Biogeographfa vol. XX VI - 2005 (Pubb/fcatoiil 30 Dicembre 2005) Biogeografia delle Alpi e Preaipi centro-orientali Coleotteri Caraboidei delle Alpi e Prealpi centrali e orientali, e loro significato biogeogtafico (Coleoptera, Caraboidea) ACHILLE CASALE* e AUGUSTO VTGNA TAGLIANTI“ ‘Dipm'tz'rm.’m0 di Zoologicz e/lntropologia Biolagiocz, Universitiz dzi Szisrczri via Muroni, 25, 1-071 00 Sasmri 6- mail.‘ cczsczle @uni5s. it q‘*Dz'pcmtz'mem‘0 di Biologiac /lrzimale e de[l’U0m0, Um'ver5z't2z dzq Roma “La! Sapienzisz” vials’ Uniyersirzi, 32, ]—00185 Roma e’*77’l£IZi[.' augusra. vignatag[iami@unir0ma1. it Key words: Central and Eastern Alps, Coleoptera, Caraboidea, historiesl-cenlogical biogeography, list of taxa. SUMMARY A synthesis of the Carabid Fauna of the Central and Eastern Alps and Pre—Alps, from a biogeographical point of view, is presented. Only the Italian side of the Alpine chain is considered, from the basin of the Toce river to the Trieste and Gorizia Karst. Main features ofthe landscape are: 1. an ancient orogenetic history and evolution, that made this area available to the colonization by carabids, of both gondwanaland and laurasian lineages, since the lower Tertiary; 2. a marked geological and geo—morphological complexity, with highest elevation at Pizzo Bernina in 4.049; 3. the presence of a very developed, dolomitiocalcareous and markedly carsified prealpine beit, rich of deep and large hypogean systems (also at high altitude), which makes this area highly difierent, from the geo-morphological point of view, from the Western Alps; 4. the geographic position, as a mountain chain of some 750 kms between central Europe and the Padanian plain, connected with the Caravanche and the Dinaric chain and close to the Adriatic sea in the Eastern part; 5. finally, a puzzled climate situation, that includes xero—therrnic areas (500-700 mm of rain per year, also in some intra-alpine sectors), close to highly rainy areas -— more than 2.500 mm per year - in the westernmost and easternmost parts of the chain. Thus, like in the Western Alps, subamediterranean vegetation types, close to cool—moist forests, coexist with high-altitude environments above the timber line. These facts explain the heterogeneity, the richness, the variety, and the interest of the carabid fauna of the area: the highest number of species (658) ascertained so far, i.e. 1/' 2 of the whole Italian carabid Fauna and about 1/5 of the carabid Fauna cited so Far for the Europe in politic sense, from the Canary islands to the Urals; the high number of euri— or stenoendemites (204, i.e. about 31 % of the species, 3/5 Ofwhich Concentrated In the pre-alpine belt), and the complex origins andfor affinities of different taxa. A large number of species (174, i.e. 26%) belongs to European chororypes (European, South—European or typically Alpine), and even more (255, i.e. 38%) to Holarctic (Holarctic, Palaearctic, West—Palaearctic, Asiatic—European, Siberian- European, Central-Asiatic-European, Central-Asiatic-European-Mediterranean, Turanic-European, Turanic-Europearb Mediterranean, European-Mediterranean) chororypes. This datum confirms the well known role, both of connection and separation, that the Alps as a whole played between Europe and the Mediterranean area. Furthermore, it is to be recalled the presence, in some localities of the Eastern Alps, ofmicrothermophilous, boreal species, like Mzscodeizz zzrmm. now widespread in Northern regions of the I-Iolarctic Region. Many orophilous, Foresedwelling, riparian and hygrophilous species, are of northern, north-eastern, turanic or dinaricvbalkanian origin. A very small, but interesting group ofthermophilous elements (mostly Harpalini), originally