Investigations on a considerable number of Wisconsin lakes have shown that, in many of them, more or less of the lower stratum, or hypolimnion, and the muddy ooze at the bottom possess no free oxygen for a certain time during the summer stagnation period (Birge and Juday, 'I I). This time varies from three or four weeks in some lakes to as many months in others. The dissolved oxygen disappears gradually from this stratum as the season advances and when it reaches a minimal amount the various organisms occupying this region respond to the change either by migrating or by adapting themselves to the new conditions. When the minimum is reached for plankton crustacea they rise to a higher level where oxygen is more abundant and when this gas becomes too scarce for some of the bottom dwelling insect larvae they migrate to the shallower water. The majority of the forms, however, are able to remain here even in the absence of free oxygen. To this latter group belong representatives of at least seventeen genera of protozoa, including rhizopods, flagellates and ciliates, but chiefly ciliates, and a number of higher forms, such as insect larvae, an ostracod, worms, and a bivalve mollusk. These forms appear to thrive as well in water which contains no free oxygen as in water which is well airated, thus being facultative anaerobes (Juday, 'o8). While making a study of the centrifuge plankton of Lake Mendota a ciliate was found in the lower stratum of water which appears to have gone beyond the facultative stage so that it has become substantially a true anaerobe. This ciliate agrees very closely in shape and structure Fig. 126, Plate X., of Conn's Protozoa of the Fresh Waters of Connecticut, which this author with hesitation places provisionally in the genus Enchelys. It has uniform ciliation, a rather pointed, obliquely truncate anterior end, and a rounded posterior extremity. Frequently
Read full abstract