Late Toarcian Research Articles

Vertebrate paleontology of the Cloverly Formation (Lower Cretaceous), IV: the oldest edentulous frog (Salientia) from Laurasia

ABSTRACT Edentulism—lack of teeth—is a derived condition among salientians (total-group frogs and toads) that has arisen independently at least 22 times within the crown group, Anura. Despite this frequency, edentulism (in part or complete) is seldom documented in the salientian fossil record, and thus its evolutionary history remains obscure. A ∼90 Ma gap presently exists between the edentulous salientians Notobatrachus reigi (late Toarcian; Argentina) and the five known species from the latest Cretaceous of Argentina, South Africa, Canada, and the U.S.A. Here we report a new instance of edentulism in an Early Cretaceous salientian, Ostrombatrachos nodos gen. et sp. nov., based on a maxilla from an early Albian age (∼ 111 Ma) locality in the Cloverly Formation of Wyoming, U.S.A. Although incomplete, the holotype maxilla exhibits a distinctive suite of features (i.e., edentulous; pit-and-ridge external ornament; shallow groove across anteriormost portion of margo orbitalis and continuing anteroventrally down lateral surface of pars facialis to interrupt the external ornament; medially expanded orbital flange; moderately broad, deep lamina horizontalis; and weakly developed processus pterygoideus) that is unknown in any other salientian. This new taxon represents the oldest occurrence of edentulous maxillae in a Laurasian salientian and helps fill the long temporal gap in edentulism among salientians. O. nodos gen. et sp. nov. does not appear to be closely related to other edentulous Mesozoic salientians, suggesting it represents yet another independent evolution of this feature.

Exquisite skeletons of a new transitional plesiosaur fill gap in the evolutionary history of plesiosauroids

Plesiosaurs are Mesozoic reptiles fully adapted to an aquatic lifestyle. Throughout their evolutionary history exceeding 140 million years plesiosaurs dispersed globally, achieved substantial diversity, occupied a variety of ecological niches, and experienced multiple faunal turnovers. Of those, the Early/Middle Jurassic transition event (∼175–171 Mya) has recently became of increased interest because it apparently profoundly affected all three major lineages of plesiosaurs. Once dominant Rhomaleosauridae started to vanish, while Pliosauridae and Plesiosauroidea diversified and gave rise to several clades that flourished for tens of millions of years. Here, we report exquisite, three-dimensionally preserved skeletons of a new plesiosaur from the Lower Jurassic of Germany. Franconiasaurus brevispinus gen. et sp. nov. lived during the late Toarcian (∼175 Mya), near the onset of the Early–Middle Jurassic turnover. Franconiasaurus displays an intriguing mixture of features, combining characters almost uniformly distributed among early plesiosaurs with those typically observed in later-diverging members of the clade. Phylogenetic analyses firmly place Franconiasaurus as the sister taxon to Cryptoclidia, bridging an evolutionary gap between early plesiosauroids, such as Plesiosaurus-like forms and microcleidids, and later-diverging representatives of the clade, such as cryptoclidids, leptocleidians, and elasmosaurids.

Spatially heterogenous seawater δ34S and global cessation of Ca-sulfate burial during the Toarcian oceanic anoxic event

The early Toarcian of the Early Jurassic saw a long-term positive carbon-isotope excursion (CIE) abruptly interrupted by a significant negative excursion (nCIE), associated with rapid global warming and an oceanic anoxic event (T-OAE, ∼183 Ma). However, the detailed processes and mechanisms behind widespread ocean deoxygenation are unclear. Here, we present high-resolution carbonate-associated sulfate sulfur-isotope (δ34SCAS) records spanning the late Pliensbachian–Toarcian (Pl–To) interval from the Tibetan Himalaya. We observe a large positive sulfur-isotope excursion (SIE) from ∼20‰ (around the Pl–To boundary) to ∼40‰ (around the end of the T-OAE nCIE), attributed to large-scale burial of reduced sulfur (pyrite and sulfurized organic matter) under widespread anoxic/euxinic conditions. Importantly, high δ34SCAS values were maintained into the mid–late Toarcian, even when global anoxic conditions diminished. The δ34S data confirm significant spatial heterogeneity in seawater δ34S compositions during the whole of the Toarcian, and provide strong evidence for a two-phase pattern of ocean deoxygenation. Upwelling of 34S-enriched equatorial deep water, affected by significant reduced-sulfur burial, likely caused the greatly amplified SIE in the formerly adjacent Tibetan area. By contrast, the dampened magnitude of the Toarcian SIE in Europe is attributed to a smaller, local reduced-sulfur sink. Box-modeling results indicate that the persistent post-T-OAE positive δ34S values were likely maintained because of a global reduction in Ca-sulfate (gypsum and anhydrite) burial driven by declining and continuously low seawater sulfate concentrations during and after the T-OAE. This geochemical pattern, albeit markedly reducing the total amount of global pyrite sequestration, increased the proportion of reduced to oxidized sulfur burial needed to generate the observed positive δ34S values.

High-precision Re[sbnd]Os dating of Lower Jurassic shale packages from the Western Canadian Sedimentary Basin

The Lower Jurassic rock record contains abundant organic-rich mudrocks suitable for refining Early Jurassic time using ReOs organic-rich shale (ORS) geochronometry, such as those associated with the Sinemurian–Pliensbachian boundary event or the Toarcian Ocean Anoxic Event (T-OAE). Despite this, most studies to date have failed to obtain high precision, geologically meaningful, ages for such Lower Jurassic rocks. The cause of imprecision in ReOs ORS age dates, at least for the Toarcian, has been linked to subtle chemostratigraphic variations in 187Os/188Osi values. Here, we report the first high-precision ReOs ages of organic-rich shale packages of the Gordondale and basal Poker Chip Members of the Lower Jurassic Fernie Formation of Northwestern Alberta, Canada from the Western Canadian Sedimentary Basin (WCSB). Our ReOs age results bracket sediment deposition between the late Sinemurian-early Pliensbachian (192.11 ± 0.77 Ma [2σ] and 193.83 ± 1.34 Ma [2σ]) and late Toarcian (180.10 ± 2.68 Ma [2σ]). When coupled with chemo- (87Sr/86Sr and 187Os/188Os isotopes) and bio-stratigraphy (ammonite and coccolith), our ReOs ages correlate with regional and global stratigraphic tie points corresponding to pivotal points of Early Jurassic time, such as the Sinemurian–Pliensbachian boundary event, T-OAE, and late Toarcian, that are otherwise weakly constrained by isotopic methods. This integrative approach helps to place these events in a more robust temporal framework. Based on seawater Os isotope data, we also speculate that a previously unidentified positive Os isotope excursion exists at the Sinemurian–Pliensbachian boundary - perhaps triggered by late-stage Central Atlantic Magmatic Province (CAMP) volcanism.

Stratigraphy, palaeoenvironment and mollusks (gastropods and bivalves) of the Toarcian succession of the Central Unit of the South Riffian Ridges (northern Morocco)

In the Central Unit of the South Riffian Ridges, the Toarcian succession is represented by two thick terrigenous detrital formations, Sned and Lmachkha formations, each one is divided into two members. The lower member of the Sned Formation, often rich in belemnite rostra and ammonites of the Tenuicostatum Zone, consists of grey marlstones, with rare marly limestone intercalations. The upper member consists of silty marlstones, with frequent thin limestone beds and contains rare ammonites of the Serpentinus Zone. Its upper part is locally rich in terebratulids. The lower member of the Lmachkha Formation consists of a silty marlstone and silty limestone alternation, and is the richest in benthic macrofauna, composed of rhynchonellids, gastropods, common bivalves, and rare ammonites of the Bifrons Zone. Seven taxa of gastropods and six taxa of bivalves were recorded from this member and described for the first time from the study area. The upper member of the Lmachkha Formation consists of silty marlstone and silty limestone alternation, and contains rare bivalves and rare ammonites of late Toarcian age. The lithofacies and pelagic macrofauna of the lower member of the Sned Formation characterise a relative deep marine environment unfavourable to benthic macrofauna. The second member indicates a gradual decrease in sea-level allowing from its upper part the recurrence of benthic macrofauna, especially terebratulids. The lithofacies and macrofauna of the lower member of the Lmachkha Formation indicate a shallow terrigenous shelf environment. The second member, containing coral reefs in the western and southern peripheries of the Central Unit, characterises a still shallow marine environment. This significant but gradual decrease in sea-level of the Central Unit from the latest early Toarcian to late Toarcian is likely related to the acceleration of sea-floor spreading in the Central Atlantic Ocean.

Volcanic rocks in Lower Jurassic marine carbonate successions in the southeastern Iberian Range (Spain): biochronostratigraphic dating

The Ligurian Tethys began to open in a roughly NE–SW direction in Pliensbachian times. West of the Ligurian oceanic crust, a carbonate-platform system constituted the eastern Iberian paleomargin. Lower and Middle Jurassic marine sediments deposited in these shallow platforms are well exposed today in the southeastern Iberian Range and host multiple volcanogenic deposits, mainly volcaniclastic in nature. Regional mapping reveals that volcanic outcrops show NW and NE (or NNE) oriented lineaments following tracks of cortical weakness. Volcanic ages were estimated by using biochronostratigraphic methods and sequence-stratigraphic correlations. As a result, ten Lower Jurassic volcanic episodes have been identified. The chronostratigraphic positions of the volcanic deposits span from the early Pliensbachian (Jamesoni Zone or later) to the late Toarcian (Thouarsense Zone). The extension-dominated regime of the eastern Iberian paleomargin also affected the southern Iberian paleomargin (Betic Cordillera) as a result of the eastward prolongation of the central Pangea fragmentation. The break-up of wide platforms and the first volcanic emissions seem to be roughly coeval in the Betic and Iberian domains and synchronous with the onset of rifting and spreading events of the Ligurian Tethys Ocean.

Redox conditions and climate control on organic matter accumulation and depletion during the Toarcian in the Qiangtang Basin, eastern Tethys

The Toarcian (Early Jurassic) sediments are regarded as the most significant hydrocarbon source rocks in the Qiangtang Basin, Qinghai–Tibet Plateau. In this study, geochemical and mineralogical data are reported from the Middle to Late Toarcian organic-lean sediments in the Qiangtang Basin, eastern Tethys. Combined with previous studies, the factors that control organic matter (OM) accumulation during the Toarcian are investigated. The Early Toarcian organic-rich sediments were deposited in stratified water column with oxygen-deficient bottom water, which was conducive to OM preservation. A warm and humid paleoclimatic condition favored the growth of organisms, which in turn could enhance the surficial biotic productivity. During the Middle to Late Toarcian, oxic waterbody was widely distributed in the Qiangtang Basin. This condition was unbeneficial to the preservation of OM. During this interval, a cold–arid climate had an adverse effect on organism growth, which in turn would reduce the initial productivity of surface water. The consistently low Ti/Al and Si/Al ratios revealed restricted but rather homogeneous clastic influx conditions during the Toarcian. This may suggest that the dilution of terrigenous material input had a little influence on OM accumulation. In addition, the differential enrichment of OM between shales and marls should attribute to carbonate dilution. These studies emphasize that redox conditions and paleoclimatic conditions were likely responsible for OM accumulation during the Toarcian.

Anatomy and Jurassic evolution of a Hercynian basement high (Caloveto High - Calabria, Southern Italy)

The Caloveto area in North-East Calabria hosts a stratigraphic succession which documents the evolution from shallow water carbonates to deeper-water pelagic and hemipelagic deposits, bearing ample evidence for severe tectonic control on Jurassic and Lower Cretaceous sedimentation. Geological mapping indicates that the shallow-water carbonates, also with coral assemblages, formed a narrow fringe around a high of the Hercynian basement, made of lowgrade metamorphic rock, which remained emergent throughout the Pliensbachian and became an intrabasinal high which stood proud of the Longobucco basin, hosting thin pelagic sedimentation through most of the Jurassic. An extensional phase in the Toarcian disrupted and foundered the benthic carbonate factory, whose drowning isdocumented by a change to Rosso Ammonitico-type deposits. This resulted in a complex network of neptunian dykes (also intruding the basement) and in-situ breccias. Starting in the late Toarcian, the rugged submarine topography was gradually leveled by onlapping marls, radiolarites and pelagic limestone, which sealed the Toarcian fault zones producing a diverse array of unconformities. Colonization by microbial communities characterizes the initial stages ofmarine sedimentation around the cores of Paleozoic basement, as banded polychrome microbialites and “swollen” phyllites, a result of the microbially-influenced/induced displacive growth of calcium carbonate crystals along split cleavage planes, constitute an unexpected field tool for identifying basin margins. Silicified marginal zones in the shallow water limestone characterize, in analogy withother Tethyan regions, the onlap unconformities of chert-rich basinal units on the submerged carbonate fringes.

Poly-phased fluid flow in the giant fossil pockmark of Beauvoisin, SE basin of France

The giant Jurassic-aged pockmark field of Beauvoisin developed in a 800 m wide depression for over 3.4 Ma during the Oxfordian; it formed below about 600 m water depth. It is composed of sub-sites organized in clusters and forming vertically stacked carbonate lenses encased in marls . This fine-scale study is focused on a detailed analysis of petrographical organization and geochemical signatures of crystals that grew up in early to late fractures of carbonate lenses, surrounding nodules, and tubes that fed them. The isotopic signature (C, O and Sr) shows that at least three different episodes of fluid migration participated to the mineralization processes. Most of the carbonates precipitated when biogenic seepage was active in the shallow subsurface during the Oxfordian. The second phase occurred relatively soon after burial during early Cretaceous and thermogenic fluids came probably from underlying Pliensbachian, Late Toarcian or Bajocian levels. The third phase is a bitumen-rich fluid probably related to these levels reaching the oil window during Mio-Pliocene. The fluids migrated through faults induced by the emplacement of Triassic-salt diapir of Propiac during the Late Jurassic and that remained polyphased drain structures over time.

First steps in reconstructing Early Jurassic sea water temperatures in the Andean Basin of northern Chile based on stable isotope analyses of oyster and brachiopod shells

The stable isotope (δ13C, δ18O) composition of a collection of Lower Jurassic brachiopods and oysters from the Andean Basin of northern Chile was analyzed. The results allow the first reconstruction of absolute water temperatures for several ammonite zones in the Lower Jurassic of South America. The temperature record starts with comparatively high values in the Late Sinemurian (average: 27.0 °C; Raricostatum Zone). Just before the Sinemurian–Pliensbachian transition, temperatures dropped to an average of 24.3 °C. The lowest temperature value in the dataset was recorded for a brachiopod shell of the latest Pliensbachian Spinatum Zone (19.6 °C). No data are available for the Early Toarcian, but results for the late Toarcian show again comparatively warm conditions (average: 24.4 °C; Thouarsense–Levesquei zones). Even though more material and analyses are necessary to corroborate the recorded temperatures, the present dataset seems to indicate the global nature of the Late Pliensbachian Cooling Event. In contrast, the global warming during the Toarcian Oceanic Anoxic Event has not been recorded due to a lack of Early Toarcian material. The δ13C record of brachiopods and oysters documents a gradual increase in values representing background conditions. Oyster shells were used for high-resolution stable isotope analyses and show seasonal temperature fluctuations over a period of around 3 years in the life time of the bivalves. If explained only by temperatures, the δ18O values point to a minimum estimate for the seasonality in the late Toarcian of slightly more than 3 °C.

New results of stable isotope and petrographic studies of Jurassic glendonites from Siberia

ABSTRACTWe present the results of an optical microscopy, cathodoluminoscopy and isotopic study on nine glendonite concretions (calcite pseudomorphs replacing metastable hexahydrate ikaite) from Lower-Middle Jurassic sediments of Northeast Russia (Anabar Bay and Lena River region). Glendonite concretions are mainly found within Late Pliensbachian, Toarcian, Aalenian, Bajocian and Lower Bathonian clastic sediments, correlating to episodes of global climatic cooling as determined by independent paleoclimate proxy data. Stable carbon and oxygen isotopic values of glendonite concretions suggest that the primary source of carbon was derived from diagenetically altered organic matter, and the source of oxygen was from seawater. The secondary diagenetic cement is characterized by a significantly lighter δ18O and significantly heavier δ13C signature than the isotopic characteristics of the bulk rock glendonite concretion. This secondary diagenetic cement is thought to have precipitated rapidly during burial diagenesis and since it occupies a significant volume of the glendonite concretion, it has the potential to significantly influence the isotopic composition of bulk rock glendonites.

Refined age and geological context of two of Australia’s most important Jurassic vertebrate taxa (Rhoetosaurus brownei and Siderops kehli), Queensland

Australia’s Jurassic vertebrate fossil record remains extremely sparse with only two dinosaur taxa and two temnospondyl amphibians identified to date. Of these, the spectacular and extremely well-preserved giant amphibian, Siderops kehli, and the only known pre-Cretaceous sauropod in Australia, Rhoetosaurus brownei, are perhaps the most important. The age of both specimens, and the stratigraphic context of Rhoetosaurus brownei, are weakly constrained and imprecisely defined, limiting our understanding of their evolutionary relationships within a broader Gondwanan context. To clarify and contextualise the evolutionary relationships and ages of these two iconic Jurassic taxa, we used UPb detrital zircon geochronology to date the sandstone matrix from around the bones of the historic museum specimens. The robust maximum depositional age for Siderops was calculated at 176.6 Ma ± 2 Ma, indicating that it is no older than late Toarcian, which refines existing biostratigraphic estimates. The Rhoetosaurus maximum depositional age was determined to be 162.6 ± 1.1 Ma, no older than early Oxfordian, demonstrating that the fossils are younger than expected, and definitely recovered from the Walloon Coal Measures.

Jurassic reef events in the Moroccan Atlas: A review

In the Moroccan Atlas, sedimentary deposits provide important data on reef events that characterize the Jurassic period. Recent work allows us to enhance knowledge of the Jurassic reefs in the Atlas, in particular their age, character and palaeogeographic distribution. Numerous localities with sponge-microbial mud mounds, coral reefs, and lithiotid bioherms are recorded from the Middle and High Atlas regions. These different biogenic constructions occupy different palaeogeographic settings: on the top of tilted blocks within the basin center; in slightly deeper positions, at the basin platform junction; and on adjacent platforms in the middle of the coastal area. The main episodes of reef building span nearly 30 million years, as follows: (1) Sinemurian, (2) early Pliensbachian, (3) late Toarcian, (4) Aalenian–early Bajocian (pars), and (5) late Bajocian. These five distinct reef events can be linked with general fluctuations of sea level and tectonism, and have palaeoclimatic implications.

The epiclastic barrier-island system of the Early‒Middle Jurassic in eastern Spain

During the Early-Middle Jurassic, several mainly volcaniclastic submarine eruptions, which followed well marked lineaments, took place in the Iberian Ranges of eastern Spain. Some of these volcanoes were occasionally emerged during sea-level lowstand periods. In this work we aim to present the signs indicating that a huge volcanic pile, located in the Sierra de Javalambre (Teruel, Spain), could have been subjected to emersion, reworking and colonization by flora. We also intend to date the volcanism and to delimit the most likely period of time when emersion, reworking and colonization were produced. The research work was supported by geological mapping, as well as by preparing stratigraphic sections and their palaeontological sampling. Biostratigraphical dating was achieved though taxonimical determination of ammonites, brachiopods, and data from pre-existing palynological studies. As a result of the shallow marine and subaerial partial erosion of the volcanic edifice, epiclastic bodies, some of which show large scale cross-bedding and bar geometry, were generated. These bedforms are interpreted as sandwaves constituting a barrier-island system protecting a lagoon on which low-energy sediments, rich in plant remains, together with washover fan facies were deposited. An Early Toarcian, Serpentinum Chronozone age for the volcanism is indicated by the brachiopod and ammonite content of infra- and inter-volcanic sediments. However, burial of the volcanic mound by the carbonates of the surrounding platforms ended in the Late Aalenian Bradfordensis Chronozone, about 10 million years after the eruption. This delay was partly due to a significant regional sedimentary gap marked by the hiatus of the Late Toarcian (from the Late Variabilis Chronozone) to the Late Aalenian (Bradfordensis-Concavum chronozones). The emersion of the volcanic pile, the colonization of the emerged islands by continental vegetation and the instauration of the epiclastic barrier-island system occurred between part of the Aalenian Murchisonae and part of the Bradfordensis chronozones. The concurrence of the porous epiclastic lithofacies together with the organic-rich lagoonal sediments could represent an analogue model of an untypical hydrocarbon system located in neritic position, which could probably be applied in other areas of extensive emerged submarine volcanoes present in the geological record.

The Early Jurassic palynostratigraphy of the Lusitanian Basin, western Portugal

A comprehensive investigation of the Early Jurassic stratigraphical palynology of the Lusitanian Basin in western Portugal was undertaken, with most emphasis placed on dinoflagellate cysts. A total of 214 samples from an upper Sinemurian to upper Toarcian composite section based on six successions were examined. The Sinemurian material examined was barren of dinoflagellate cysts; however, the Pliensbachian and Toarcian successions are characterised by relatively low diversities where Luehndea spinosa, Mancodinium semitabulatum, Mendicodinium microscabratum, Nannoceratopsis gracilis, Nannoceratopsis senex, and Scriniocassis priscus are relatively common and biostratigraphically significant. Luehndea spinosa dominates the lowermost Toarcian (Dactylioceras polymorphum ammonite Biozone), and is an index species. At the base of the Hildaites levisoni ammonite Biozone, the effects of the Toarcian Oceanic Anoxic Event (T-OAE) caused Luehndea spinosa to become extinct. At the same time, dinoflagellate cyst abundance and diversity markedly decreased. After the T-OAE, during the middle and late Toarcian, phytoplankton recovery was prolonged and slow in the Lusitanian Basin. The Luehndea spinosa and Mendicodinium microscabratum dinoflagellate cyst biozones are defined, both of which are subdivided into two dinoflagellate cyst subbiozones.

Benthic foraminiferal bioevents in lower to upper Toarcian strata of Southern Beaujolais (SE France)

This paper documents changes in benthic foraminiferal assemblages compared with high resolution ammonite biozonation along the lower Toarcian to upper Toarcian marine succession of Southern Beaujolais in southeastern France. Eight ammonite and three benthic foraminiferal zones including five subzones are distinguished based on the occurrence of twelve foraminiferal events. Each benthic foraminiferal subzone is characterized by its taxonomic and morphogroup composition, which represents the paleoecological response of these taxa and morphotypes of benthic foraminifera in the Early Jurassic and early Middle Jurassic. Major changes in abundance and diversity occur at the end of the Toarcian Oceanic Anoxic Event (T-OAE) and near the Early-Middle Jurassic transition. The low-abundance foraminiferal assemblages recorded in the Serpentinus ammonite Zone are interpreted as reflecting adverse environmental conditions after the T-OAE. The later recovery and development of the foraminiferal assemblages is documented in the Bifrons up to the Aalensis zones and is attributed to improved bottom water oxygenation. Common occurrences of agglutinated foraminifera represented mostly by Trochammina pulchra Ziegler in the Dispensum Zone point to an influx of cooler water masses during the late Toarcian. The morphogroup analysis carried out on the foraminifera and their paleoecological interpretations shed light on the changes in the stratigraphic record at the end of the T-OAE up to the Toarcian/Aalenian boundary.

The planktonic foraminifera of the Jurassic. Part II: Stratigraphy, palaeoecology and palaeobiogeography

The stratigraphy, palaeoecology and palaeobiogeography are outlined for two genera and ten species of Jurassic planktonic foraminifera described in Gradstein et al. (Swiss J Palaeontol 2017. doi: 10.1007/s13358-017-0131-z). The two genera are Globuligerina and Conoglobigerina. Globuligerina probably appeared in late Toarcian (late Early Jurassic) and Conoglobigerina first occurred in Middle Oxfordian (early Late Jurassic). Within the two genera ten species are defined, including from older to younger: Globuligerina dagestanica (Morozova), G. avariformis (Kasimova), G. balakhmatovae (Morozova), G. oxfordiana (Grigelis), G. bathoniana (Pazdrowa), G. jurassica (Hofman), G. oxfordiana (Grigelis) calloviensis Kuznetsova emended, G. tojeiraensis Gradstein, Conoglobigerina helvetojurassica (Haeusler), C. grigelisi Gradstein and C. gulekhensis (Gorbachik and Poroshina). Globuligerina balakhmatovae, G. oxfordiana and G. bathoniana are longer ranging, although there are subtle evolutionary trends in these taxa that may refine stratigraphic usage. Other taxa have shorter stratigraphic ranges. Using stratigraphic trends, test morphology and wall texture changes, the species are tentatively assembled in four evolutionary groups, termed A–D, and in six zones from Late Toarcian through Tithonian. The greater species diversity and possibly also specimen abundance took place in the Kimmeridgian, a time of high global sea level and greenhouse palaeoclimatic conditions. Jurassic planktonic foraminifera preferred marine continental margin conditions instead of the distal open ocean and spread from offshore Eastern Canada to offshore Western Australia along the margins of the Tethys Ocean. The group did not venture into austral or boreal realms, likely reasons why migration probably failed to the west coasts of South and North America.

The planktonic foraminifera of the Jurassic. Part I: material and taxonomy

The taxonomy is modernized and updated for Jurassic planktonic foraminifera using all available literature and stratigraphic series of sediment samples from Canada, Portugal, France, Switzerland, Poland, Lithuania, Russia and Dagestan. The regional stratigraphy of all sampled sections is outlined. Many of the 65–200 µm sample residues yielded well-preserved free specimens, with local assemblages containing up to six species. Thin sections were studied also but do not allow detailed taxonomy. Two genera are described: Globuligerina and Conoglobigerina. Globuligerina probably appeared in the late Toarcian (late Early Jurassic) and Conoglobigerina first occurred in middle Oxfordian (early Late Jurassic). A principal difference between Globuligerina and Conoglobigerina is that the former has a smooth to pustulose wall surface texture and the latter a reticulate wall surface texture. Within the two genera, ten species are described, including from older to younger: Globuligerina dagestanica (Morozova), G. avariformis (Kasimova), G. balakhmatovae (Morozova), G. oxfordiana (Grigelis), G. bathoniana (Pazdrowa), G. jurassica (Hofman), G. oxfordiana (Grigelis) calloviensis Kuznetsova emended, G. tojeiraensis Gradstein, Conoglobigerina helvetojurassica (Haeusler), C. grigelisi Gradstein and C. gulekhensis (Gorbachik and Poroshina). The coarsely perforate ‘Globuligerina’ stellapolaris needs more investigation to establish its identity as a Jurassic or a Late Cenozoic taxon (Gradstein 2017a). Globuligerina balakhmatovae, G. oxfordiana and G. bathoniana occur from Bajocian through early Kimmeridgian, or even Tithonian. Subtle morphological trends may refine stratigraphic usage. Other taxa have shorter stratigraphic ranges. Nineteen plates and ten type species figures detail morphology of all taxa. For key taxa, both the types and specimens from other localities are illustrated.

Litho- and biostratigraphy of the Opalinus Clay and bounding formations in the Mont Terri rock laboratory (Switzerland)

A 250 m-deep inclined well, the Mont Terri BDB-1, was drilled through the Jurassic Opalinus Clay and its bounding formations at the Mont Terri rock laboratory (NW Switzerland). For the first time, a continuous section from (oldest to youngest) the topmost members of the Staffelegg Formation to the basal layers of the Hauptrogenstein Formation is now available in the Mont Terri area. We extensively studied the drillcore for lithostratigraphy and biostratigraphy, drawing upon three sections from the Mont Terri area. The macropaleontological, micropaleontological, and palynostratigraphical data are complementary, not only spatially but they also cover almost all biozones from the Late Toarcian to the Early Bajocian. We ran a suite of geophysical logs to determine formational and intraformational boundaries based on clay content in the BDB-1 well. In the framework of an interdisciplinary study, analysis of the above-mentioned formations permitted us to process and derive new and substantial data for the Mont Terri area in a straightforward way. Some parts of the lithologic inventory, stratigraphic architecture, thickness variations, and biostratigraphic classification of the studied formations deviate considerably from occurrences in northern Switzerland that crop out further to the east. For instance, with the exception of the Sissach Member, no further lithostratigraphic subdivision in members is proposed for the Passwang Formation. Also noteworthy is that the ca. 130 m-thick Opalinus Clay in the BDB-1 core is 20 m thinner than that equivalent section found in the Mont Terri tunnel. The lowermost 38 m of the Opalinus Clay can be attributed chronostratigraphically solely to the Aalensis Zone (Late Toarcian). Deposition of the Opalinus Clay began at the same time farther east in northern Switzerland (Aalensis Subzone, Aalensis Zone), but in the Mont Terri area the sedimentation rate was two or three orders of magnitude higher.

Extraordinary diversity of feather stars (Echinodermata: Crinoidea: Comatulida) from a Lower Jurassic (Pliensbachian–Toarcian) rock reef of Feuguerolles (Normandy, France)

Comatulid feather stars are rare fossils in the Early Jurassic, providing only patchy insights into the early evolution of the group. Here, we describe new finds of comatulids from the late Pliensbachian to late Toarcian of Feuguerolles, Calvados, France. They include new species belonging to four superfamilies. The dominant genus is Palaeocomaster, with P. structus n. sp. represented by centrodorsals. For Palaeocomaster, the family Palaeocomasteridae n. fam. is proposed; it is placed in the Superfamily Comatuloidea. Solanocrinites is represented by S. jagti n. sp. The new taxa Andymetra toarcensis n. sp. and Spinimetra chesnieri n. g. et n. sp. are Family incertae sedis. The diagnoses of Comatuloidea and Solanocrinitoidea are modified based on the new material. The paracomatulid Singillatimetra truncata n. sp. is represented by a columnal, as is Forcipicrinus normannicus n. g. et n. sp. tentatively assigned to Pentacrinitina, Family incertae sedis. The material indicates that an important early comatulid radiation took place prior to the Toarcian Oceanic Anoxic Event, creating some basic types of these free-living crinoids. Along with the stratigraphic distribution patterns of the co-occurring stalked crinoids including the isocrinids Pentacrinites collenoti, Isocrinus (Chladocrinus) basaltiformis, the millericrinid Amaltheocrinus amalthei, and the cyrtocrinids Cotylederma docens, Eudesicrinus mayalis, Praetetracrinus inornatus, Sacariacrinus n. sp. and Tetracrinus n. sp., the new feather star finds suggest that the Toarcian Oceanic Anoxic Event was not of major influence to crinoid diversity.