Li abundance was determined through synthetic line profile fitting of the <sup>7<sup/>Li resonance doublet line at 670.8 nm. The results have been corrected for departures from LTE. <i>Results. <i/> MOA-2010-BLG-285S is, at [Fe/H] = -1.23, the most metal-poor dwarf star detected so far in the Galactic bulge. Its old age (12.5 Gyr) and enhanced [<i>α<i/>/Fe] ratios agree well with stars in the thick disk at similar metallicities. This star represents the first unambiguous detection of Li in a metal-poor dwarf star in the Galactic bulge. We find an NLTE corrected Li abundance of log<i>ϵ<i/>(Li) = 2.16, which is consistent with values derived for Galactic disk and halo dwarf stars at similar metallicities and temperatures. <i>Conclusions. <i/> Our results show that there are no signs of Li enrichment or production in the Galactic bulge during its earliest phases. Observations of Li in other galaxies (<i>ω<i/> Cen) and other components of the Galaxy suggest further that the Spite plateau is universal.

MULTIWAVELENGTH OBSERVATIONS OF THE HOT DB STAR PG 0112+104

We present a comprehensive multiwavelength analysis of the hot DB white dwarf PG 0112+104. Our analysis relies on newly-acquired FUSE observations, on medium-resolution FOS and GHRS data, on archival high-resolution GHRS observations, on optical spectrophotometry both in the blue and around Halpha, as well as on time-resolved photometry. From the optical data, we derive a self-consistent effective temperature of 31,300+-500 K, a surface gravity of log g = 7.8 +- 0.1 (M=0.52 Msun), and a hydrogen abundance of log N(H)/N(He) < -4.0. The FUSE spectra reveal the presence of CII and CIII lines that complement the previous detection of CII transitions with the GHRS. The improved carbon abundance in this hot object is log N(C)/N(He) = -6.15 +- 0.23. No photospheric features associated with other heavy elements are detected. We reconsider the role of PG 0112+104 in the definition of the blue edge of the V777 Her instability strip in light of our high-speed photometry, and contrast our results with those of previous observations carried out at the McDonald Observatory.

THE ATMOSPHERE PARAMETERS AND THE LINE PROFILE VARIATIONS OF ρ PUPPIS

We investigate <TEX>${\rho}$</TEX> Pup using the high resolution spectral observations taken from the VLT archive and observations at a 1.8m-Korean telescope with BOES spectrograph. The atmospheric parameters are determined using the iron-line abundance analysis. We derive an effective temperature value of <TEX>$T_{eff}=6890{\pm}250K$</TEX>, surface gravity of log g=<TEX>$3.28{\pm}0.3$</TEX> dex, microturbulent velocity of <TEX>${\upsilon}_{micro}=4.1{\pm}0.4km\;s^{-1}$</TEX>, and the iron abundance of log N=<TEX>$7.82{\pm}0.15$</TEX>. The projected rotational velocity of the star is close to <TEX>${\upsilon}$</TEX> sin i=3.5km <TEX>$s^{-1}$</TEX>. Asymmetric line profiles in the observed spectra and variation of this asymmetry with time show that both strong radial pulsation and weak non-radial pulsations are present in <TEX>${\rho}$</TEX> Pup.

Site preference and occurrence patterns of Picea jezoensis and Abies sachalinensis on decayed logs in natural coniferous forests in Hokkaido, northern Japan

The objectives of this study were to investigate differences in the site preferences of seedlings of Picea jezoensis and Abies sachalinensis on decayed logs, and to examine the occurrence patterns of seedlings and saplings of the two species and whether they occur together or separately on logs. We characterized the habitats of 1–2-year-old seedlings of the two species on logs and examined the relationship of the two species on logs by growth stages in two plots. One plot had been disturbed about 50 years ago whereas the other had not for a long time. Although the thickness of moss and the litter layer in the habitats of 1–2-year-old seedlings were significantly different between the two species, the two species could occur together. In one study plot, seedlings and saplings of the two species occurred together. The initial occurrence pattern of the seedlings affected the occurrence patterns of the saplings on logs. The occurrence patterns of the seedlings and saplings of the two species on logs seemed to be affected by the abundance of seed trees. In the other study plot saplings of the two species occurred separately, but one species was not always competitively superior to the other species. Disturbance history affected the occurrence patterns of the saplings of the two species on decayed logs at the two study plots. Consequently, it is concluded that seed dispersal and the abundance of available logs, which are usually affected by disturbance, are significant factors in the natural regeneration of conifers in Hokkaido.

A meta-analysis of species-abundance distributions

The species–abundance distribution (SAD) describes the abundances of all species within a community. Many different models have been proposed to describe observed SADs. Best known are the logseries, the lognormal, and a variety of niche division models. They are most often visualized using either species richness – log abundance class (Preston) plots or abundance – species rank order (Whittaker) plots. Because many of the models predict very similar shapes, model distinction and testing become problematic. However, the variety of models can be classified into three basic types: one that predicts a double S-shape in Whittaker plots and a unimodal distribution in Preston plots (the lognormal type), a second that lacks the mode in Preston plots (the logseries type), and a third that predicts power functions in both plotting types (the power law type). Despite the interest of ecologists in SADs no formal meta-analysis of models and plotting types has been undertaken so far. Here we use a compilation of 558 species–abundance distributions from 306 published papers to infer the frequency of the three SAD shapes in dependence of environmental variables and type of plotting. Our results highlight the importance of distinguishing between fully censused and incompletely sampled communities in the study of SADs. We show that completely censused terrestrial or freshwater animal communities tend to follow lognormal type SADs more often than logseries or power law types irrespective of species richness, spatial scale, and geographic position. However, marine communities tend to follow the logseries type, while plant communities tend to follow the power law. In incomplete sets the power law fitted best in Whittaker plots, and the logseries in Preston plots. Finally our study favors the use of Whittaker over Preston plots.

Hierarchical modelling and estimation of abundance and population trends in metapopulation designs

1. Population assessment in changing environments is challenging because factors governing abundance may also affect detectability and thus bias observed counts. We describe a hierarchical modelling framework for estimating abundance corrected for detectability in metapopulation designs, where observations of 'individuals' (e.g. territories) are replicated in space and time. We consider two classes of models; first, we regard the data as independent binomial counts and model abundance and detectability based on a product-binomial likelihood. Secondly, we use the more complex detection-non-detection data for each territory to form encounter history frequencies, and analyse the resulting multinomial/Poisson hierarchical model. Importantly, we extend both models to directly estimate population trends over multiple years. Our models correct for any time trends in detectability when assessing population trends in abundance. 2. We illustrate both models for a farmland and a woodland bird species, skylark Alauda arvensis and willow tit Parus montanus, by applying them to Swiss BBS data, where 268 1 km(2) quadrats were surveyed two to three times during 1999-2003. We fit binomial and multinomial mixture models where log(abundance) depended on year, elevation, forest cover and transect route length, and logit(detection) on year, season and search effort. 3. Parameter estimates were very similar between models with confidence intervals overlapping for most parameters. Trend estimates were similar for skylark (-0.074 +/- 0.041 vs. -0.047 +/- 0.019) and willow tit (0.044 +/- 0.046 vs. 0.047 +/- 0.018). As expected, the multinomial model gave more precise estimates, but also yielded lower abundance estimates for the skylark. This may be due to effects of territory misclassification (lumping error), which do not affect the binomial model. 4. Both models appear useful for estimating abundance and population trends free from distortions by detectability in metapopulation designs with temporally replicated observations. The ability to obtain estimates of abundance and population trends that are unbiased with respect to any time trends in detectability ought to be a strong motivation for the collection of replicate observation data.

Finite generation of the log canonical ring in dimension four

We treat two different topics on the log minimal model program, especially for four-dimensional log canonical pairs. (a) Finite generation of the log canonical ring in dimension four. (b) Abundance theorem for irregular fourfolds. We obtain (a) as a direct consequence of the existence of four-dimensional log minimal models by using Fukuda's theorem on the four-dimensional log abundance conjecture. We can prove (b) only by using traditional arguments. More precisely, we prove the abundance conjecture for irregular $(n+1)$-folds on the assumption that the minimal model conjecture and the abundance conjecture hold in dimension $\leq n$.

The copepod communities of the north and south Pacific central gyres and the form of species-abundance distributions

Epiplanktonic copepods were sampled on 10 cruises in the Pacific central gyres, 7 in the north gyre and 3 in the south gyre, between 1964 and 1973. These gyres are the largest biomes, stable, ancient, down-welling, oligotrophic and with little temporal variation. The data from each cruise were standardized to numbers per 10 3 m 3 ; no data from the south gyre cruises has been published before. The structure of the communities was analysed with species-abundance curves and ordination. One hundred and eighty-two species were found in all, 118―158 per cruise, 73 in all cruises. Double-centred ordination of those 73 showed three distinct sets of cruises: south (Isaacs-Kidd net), north (Isaacs-Kidd net) and north (bongo net). The distance in species-space between the north and south gyres is the same but orthogonal to the distance between samples collected by the two nets! Sixteen species abundance distributions (SADs), from 10 cruises and 6 combinations of them, were used to test the hypotheses that such distributions are mildly platykurtic and increasingly left-skew with increasing sample size. All SADs were sigmoidal on rank abundance plots of the log abundance, and agreed with the hypotheses, clarifying the mathematical form. Such replicated, large sample, SADs are rare.

General patterns in the size scaling of phytoplankton abundance in coastal waters during a 10-year time series

To ascertain the general patterns in phytoplankton size structure of a temperate, coastal ecosystem, we determined the scaling relationship between total abundance and cell size (size spectrum) for nano- and micro-phytoplankton in a shelf station off NW Iberian Peninsula on a monthly basis during the period 1993 –2002. The inverse linear relationship between log abundance and log cell size was persistent throughout the water column and across seasonal and inter-annual time scales. In addition, and despite the high productivity and marked temporal variability in water column structure at our study site, departures from linearity in the size spectra were rare. The slope (20.96) of the overall size spectrum for the entire time series indicated that roughly equal amounts of biomass were present over different logarithmic size classes in the size range considered. The phytoplankton size spectra had similar average slopes during winter mixing, early upwelling, summer stratification and autumn downwelling, suggesting that, under these oceanographic conditions, both nano- and micro-phytoplankton respond similarly to environmental variability. In contrast, significantly less negative slopes were observed during upwelling relaxation, indicating an increased importance of larger cells. Our results illustrate the utility of individual size distributions to provide a synthetic description of phytoplankton community structure in dynamic, non steady-state marine ecosystems.

A unified measure of the number, volume and diversity of dead trees and the response of fungal communities

Summary 1. Much of ecological research focuses on the responses of species and species communities to variation in the amount and quality of resources that are required for survival and reproduction. In such research, it is critical to measure the availability of resources in a manner that is relevant in relation to the ecological requirements of the species. 2. We have developed a measure for resource availability that integrates the contributions of the number, volume and diversity of resource units to quantify the amount of habitat that is available for a species community. We apply this measure to data on the occurrence of 116 species of wood‐decaying polyporous fungi in 47 study plots of boreal forest within an area of 150 × 150 km. 3. We show that species richness and pooled abundance of common species is explained well by the number of downed logs, whereas the occurrence of 41 red‐listed species is best explained by the total volume of logs and by the abundance of large logs in particular. The occurrence of common species is explained by the local availability of dead wood, whereas the occurrence of red‐listed species is additionally affected by the spatial connectivity of the focal forest stand to the surrounding larger expanses of old‐growth forest. 4. Our results elicit the contrasting ecologies of common and red‐listed species in relation to how the number of logs, their size distribution and diversity, and forest connectivity affect species occurrences. The results suggest that the most cost‐effective means of preventing further declines of threatened species is to increase the amount of large downed logs through restoration and biodiversity‐oriented management in the vicinity of existing areas of natural‐like forests. 5. Synthesis. Our results illustrate that the most relevant way of measuring resource availability can differ greatly even within a taxonomically coherent community seemingly sharing the same resources. Our approach for modelling resource availability applies to the resources that occur as discrete objects with variation in the size and quality of individual resource units.

Size‐spectra dynamics from stochastic predation and growth of individuals

In aquatic ecosystems, where organisms typically feed and grow by eating smaller individuals, a characteristic size spectrum emerges, such that large organisms are much more rare than small ones. Here, a stochastic individual-based model for the dynamics of size spectra is described, based on birth, growth, and death of individuals, using simple assumptions about feeding behavior. It is shown that the deterministic limit derived from the stochastic process is a partial differential equation previously used to describe the dynamics of size spectra. The equation has two classes of dynamics in the long term. The first is a steady state. A derivation under simple mass-balance assumptions shows that, at steady state, the linear size spectrum relating log abundance to log mass has a slope of approximately -1, similar to that often observed in natural size spectra. The second class of dynamics, not previously described, is a traveling-wave solution in which waves move along the size spectrum from small to large body size. Traveling waves become more likely when predators prefer prey much smaller than themselves and when they are specialized in the range of prey body sizes consumed. Wavelength depends on the size of prey relative to the size of predator, and wave speed depends on how fast mass moves through the spectrum.

How does abundance scale with body size in coupled size‐structured food webs?

1. Widely observed macro-ecological patterns in log abundance vs. log body mass of organisms can be explained by simple scaling theory based on food (energy) availability across a spectrum of body sizes. The theory predicts that when food availability falls with body size (as in most aquatic food webs where larger predators eat smaller prey), the scaling between log N vs. log m is steeper than when organisms of different sizes compete for a shared unstructured resource (e.g. autotrophs, herbivores and detritivores; hereafter dubbed 'detritivores'). 2. In real communities, the mix of feeding characteristics gives rise to complex food webs. Such complexities make empirical tests of scaling predictions prone to error if: (i) the data are not disaggregated in accordance with the assumptions of the theory being tested, or (ii) the theory does not account for all of the trophic interactions within and across the communities sampled. 3. We disaggregated whole community data collected in the North Sea into predator and detritivore components and report slopes of log abundance vs. log body mass relationships. Observed slopes for fish and epifaunal predator communities (-1.2 to -2.25) were significantly steeper than those for infaunal detritivore communities (-0.56 to -0.87). 4. We present a model describing the dynamics of coupled size spectra, to explain how coupling of predator and detritivore communities affects the scaling of log N vs. log m. The model captures the trophic interactions and recycling of material that occur in many aquatic ecosystems. 5. Our simulations demonstrate that the biological processes underlying growth and mortality in the two distinct size spectra lead to patterns consistent with data. Slopes of log N vs. log m were steeper and growth rates faster for predators compared to detritivores. Size spectra were truncated when primary production was too low for predators and when detritivores experienced predation pressure. 6. The approach also allows us to assess the effects of external sources of mortality (e.g. harvesting). Removal of large predators resulted in steeper predator spectra and increases in their prey (small fish and detritivores). The model predictions are remarkably consistent with observed patterns of exploited ecosystems.