Late Pleistocene Sites Research Articles

Species identification of voles and lemmings from Late Pleistocene deposits in Pin Hole Cave (Creswell Crags, UK) using collagen fingerprinting

Microfaunal remains are commonly used as palaeoenvironmental proxies and have been proposed as a means to identify relative ages of Late Quaternary deposits through biostratigraphy (i.e., utilising ‘Mammal Assemblage Zones’). However, assemblages of faunal remains can include a diverse range of taxa which are often difficult to distinguish using morphological aspects of the surviving bones, particularly when diagnostic features are eroded or missing due to fragmentation. Here we investigate the application of a recently developed method of species identification by collagen fingerprinting to arvicoline rodents (voles and lemmings), the dominant mammalian taxonomic group present at the Late Pleistocene site of Pin Hole Cave, Creswell Crags U.K., which is currently the designated British type locality for the Pin Hole Mammal Assemblage Zone (Oxygen Isotope Stage 3) fauna. We also further explore the potential for studying collagen decay rate via deamidations, and their applicability across the species boundary, in terms of its use for relative ageing of remains. Our results demonstrate the ease with which some taxa can be objectively distinguished to genus (e.g. Microtus from Myodes and Lemmus from Dicrostonyx) and in some cases species (e.g., M. gregalis from M. oeconomus), but that the potential for relative ageing is complicated by a range of taphonomic factors. The results highlight the potential for this new technique in much larger-scale palaeoenvironmental studies investigating temporal changes in vertebrate biodiversity.

The Middle to Late Pleistocene herpetofaunal assemblages from the Jarama and Manzanares valleys (Madrid, central Spain): An ecological synthesis

The successive fossil amphibian and reptile assemblages from the Middle to Late Pleistocene sites from the Manzanares and Jarama River Valleys (Madrid, central Spain) permitted the reconstruction of part of the climate instability with high-amplitude and rapid shifts of the last 450 ka and their associated landscapes: Áridos-1 (MIS11b), Valdocarros II (MIS8a/7e), Estanque de Tormentas de Butarque ETB-H02 (MIS7d or MIS6), PRERESA (MIS7/6 or MIS5a) and HAT (MIS5a). This work aims to present a regional synthesis of the palaeoclimatic and palaeoenvironmental data obtained from the herpetofaunal assemblages for these two valleys and their influence on the biodiversity during the Pleistocene. As a whole, these archaeo-paleontological localities document a total of 20 taxa (8 anurans and 12 reptiles) thus representing 76.9% of the modern authochtonous herpetofauna of the southeast of the Region of Madrid. Taking as a reference the modern situation in the area, the successive herpetofaunal assemblages permits a hypothetical landscape reconstruction where three different periods are represented with a glacial landscape (ETB-H02), a landscape of transition from cool to temperate climatic conditions (Valdocarros II), and an interglacial landscape (Áridos-1, HAT, PRERESA, and today). Environment is particularly open during dry periods, independently of if it is cold or warm. The main difference between an interglacial and a glacial period is the opposite representation of woodlands vs. moist environments: the last ones being more represented during cold periods than during warm periods. Finally, as documented by the succession from Valdocarros II, periods of transition between cold and warm climate are more forested but at the expense of humid meadows progressively. According to the relation between richness, biodiversity and climatic and environmental factors, a clear correlation appears between reptile richness and woodlands. In a similar way, mean annual precipitation (MAP) is revealed to be the most influent factor on reptile local diversity certainly because of its implications on vegetal cover extension and ecosystem productivity and resources. For amphibians the MAP does not influence dominancy but species richness: anuran richness being higher for lower MAP. Such an unusual pattern is certainly due to the fact that there are no strict forest-dweller anurans within the archaeological assemblages and that most of the anurans present in the sites are well adapted to arid conditions.

Discovery of circa 115,000-year-old bone retouchers at Lingjing, Henan, China.

Most Chinese lithic industries dated between 300,000 and 40,000 are characterized by the absence of Levallois debitage, the persistence of core-and-flake knapping, the rarity of prepared cores, their reduction with direct hard hammer percussion, and the rarity of retouched flakes. Here we report the discovery of seven bone soft hammers at the early hominin Lingjing site (Xuchang County, Henan) dated to 125,000–105,000. These artefacts represent the first instance of the use of bone as raw material to modify stone tools found at an East Asian early Late Pleistocene site. Three types of soft hammers are identified. The first consists of large bone flakes resulting from butchery of large herbivores that were utilized as such for expedient stone tools retouching or resharpening. The second involved the fracture of weathered bone from medium size herbivores to obtain elongated splinters shaped by percussion into sub-rectangular artefacts. Traces observed on these objects indicate intensive and possibly recurrent utilization, which implies their curation over time. The last consists of antler, occasionally used. Lingjing bone tools complement what we know about archaic hominin cultural adaptations in East Asia and highlight behavioural consistencies that could not be inferred from other cultural proxies. This discovery provides a new dimension to the debate surrounding the existence of the Middle Palaeolithic in the region. The attribution of East Asian sites to the Middle Palaeolithic assumes that cultural traits such as the Levallois method represent evolutionary hallmarks applicable to regions of the world different from those in which they were originally found. Here, we promote an approach that consists in identifying, possibly from different categories of material culture, the original features of each regional cultural trajectory and understanding the behavioural and cognitive implications they may have had for past hominin populations.

Los micromamíferos (Eulipotyphla, Chiroptera, Rodentia y Lagomorpha) del yacimiento del Pleistoceno Superior de la cueva de El Castillo (Cantabria, España)

El material de micromamíferos del yacimiento del Pleistoceno Superior de la cueva de El Castillo, objeto de estudio detallado en este trabajo, proviene de los niveles auriñacienses 18b y 18c (datados en 40.000-45.000 BP), nivel 19, y niveles musterienses 20b, 20c, 20d, 20e (datados en 41.000-49.000 BP), 21a y 21b. La asociación de micromamíferos es la siguiente: Erinaceus europaeus, Crocidura russula, Sorex coronatus, Sorex minutus, Neomys fodiens, Talpa europaea, Galemys pyrenaicus, cf. Miniopterus schreibersii, Chiroptera indet., Pliomys lenki, Microtus arvalis – Microtus agrestis, Microtus lusitanicus, Microtus oeconomus, Chionomys nivalis, Arvicola terrestris, Apodemus sylvaticus – Apodemus flavicollis y Lepus sp. La mayoría de estas especies están presentes en la fauna actual de Cantabria, excepto Pliomys lenki que se extinguió en el último tercio del Pleistoceno Superior, y Microtus oeconomus, que persistió en la península ibérica durante el Holoceno bien avanzado hasta su desaparición en tiempos históricos, y que actualmente está en regiones euroasiáticas más septentrionales. Hay una gran continuidad de la gran mayoría de los taxones en todos los niveles. La fauna en su conjunto parece indicar un medio predominantemente abierto, por lo general de praderas húmedas (y en mucha menor medida praderas secas) con una buena cobertura vegetal a nivel del suelo, en el que habría también algunas zonas arboladas o boscosas, y algún curso de agua. Los indicadores termófilos son escasos y poco abundantes, lo que podría indicar que el clima podría ser algo menos templado que en otros periodos del Pleistoceno Superior y quizás también que actualmente en la zona.

Equid prey acquisition and Archaic Homo adaptability at the early Late Pleistocene site of Xujiayao, China

AbstractBesides active hunting by hominins, there are two methods of scavenging that can provide primary access to intact ungulate carcasses: first‐access scavenging from nonpredator‐related accidents and early‐access aggressive scavenging from carnivore kills. Patterns in mortality profiles of prey animals at archaeological sites provide evidence of prey acquisition strategies, which is an important factor in reconstructing hominin adaptability. The Xujiayao site is an early Late Pleistocene site in the Nihewan Basin, China. Its faunal assemblage is dominated by Equus przewalskii and Equus hemionus. Evidence from previous taphonomic analysis has implied that Archaic Homo was the dominant taphonomic agent in the accumulation of the animal remains. This research applies mortality profile analysis to determine a follow‐up question: how were the equid carcasses acquired? The results of this study indicate that Archaic Homo at Xujiayao probably used both active hunting and scavenging to acquire equids in the early period (lower cultural layer) but mainly used active hunting in the later period (upper cultural layer) due to the fact that the climate changed to much colder conditions in the later period. This research provides new information on the subsistence adaptations of Archaic Homo in northern latitudinal climates of China in the early Late Pleistocene.

Quaternary fauna of the Eastern Adriatic (Croatia) with the special review on the Late Pleistocene sites

This paper presents an updated overview of fauna from the Early to Late Pleistocene/Holocene localities on the Eastern Adriatic coast and hinterland of Croatia. In this term 15 sites: four of Early (Šandalja I, Razvođe, Tatinja draga, and Podumci), two of Middle (Dubci, and Milovac), and nine of the Late Pleistocene age (Marlera I, Mujina pećina, Romualdova pećina, Vrtare male, Šandalja I + II, Vela spila, Ljubićeva pećina, Hijenska pećina, and Kopačina) have been selected and presented. The fossil mammal taxa prevail in all samples, followed by avifauna, while other groups of animals are less represented. On the basis of the palaeoecological remarks of different taxa the palaeoenvironmental and climate changes have been discussed through particular stages of the Early, Middle, and Late Pleistocene, and the beginning of the Holocene.

Late Pleistocene Human Migrations in China

Many archaeological and paleoanthropological discoveries have been made in China over the last 2 decades. Among these findings, I particularly note the recently excavated Late Pleistocene sites in the loess plateau in northern China and others found in a number of river basins in southern China. They all provide significant new information concerning Late Pleistocene human migrations across this vast region. A result of these excavations is the confirmation that flake- and pebble-tool industries dominated the region before the late marine isotope stage (MIS) 3. Small-flake-tool assemblages emerged suddenly during the late MIS 3 in South China. Blade industry first arrived in northwest China at the end of MIS 3, and microblade assemblages appeared in the loess plateau and the surrounding areas at a later stage. In this paper I briefly introduce the progress in Chinese Paleolithic archaeology and discusses Late Pleistocene human migrations and related issues.

Changing competition dynamics among predators at the late Early Pleistocene site Barranc de la Boella (Tarragona, Spain)

The late Early Pleistocene site Barranc de la Boella provides an unparalleled opportunity to assess the context of the activities of the hominin populations that inhabited the Iberian Peninsula at 1Ma. Recently, strong evidence for access to mammoth meat has been described at the Pit 1 locality. At the la Mina and el Forn excavation areas, little evidence exists for the anthropogenic processing of macromammals. However, the presence of humans is recorded, and the available evidence suggests these populations had access to several interesting resources. By analysing and comparing five separate assemblages at Barranc de la Boella, we assess the fluctuating presence of hominins and carnivores and the levels of competition among predators in each assemblage. Our analysis reveals different levels of competition intensity during the formation of assemblages when hominin groups were present, as evidenced by the abundance and diversity of stone artefacts. The analyses of skeletal component ratios indicate several competitive contexts, and the greatest presence of hominin groups is associated with the most competitive scenarios. The palaeoenvironment at Barranc de la Boella was rich in resources that hominins could exploit. The presence of hominin and carnivore groups appears to have been higher in levels with more inferred competition. This scenario supports prior research that concludes that carnivore abundance and highly competitive contexts were two constants in the lives of these hominin groups. Thus, the criteria determining whether hominins could inhabit a given landscape were most likely related to the presence or absence of resources, such as animal resources, water and raw materials, rather than the dynamics of the carnivore populations, to which the hominins were able to become habituated.

Latest Early Pleistocene wolf-like canids from the Iberian Peninsula

Several species of the genus Canis (Carnivora: Canidae) have been recorded from the European Early Pleistocene, but the phylogenetic relationships among them and in relation to extant members of this genus are still unclear. This is particularly true for the medium-sized and wolf-like extinct species Canis mosbachensis. It has been considered by many researchers as a descendant of the larger Canis etruscus and as a likely putative ancestor of extant wolves (Canis lupus). Other scholars, in contrast, have advocated instead for a closer relationship between C. mosbachensis and the extinct Canis arnensis, and even a close relationship between C. mosbachensis and C. lupus has been questioned. Here we describe the previously unpublished medium-sized Canis remains from the late Early Pleistocene site of Vallparadís Estació, along with additional new Canis material from the roughly coeval site of Cueva Victoria (both in the Iberian Peninsula), and compare them qualitatively and morphometrically with both extant and extinct species of this genus. The described material most closely resembles in craniodental size and shape the remains from Central and Southern Europe that have been previously assigned to C. mosbachensis, to which they are hence formally attributed. The excellent preservation of the newly described specimens (which include the most complete skull of this taxon) enables the description of features previously unknown for this species, which further support a close phylogenetic link with living wolves. Based on the described material, we review the role played by C. mosbachensis in the evolutionary history of European fossil canids, and conclude that this extinct species is most closely related to C. lupus and other closely-allied species, such as Canis anthus and Canis latrans.

Macaca sylvanus Linnaeus 1758 from the Middle Pleistocene of Quecchia Quarry (Brescia, Northern Italy)

During the Plio-Pleistocene the Barbary macaque, Macaca sylvanus Linnaeus, 1758 was widely distributed throughout Europe and North Africa (Szalay and Delson, 1979; Delson, 1980; Rook et al., 2001), and it became extinct in Europe during the Late Pleistocene (Elton and O’Regan, 2014). Nowadays this primate is still present in North Africa and a small population has been reintroduced at Gibraltar (Modolo et al., 2005). The taxonomy of the Plio-Pleistocene European macaques is still debated but many authors agree in considering all the fossils as belonging to the M. sylvanus lineage, while the Pleistocene endemic M. majori Azzaroli, 1946 from Sardinia (Italy) has been considered a distinct valid species (Rook and O’Higgins, 2005). The occurrence of this species is documented from Late Pliocene to Late Pleistocene sites located in Northern and Central Italy. Here we describe unpublished dentognathic remains of Macaca sylvanus from the Middle Pleistocene site of Quecchia Quarry (Botticino, Brescia, North Italy).

Late Pleistocene lithic technology in the Ivane valley: A view from the rainforest

Lithic assemblages in late Pleistocene sites within rainforest environments in Southeast Asia and Australasia are characterised as being simple core-and-flake technologies with little evidence for formal tools. This is usually attributed to the Bamboo Hypothesis, which proposes that modern humans first moving into these rainforest environments would have exploited available plant resources such as wood or bamboo at the expense of lithic technology. This paper challenges this portrayal by reporting the presence of formal tools in the Ivane valley of Papua New Guinea, dating from first occupation between 43,000–49,000 years cal BP.

Variations in Microtus arvalis and Microtus agrestis (Arvicolinae, Rodentia) Dental Morphologies in an Archaeological Context: the Case of Teixoneres Cave (Late Pleistocene, North-Eastern Iberia)

Morphological and morphometric variations in the first lower molars of Microtus arvalis and Microtus agrestis from the late Pleistocene site of Teixoneres Cave (Barcelona, Spain) have been investigated in order to understand the modifications in dental patterns occurring in these two species in a peripheral region of their distribution area. It was possible to identify along the sequence differences in size and frequencies of morphotypes within the two populations, corresponding to environmental and climatic oscillations. Hypotheses to explain these intraspecific changes are discussed, and the variations are ascribed primarily to small-scale intraregional movements of these two rodent populations.

Early Maritime Desert Dwellers in Namaqualand, South Africa: A Holocene Perspective on Pleistocene Peopling

ABSTRACTSouth Africa's northern Namaqualand coastal desert is the southern extension of the Namib. Today, this region is semi-desert with patchy subsistence resources and scarce, unpredictable rainfall. Yet this ancient desert landscape possesses residues of human activity stretching back into the Middle Pleistocene, evidenced by heavily weathered surface finds, including handaxes and Victoria West cores. Such old finds in so harsh an environment raise important questions: how do human movements into this area relate to local palaeoenvironmental changes, and how has this relationship changed through time? While no dated Middle Pleistocene sites presently exist to reconstruct the earliest hominin dispersals, several late Pleistocene sites now have chronostratigraphic sequences that can be brought to bear on these questions. This article presents chronological and subsistence-settlement data for one such site, Spitzkloof A Rockshelter in northern Namaqualand's rugged Richtersveld. Humans are shown to have visited the site very sporadically between ∼50,000 and 17,000 cal BP. Unlike most of the subcontinent, the most intensive occupations occur during early Marine Isotope Stage 2, when multiple proxies suggest enhanced humidity associated with intensified winter rainfall. We examine these data using the region's better-developed Holocene archaeological record to create predictions about the earliest coastal desert dwellers.

Villafranchian-Galerian mammal faunas transition in South-Western Europe. The case of the late Early Pleistocene mammal fauna of the Frantoio locality, Arda River (Castell’Arquato, Piacenza, Northern Italy)

We investigated the fossil mammal assemblage from the late Early Pleistocene (early Galerian) site of the Arda River. Seven taxa were identified: one carnivore (Ursus dolinensis), five artiodactyls (Sus strozzii, Pseudodama farnetesis, Praemegaceros sp., Bison sp., and Hippopotamus sp.) and one perissodactyl (Stephanorhinus hundsheimensis). The Arda River faunal assemblage includes taxa already present in the Villafranchian, such as Pseudodama and Sus strozzii. The bear U. dolinensis and the bison, a more evolved form than Bison (Eobison), are taxa typical of the beginning of the Galerian mammal age. Paleontological, sedimentological and paleobotanical data point towards the presence of woodlands settled on an alluvial plain with patches of grassland dated around 1.0 Ma. The continental sediments of the Arda River represent one of the first steps of the forming of the alluvial Po valley.

Optimality models and the food quest in Pleistocene Tasmania

The application of behavioural ecology models to deep time archaeological sites in Australia is often rendered difficult by the poor resolution of the archaeological record coupled with the imprecision in reconstructing past environments at regional levels. Here we report on a series of high latitude late Pleistocene sites in south-west Tasmania, where the conjunction of cultural activities and long term environmental change have preserved rich deposits in a number of cave sites. These data are used to test the utility of several models within optimal foraging theory. The data indicate the intensive and selective pursuit of two mid-size mammals, the Bennett’s wallaby and the wombat, at the expense of other prey species that were also likely present. Hunting patterns involved the scheduled seasonal use of lowland and upland valley sites underwriting a regional strategy of moving between focal hunting locations to maximise returns. This strategy continued with little change from c. 40,000BP to 15,000BP, providing a stable, predictable and secure food source for people through a period of extreme climatic change. The benefits and limitations of the behavioural ecology models used in this study indicate their utility where the data are sufficiently robust.

Biochronological data for the middle pleistocene site of Grotte de la Carrière (Lachambre karstic complex, Corneilla-De-Conflent, Eastern Pyrenees, Southern France) inferred from the small-mammal assemblage

The Embulla Mountain, located between the municipalities of Ria and Corneilla-de-Conflent (Eastern Pyrenees), possesses a large karstic complex with more than 20 caves situated at different altitudinal levels. Lachambre Cave, with more than 26 km of galleries, is the largest of these. The research carried out by the authors since 2011 has led to the discovery of several Middle to Late Pleistocene sites. Grotte de la Carriere is one of these newly discovered sites. The first paleontological survey, undertaken in May 2015, led to the unearthing of more than 2000 large- and small-vertebrate remains in eight different stratigraphical layers. The small-mammal assemblage from the richest layer of the sequence (layer 4) includes at least two insectivores (Talpa europaea and Sorex sp.) and eight rodents (Microtus arvalis, Microtus agrestis jansoni, Iberomys brecciensis, Clethrionomys glareolus, Pliomys coronensis, Apodemus gr. sylvaticus-flavicollis, Allocricetus bursae and Sciurus sp.). This association is very similar to other Middle Pleistocene sites in southern France, such as Caune de l'Arago and Grotte du Lazaret, as well as being similar to other Iberian Middle Pleistocene sites, such as Gran Dolina (TD10), Galeria and Sima del Elefante (TE18-19) in the Sierra de Atapuerca. These data suggest an approximate age for Grotte de la Carriere - layer 4 of between 220 and 450 ka. The chronology proposed on the basis of the micromammal assemblage also coincides with the large mammals recovered and identified from this layer, mainly composed of the species Ursus deningeri, Panthera fossilis and Canis mosbachensis.

Extinct Beringian wolf morphotype found in the continental U.S. has implications for wolf migration andevolution.

Pleistocene diversity was much higher than today, for example there were three distinct wolf morphotypes (dire, gray, Beringian) in North America versus one today (gray). Previous fossil evidence suggested that these three groups overlapped ecologically, but split the landscape geographically. The Natural Trap Cave (NTC) fossil site in Wyoming, USA is an ideally placed late Pleistocene site to study the geographical movement of species from northern to middle North America before, during, and after the last glacial maximum. Until now, it has been unclear what type of wolf was present at NTC. We analyzed morphometrics of three wolf groups (dire, extant North American gray, Alaskan Beringian) to determine which wolves were present at NTC and what this indicates about wolf diversity and migration in Pleistocene North America. Results show NTC wolves group with Alaskan Beringian wolves. This provides the first morphological evidence for Beringian wolves in mid‐continental North America. Their location at NTC and their radiocarbon ages suggest that they followed a temporary channel through the glaciers. Results suggest high levels of competition and diversity in Pleistocene North American wolves. The presence of mid‐continental Beringian morphotypes adds important data for untangling the history of immigration and evolution of Canis in North America.

Mountain strongholds for woody angiosperms during the Late Pleistocene in SE Iberia

Mediterranean mountains played an essential role during glacial periods as vegetation refugia. The SE Iberia Late Pleistocene woody angiosperm fossil and floristic evidences are reviewed in the context of phylogeographical studies aiming to identify (i) spatial patterns related to woody angiosperms glacial survival, (ii) structural and functional characteristics of montane refugia, and (iii) gaps in knowledge on the woody angiosperm patterns of survival in Mediterranean mountains. The distribution of palaeobotanical data for SE Iberia refugia has been found to be taphonomically biased due to the scarcity of available and/or studied high-altitude Late Pleistocene sites. However, Siles Lake data together with floristic inference provide evidences for woody angiosperms' survival in a high-altitude Mediterranean area. The main features boosting survival at montane contexts are physiographic complexity and water availability. Phylogeography studies have mainly been conducted at a continental scale. Although they cohere with palaeobotanical data to a broad scale, a general lack of sampling of SE Iberian range-edge populations, as well as misconceptions about the origin of the populations sampled, impede to infer the proper location of woody angiosperms' mountain refugia and their importance in the post-glacial European colonisation. We conclude that floristic, geobotanical, palaeobotanical, ethnographical and genetic evidence should be merged to gain a deeper understanding on the role played by Mediterranean mountains as glacial refugia in order to explain the current distribution of many plants and the large biodiversity levels encountered in Mediterranean mountain areas. This is hallmark for effective and efficient conservation and management.

New data on Late Pleistocene small vertebrates from northern France

Quaternary small vertebrate assemblages from northern France, and more generally from northwestern Europe, are poorly recognized, and studied less than those from more southern, Mediterranean areas. However, important sites of human occupation are often found in these northern areas, with significant issues related to the occupation of these sites by Neanderthals and previous humans, as well as faunal dynamics under climatic pressure. In this paper, we present preliminary results from the study of small vertebrate assemblages from two Late Pleistocene sites of northern France: Mutzig (Alsace) and Le Rozel (Normandy). Both are ancient rock-shelters that have been excavated recently and have yielded evidence of Neanderthal occupation, but in very different contexts: Le Rozel is located in a coastal area of northwestern France contemporaneous with a temperate period, while Mutzig is located at the foot of the Vosges mountains in northeastern France and was occupied during a cold period. Consequently, even if these two sites are chronologically close to each other, differences in faunal composition are observed and discussed in relation to the geographic, climatic and biochronological context, in comparing the results from small vertebrates with those from other disciplines (numerical dating, large mammals).

The oldest hominin butchery in European mid-latitudes at the Jaramillo site of Untermassfeld (Thuringia, Germany)

The late Early Pleistocene site of Untermassfeld, dated to the Jaramillo subchron (ca. 1.07 millions of years ago), is well known for its rich Epivillafranchian fauna. It has also recently yielded stone artefacts attesting hominin occupation. Now, we report here, for the first time, evidence of hominin butchery such as cut marks and intentional hammerstone-related bone breakage. This probable subsistence behaviour was detected in a small faunal subsample recovered from levels with Mode 1 stone tools. The butchered faunal assemblage was found during fieldwork and surveying in fluvial riverbanks (Lower Fluviatile Sands) and channel erosion sediments (Upper Fluviatile Sands). The frequent occurrence of butchery traces on bones of large-sized herd animals (i.e., Bison) may imply a greater need for meat in seasonal habitats characterised by a depletion of nutritive plants in winter. Early access to carcasses, before their consumption by carnivores, provided hominins with sufficient quantities of meat. This access was acquired with a Mode 1 lithic industry, to ensure food procurement and survival at high latitudes in Europe. Stone tools and faunal remains with signs of anthropic intervention recovered at Untermassfeld are evidence of the oldest hominin settlement at continental mid-latitudes (50° N).