Juniper Encroachment Research Articles

Climate warming and precipitation redistribution modify tree–grass interactions and tree species establishment in a warm‐temperate savanna

Savanna tree-grass interactions may be particularly sensitive to climate change. Establishment of two tree canopy dominants, post oak (Quercus stellata) and eastern redcedar (Juniperus virginiana), grown with the dominant C4 perennial grass (Schizachyrium scoparium) in southern oak savanna of the United States were evaluated under four climatic scenarios for 6years. Tree-grass interactions were examined with and without warming (+1.5°C) in combination with a long-term mean rainfall treatment and a modified rainfall regime that redistributed 40% of summer rainfall to spring and fall, intensifying summer drought. The aim was to determine: (1) the relative growth response of these species, (2) potential shifts in the balance of tree-grass interactions, and (3) the trajectory of juniper encroachment into savannas, under these anticipated climatic conditions. Precipitation redistribution reduced relative growth rate (RGR) of trees grown with grass. Warming increased growth of J. virginiana and strongly reduced Q. stellata survival. Tiller numbers of S. scoparium plants were unaffected by warming, but the number of reproductive tillers was increasingly suppressed by intensified drought each year. Growth rates of J. virginiana and Q. stellata were suppressed by grass presence early, but in subsequent years were higher when grown with grass. Quercus stellata had overall reduced RGR, but enhanced survival when grown with grass, while survival of J. virginiana remained near 100% in all treatments. Once trees surpassed a threshold height of 1.1m, both tiller number and survival of S. scoparium plants were drastically reduced by the presence of J. virginiana, but not Q. stellata. Juniperus virginiana was the only savanna dominant in which neither survival nor final aboveground mass were adversely affected by the climate scenario of warming and intensified summer drought. These responses indicate that climate warming and altered precipitation patterns will further accelerate juniper encroachment and woody thickening in a warm-temperate oak savanna.

Winter resource selection by female mule deer Odocoileus hemionus: functional response to spatio‐temporal changes in habitat

Populations of mule deer Odocoileus hemionus have declined throughout most of their historic range in recent decades, and habitat alteration has been hypothesized as one potential cause of those declines. Consequently, understanding how patterns of behaviour change as landscapes are altered through time may provide important insights into mechanisms underlying observed demographic trends in populations of mule deer. We examined resource selection in relation to habitat change by mule deer on the Tex Creek winter range in southeastern Idaho, USA. We created a GIS‐based map of habitats available to mule deer during two time periods, past (1985‐1986) and current (2007‐2009), to document changes in habitat over time. We then modeled past and current patterns of resource selection by mule deer based on locations obtained from visual observation, radio‐telemetry and GPS collars. Abundance and distribution of juniper Juniperus spp., aspen Populus tremuloides, sagebrush steppe Artemisia spp. and riparian habitat did not change significantly between past and current time periods. In contrast, we documented an increase in grassland from 3.5 to 30.7% of our study area, and a corresponding decrease in agricultural land, which provides high‐quality forage for mule deer, from 37.8 to 12.5% of our study area. Patterns of resource selection largely were similar between the two time periods. Nevertheless, mule deer significantly increased selection of agricultural fields and areas far from roads between the 1980s and 2007‐2009. In addition, juniper stands were strongly selected in all years, and importance values (use × availability rescaled to 100%) for grassland and sagebrush steppe increased between past and current time periods. Our results indicated that mule deer responded behaviourally to declining availability of high‐quality forage (i.e. agricultural land) by increasing selection of agricultural fields. Such functional responses in habitat selection may have important consequences for dynamics of mule deer populations, but additional research is necessary to link those responses to demography and population performance. Although juniper encroachment often is thought to have affected mule deer negatively on many summer ranges, strong selection for juniper in our study highlights the potential importance of that species on winter range, likely because it provides both thermal and hiding cover.

Relationships between landscape patterns and fire occurrence within a successional gradient in sagebrush steppe–juniper woodland

Expansion of western juniper (Juniperus occidentalis Hook. var. occidentalis) has altered vegetation composition, fire behaviour and fire potential throughout south-western Idaho and eastern Oregon. Utilising GIS-derived products and fire-simulation software, the influence of the spatial arrangement of different woodland developmental stages on simulated surface fire occurrence was evaluated. Custom fuel models and a recent vegetation map processed in FARSITE under moderate fire conditions were used to create a fire-occurrence grid in three sixth-order watersheds on the Owyhee Plateau of south-western Idaho. Landscape pattern metrics were selected to quantify the spatial arrangement of plant communities within a neighbourhood around points from each successional stage randomly placed within each watershed. Linear regression analysis of fire occurrence and each of the selected landscape metrics was compared for four successional stages of western juniper encroachment to assess the effect of landscape-scale vegetation arrangement on fire occurrence. The landscape structure had little influence on whether an early-successional area burns in a surface fire, whereas the surrounding landscape structure influenced whether a late-successional or mature woodland area burned. Landscape metrics that showed significance in late-successional and mature woodland stages include patch density, mean area and Simpson’s diversity.

Hydrologic Vulnerability of Sagebrush Steppe Following Pinyon and Juniper Encroachment

Abstract Woodland encroachment on United States rangelands has altered the structure and function of shrub steppe ecosystems. The potential community structure is one where trees dominate, shrub and herbaceous species decline, and rock cover and bare soil area increase and become more interconnected. Research from the Desert Southwest United States has demonstrated areas under tree canopies effectively store water and soil resources, whereas areas between canopies (intercanopy) generate significantly more runoff and erosion. We investigated these relationships and the impacts of tree encroachment on runoff and erosion processes at two woodland sites in the Intermountain West, USA. Rainfall simulation and concentrated flow methodologies were employed to measure infiltration, runoff, and erosion from intercanopy and canopy areas at small-plot (0.5 m 2 ) and large-plot (13 m 2 ) scales. Soil water repellency and vegetative and ground cover factors that influence runoff and erosion were quantified. Runoff and erosion from rainsplash, sheet flow, and concentrated flow processes were significantly greater from intercanopy than canopy areas across small- and large-plot scales, and site-specific erodibility differences were observed. Runoff and erosion were primarily dictated by the type and quantity of ground cover. Litter offered protection from rainsplash effects, provided rainfall storage, mitigated soil water repellency impacts on infiltration, and contributed to aggregate stability. Runoff and erosion increased exponentially ( r 2 = 0.75 and 0.64) where bare soil and rock cover exceeded 50%. Sediment yield was strongly correlated ( r 2 = 0.87) with runoff and increased linearly where runoff exceeded 20 mm·h −1 . Measured runoff and erosion rates suggest tree canopies represent areas of hydrologic stability, whereas intercanopy areas are vulnerable to runoff and erosion. Results indicate the overall hydrologic vulnerability of sagebrush steppe following woodland encroachment depends on the potential influence of tree dominance on bare intercanopy expanse and connectivity and the potential erodibility of intercanopy areas.

Estimating Juniper Cover From National Agriculture Imagery Program (NAIP) Imagery and Evaluating Relationships Between Potential Cover and Environmental Variables

Western juniper (Juniperus occidentalis subsp. occidentalis Hook.) woodlands are expanding from their historic range and causing significant declines of other plant communities. However, landscape-scale restoration projects are hindered by timeconsuming and expensive methods to inventory juniper cover and prioritize landscapes based on developmental phase of juniper encroachment. We investigated the ability of feature-extraction software to estimate western juniper cover from color aerial photographs obtained from the National Agriculture Imagery Program (NAIP) and explored the relationships between juniper cover at stand closure (potential juniper cover) and environmental/site indices (heat load, site exposure, and integrated moisture index) and characteristics measured from commonly available geospatial data layers. Estimates of juniper cover derived from NAIP imagery and ground measurements were similar (R 2 5 0.74; P , 0.01). Neither method consistently estimated juniper cover higher or lower than the other method (P 5 0.79). Environmental indices were either not correlated or weakly correlated with juniper cover at stand closure. However, the environmental/site characteristics (slope, aspect, and elevation) could be used to explain 40% of the variation in juniper cover at stand closure (R 2 5 0.40; P , 0.01). Thus, commonly available geospatial data layers can be used to assist in determining potential juniper cover. This information can then be compared to current juniper cover to determine juniper woodland developmental phase. Knowing the developmental phase is important because management strategies and effectiveness of restoration treatments differ among phases of juniper encroachment. Our results suggest that NAIP imagery can be a valuable tool to estimate juniper cover over large areas effectively to make landscape-scale restoration more feasible. The model of the relationship between environmental/site characteristics measured from commonly available geospatial data layers and potential juniper can be used to assist in restoration planning and prioritization, but could be improved with further refinement.

Characterizing Western Juniper Expansion via a Fusion of Landsat 5 Thematic Mapper and Lidar Data

Juniper encroachment into shrub steppe and grassland systems is one of the most prominent changes occurring in rangelands of western North America. Most studies on juniper change are conducted over small areas, although encroachment is occurring across large regions. Development of image-based methods to assess juniper encroachment over large areas would facilitate rapid monitoring and identification of priority areas for juniper management. In this study, we fused Landsat 5 Thematic Mapper and Light Detection and Ranging (lidar)–based juniper classifications to evaluate juniper expansion patterns in the Reynolds Creek Experimental Watershed of southwestern Idaho. Lidar applications for characterizing juniper encroachment attributes at finer scales were also explored. The fusion-based juniper classification model performed well (83% overall accuracy). A comparison of the resulting juniper presence/absence map to a 1965 vegetation cover map indicated 85% juniper expansion, which was consistent with tree-ring data. Comparisons of current and previous canopy-cover estimates also indicated an increase in juniper density within the historically mapped juniper distribution. Percent canopy cover of juniper varied significantly with land-cover types highlighting areas where intensive juniper management might be prioritized. El avance del Juniperus sobre las estepas arbustivas y los ecosistemas graminosos es uno de los cambios más prominentes ocurriendo actualmente en pastizales del Oeste de América del Norte. La mayoría de los estudios sobre cambios asociados al Juniperus se conducen en áreas pequeñas a pesar de que la invasión de esta especie está ocurriendo a escala de grandes regiones. El desarrollo de métodos basados en imágenes para relevar el avance del Juniperus a escala de áreas extensas facilitaría el monitoreo expeditivo y la identificación de áreas de manejo de Juniperus. En este ensayo fusionamos clasificaciones de Juniperus de imágenes Landsat 5 TM y LIDAR para evaluar los patrones de expansión del Juniperus en la cuenca experimental de Reynolds Creek del sudoeste de Idaho. Aplicaciones del LIDAR para caracterizar atributos de la invasión de Juniperus a escalas más finas también fueron exploradas. El modelo basado en la fusión de la clasificación de Juniperus tuvo un buen desempeño (83% de exactitud general). Una comparación del mapa de presencia/ausencia de Juniperus obtenido con este análisis con un mapa de cobertura de vegetación de 1965 indicó un 85% de expansión de Juniperus, patrón que fue consistente con datos de anillos de crecimiento de los árboles. La comparación de estimaciones de cobertura de canopeo actuales y pasadas también indicó un aumento en la densidad de Juniperus dentro del área de distribución histórica de Juniperus mapeada. El porcentaje de cobertura del canopeo de Juniperus varió significativamente con el tipo de cobertura del terreno remarcando áreas en las que el manejo intensivo del Juniperus podría ser priorizada.

Intercanopy community structure across a heterogeneous landscape in a western juniper‐encroached ecosystem

AbstractQuestions: How does landscape position influence biotic and abiotic attributes of western juniper (Juniperus occidentalis) encroached ecosystems? How does intercanopy plant community structure respond to changes in soil moisture and temperature based on juniper cover and topographic position?Location: Steens Mountain, southeast Oregon, USA.Objectives: Competition with western juniper modifies plant community composition, alters soil hydrology, and reduces plant productivity. Research is needed to understand these influences across heterogeneous landscapes. This study characterizes the relationship between juniper encroachment and soil water, soil temperature, topographic position, and intercanopy plant community structure.Methods: Using a completely randomized block design, plant density and cover, percent bare ground, percent soil moisture, soil temperature, heat accumulation, and elevation were sampled in 10 m2 plots representing low (<1%), moderate (∼14%), and high (∼27%) juniper cover at four aspects. The relationship and difference between vegetation patterns and environmental variables were analyzed using AOV, NMS, and MRPP (α=0.1). Indicator species analysis tested for shifts in dominant species along ecological gradients.Results: Soil moisture remained higher in low juniper cover sites than moderate and high juniper cover sites. North‐facing sites had highest soil moisture at 5 cm depth with low and moderate juniper cover levels. With increasing soil temperature from May to June, soil moisture declined by 19.7% at 5 cm depth. Achnatherum lemmonii and Pseudoreogneria spicata occurred in closed juniper stands while Achnatherum occidentale and Leymus cinereus were common when encroachment was limited.Application: This approach can be used to predict ecosystem response to western juniper encroachment across heterogeneous landscapes.

A Process-Based Application of State-and-Transition Models: A Case Study of Western Juniper (Juniperus occidentalis) Encroachment

Abstract A threshold represents a point in space and time at which primary ecological processes degrade beyond the ability to self-repair. In ecosystems with juniper (Juniperus L. spp.) encroachment, ecological processes (i.e., infiltration) are impaired as intercanopy plant structure degrades during woodland expansion. The purpose of this research is to characterize influences of increasing juniper on vegetation structure and hydrologic processes in mountain big sagebrush–western juniper (Artemisia tridentata Nutt. subsp. vaseyana [Rydb.] Beetle–Juniperus occidentalis Hook.) communities and to identify and predict states and thresholds. Intercanopy plant cover and infiltration rates were sampled in relation to juniper canopy cover. Study plots, arranged in a randomized complete-block design, represented low shrub–high juniper, moderate shrub–moderate juniper, and high shrub–low juniper percentage of canopy cover levels at four primary aspects. In field plots, percentage of plant cover, bare ground, and s...

Production of vegetation spatial-structure maps by per-object analysis of juniper encroachment in multitemporal aerial photographs

The remote sensing of vegetation, which has predominantly applied methods that analyze each image pixel as independent observations, has recently seen the development of several methods that identify groups of pixels that share similar spectral or structural properties as objects. The outputs of “per-object” rather than “per-pixel” methods represent characteristics of vegetation objects, such as location, size, and volume, in a spatially explicit manner. Before decisions can be influenced by data products derived from per-object remote sensing methods, it is first necessary to adopt methodologies that can quantify the spatial and temporal trends in vegetation structure in a quantitative manner. In this study, we present one such methodological framework where (i) marked point patterns of vegetation structure are produced from two per-object methods, (ii) new spatial-structural data layers are developed via moving-window statistics applied to the point patterns, (iii) the layers are differenced to highlight spatial-structural change over a 60 year period, and (iv) the resulting difference layers are evaluated within an ecological context to describe landscape-scale changes in vegetation structure. Results show that this framework potentially provides information on the population, growth, size association (nonspatial distribution of large and small objects), and dispersion. We present an objective methodological comparison of two common per-object approaches, namely image segmentation and classification using Definiens software and two-dimensional wavelet transformations.

Assessment of Juniper Encroachment With The Use of Satellite Imagery and Geospatial Data

Juniper encroachment into otherwise treeless shrub lands and grasslands is one of the most pronounced environmental changes observed in rangelands of western North America in recent decades. Most studies on juniper change are conducted over small areas, although encroachment is occurring throughout regions. Whether changes in juniper cover can be assessed over large areas with the use of long-term satellite data is an important methodological question. A fundamental challenge in using satellite imagery to determine tree abundance in rangelands is that a mix of trees, sagebrush, and herbaceous cover types can occur within a given image pixel. Our objective was to determine if spectral mixture analysis could be used to estimate changes in Rocky Mountain juniper (Juniperus scopulorum Sarg) and Utah juniper (Juniperus osteosperma [Torr.] Little) cover over 20 yr and 20 000 ha in southeast Idaho with the use of Landsat imagery. We also examined the spatial patterns and variation of encroachment within our study area using Geographic Information Systems–based data sets of grazing use, land-cover types, and topography. Juniper cover determined from 15-cm-resolution digital aerial orthophotography was used to train and validate juniper presence/absence classification in 1985 and 2005 Landsat images. The two classified images were then compared to detect changes in juniper cover. The estimated rate of juniper encroachment over our study area was 22–30% between 1985 and 2005, consistent with previous ground-based studies. Moran’s I analysis indicated that juniper encroachment pattern was spatially random rather than clustered or uniform. Juniper encroachment was significantly greater in grazed areas (P 5 0.02), and in particular in grazed shrub land cover type (P 5 0.06), compared to ungrazed areas. Juniper encroachment was also greater on intermediate slopes (10–35% slopes) compared to steeper or flatter terrain, and encroachment was somewhat less on north-facing (P 5 0.03) and more on west-facing (P 5 0.02) slopes compared to other aspects.

Shift in location of pygmy rabbit ( Brachylagus idahoensis) habitat in response to changing environments

We show how land use and potential climate changes occurring in sagebrush communities in the Great Basin of North America are affecting the pygmy rabbit, a sagebrush obligate species. We revisited 105 sites where pygmy rabbits were collected before 1950 and determined the current presence of the species using infrared-triggered camera surveys. Pygmy rabbits were present at 36% of the sites. Fourteen percent of the sites showed signs of pinyon–juniper woodland encroachment, with only one of these sites still harboring pygmy rabbits. Sites also showed current evidence of fires (16%), urbanization (13%), and agricultural conversion (6%). At a local scale, fire frequency reduction due to livestock grazing and fire suppression have shifted pygmy rabbit habitat to lower elevations. However, overall there was more extirpation at lower elevation sites, and the increase in mean elevation of 157 m closely corresponded to the predicted elevation increase with the rise in average global temperature seen over the last century. Warming temperatures may continue to raise lower elevation boundaries of habitable sites, while higher elevation sites are being increasingly impacted by pinyon–juniper encroachment. Predicted warming temperatures also may engender greater wildfire frequencies exacerbating habitat loss, the greatest threat to pygmy rabbit persistence.

Infiltration, Runoff, and Sediment Yield in Response to Western Juniper Encroachment in Southeast Oregon

Abstract Infiltration was measured in a western juniper ( Juniperus occidentalis Hook.) watershed to characterize the hydrologic processes associated with landscape position. Infiltration rate, runoff, and sediment content were measured with the use of a small-plot rainfall simulator. Study sites were located in each of the four primary aspects (north, south, east, and west). Research sites were located in two ecological sites—South Slopes 12–16 PZ and North Slopes 12–16 PZ. Within aspect, plots were located in three juniper cover levels: high (> 22%), moderate (13%–16%), and low ( −1 rate, levels comparable to an infrequent, short-duration, high-intensity precipitation event. Runoff was measured at 5-min intervals for 60 min. Comparing canopy cover levels, steady-state infiltration rates on control plots (9.0 cm · h −1 ) was 68% greater than high juniper cover sites (2.87 cm · h −1 ) and 34% greater than moderate juniper cover sites (5.97 cm · h −1 ) on south-facing slopes. On north-facing slopes, no differences in infiltration rates were observed between juniper cover levels, demonstrating differential hydrologic responses associated with ecological site. Generally, all water applied to control plots infiltrated. Highest infiltration rates were positively associated with increased surface litter and shrub cover. In addition, depth of water within the soil profile was lowest in high juniper cover plots. This suggests that less water is available to sustain understory and intercanopy plant growth in areas with high juniper cover. Accelerated runoff and erosion in juniper dominated sites (high level) across east-, west-, and south- facing slopes can lead to extensive degradation to the hydrology of those sites. These data suggest that sustained hydrologic processes are achieved with reduced western juniper canopy cover.

Markov models of land cover dynamics in a southern Great Plains grassland region

Grassland regions of the southern Great Plains are fragmented by agricultural activity and many habitat remnants have experienced encroachment by juniper (Juniperus virginiana L.). Recently, many cropland areas have been converted to monoculture grassland (pastures) and enrolled into the Conservation Reserve Program (CRP). Our objectives were to develop spatial and temporal Markov models to characterize land cover dynamics relative to juniper expansion and CRP using aerial photography from 1965, 1981, and 1995. We used landscapes surrounding three Breeding Bird Survey routes with varying levels of juniper encroachment in Oklahoma as study areas. As expected, land cover changes from 1965 to 1995 included increases in juniper woodland, mixed juniper-deciduous woodland, and pastures from CRP activity. Markov models revealed that juniper had a low likelihood of self-replacement in early stages of encroachment. In all areas, relatively little native grassland was lost to juniper encroachment, but other native habitat types such as deciduous woodland were heavily impacted. Transition probabilities for land cover dynamics varied significantly both spatially and temporally. Projections of these raw transition matrices produced widely varying models of future land cover conditions. By modifying the matrices to account for recent and potential socio-political and ecological changes occurring in this region, a number of more plausible land cover scenarios were produced than those resulting from simple projections of raw transition matrices.

INFLUENCE OF FIRE AND JUNIPER ENCROACHMENT ON BIRDS IN HIGH-ELEVATION SAGEBRUSH STEPPE

Abstract We examined relationships between high-elevation sagebrush (Artemisia spp.) steppe habitats altered by prescribed fire and western juniper (Juniperus occidentalis) encroachment on breeding distributions of Brewer's Sparrows (Spizella breweri), Vesper Sparrows (Pooecetes gramineus), Green-tailed Towhees (Pipilo chlorurus), and Sage Thrashers (Oreoscoptes montanus) on Steens Mountain in southeastern Oregon. In 2000 we conducted fixed-radius point count surveys at 172 sites encompassing burned and unburned sagebrush habitat and a range of juniper densities. For each bird species we developed habitat models using local variables measured in the field and landscape variables derived from remotely sensed data. Akaike's Information Criterion (AICc) was used to select the best-approximating model from a suite of a priori candidate models. Brewer's Sparrows, Sage Thrashers, and Green-tailed Towhees were positively related to increasing local sagebrush cover or percent sagebrush in the landscape, whereas V...

Herbaceous Restoration of Juniper Dominated Grasslands With Chaining and Fire

Juniper (Juniperus spp.) encroachment in grasslands usually progresses toward a stable woody state of mature trees that requires a significant disturbance to shift succession in another direction. Fire alone is often inadequate to shift succession in dense stands of mature juniper and must be preceded by a mechanical treatment such as chaining to reduce juniper competition and increase herbaceous growth that fuels a subsequent fire. However, little long-term data are available that measure combined effects of mechanical and fire treatments on restoration of juniper-dominated grasslands. Here, on a degraded (40% bare ground) north Texas site dominated by redberry juniper (Juniperus pinchotii Sudw), we quantify long-term herbaceous responses to mechanical chaining followed by fire. Two types of chaining, ground-level and elevated, were evaluated and all chained plots were burned 4 years after chaining. Herbaceous and woody responses were measured for several years after both chaining and fire treatments and compared to untreated controls. At study termination, both of the chaining + fire treatments reduced juniper cover from 32% to < 6%, but mortality was < 10%, because most plants basal-sprouted. Total grass production did not increase in chained treatments over the untreated until 3 growing seasons after chaining. Grass production declined the first growing season following the fire treatment, but increased in treated plots to 3 times the untreated the second and third year postfire. Total grass cover in treated plots did not increase over the untreated until the second year after the fire treatment. There was no difference in juniper or herbaceous responses between the 2 chain types. Results suggest increases in herbaceous production from chaining alone were due to increased growth of existing vegetation patches whereas the fire treatment appeared to stimulate herbaceous recruitment into bare soil areas.

Herbaceous Restoration of Juniper Dominated Grasslands With Chaining and Fire

Juniper (Juniperus spp.) encroachment in grasslands usually progresses toward a stable woody state of mature trees that requires a significant disturbance to shift succession in another direction. Fire alone is often inadequate to shift succession in dense stands of mature juniper and must be preceded by a mechanical treatment such as chaining to reduce juniper competition and increase herbaceous growth that fuels a subsequent fire. However, little long-term data are available that measure combined effects of mechanical and fire treatments on restoration of juniper-dominated grasslands. Here, on a degraded (40% bare ground) north Texas site dominated by redberry juniper (Juniperus pinchotii Sudw), we quantify long-term herbaceous responses to mechanical chaining followed by fire. Two types of chaining, ground-level and elevated, were evaluated and all chained plots were burned 4 years after chaining. Herbaceous and woody responses were measured for several years after both chaining and fire treatments and compared to untreated controls. At study termination, both of the chaining + fire treatments reduced juniper cover from 32% to &lt; 6%, but mortality was &lt; 10%, because most plants basal-sprouted. Total grass production did not increase in chained treatments over the untreated until 3 growing seasons after chaining. Grass production declined the first growing season following the fire treatment, but increased in treated plots to 3 times the untreated the second and third year postfire. Total grass cover in treated plots did not increase over the untreated until the second year after the fire treatment. There was no difference in juniper or herbaceous responses between the 2 chain types. Results suggest increases in herbaceous production from chaining alone were due to increased growth of existing vegetation patches whereas the fire treatment appeared to stimulate herbaceous recruitment into bare soil areas.

Managing Native Invasive Juniper Species Using Fire

Junipers (Juniperus spp.) are native woody shrubs that have expanded beyond their normal historical ranges in the western and southwestern United States since the late 1800s. Most ecologists and resource managers agree that juniper has become a deleterious native invasive plant that threatens other vegetation ecosystems, such as grasslands, through a steady encroachment and ultimate domination. The use of fire in managing junipers is based on a management goal to increase the disturbance return interval and thereby reduce the abundance and/or competitive impact of juniper in an ecosystem. In this paper, we discuss rates of juniper encroachment in relation to presettlement fire regimes, juniper encroachment and soil health, postfire vegetation responses, and long-term potential of different juniper treatment scenarios that involve prescribed fire.

Predicting juniper encroachment and CRP effects on avian community dynamics in southern mixed-grass prairie, USA

The probability of occurrence of 30 bird species was modeled as a function of landscape covertype in northwestern Oklahoma, USA. This grassland region has been extensively fragmented by agricultural activity, and remnant grassland patches are undergoing severe degradation from encroaching juniper ( Juniperus virginiana L.). In addition, many marginal or highly erodable croplands have been placed into perennial pasture dominated by exotic grasses under the Conservation Reserve Program (CRP). Based on temporal patterns of landscape change observed between 1965 and 1995, we estimated the covertype composition of the landscapes in the year 2015 under various CRP administrative and juniper expansion/control scenarios. We then used logistic regression to predict bird responses to these landscape composition estimates. Our estimates suggest that at the current rate of expansion, juniper will overtake substantial areas of remnant grassland even with extensive control measures. As a result, some obligate and facultative grassland birds are projected to decline, while numerous species tolerant of or partially reliant on woody vegetation will increase. Landscape dynamics due to changes in the CRP might be significant and could be designed to benefit declining grassland birds, but these benefits thus far are relatively minor compared to the effects encroaching juniper woodlands will have on the landscape and the avian community.

Landscape cover type and pattern dynamics in fragmented southern Great Plains grasslands, USA

We documented land cover and landscape pattern changes in an area of northwestern Oklahoma, USA using aerial photography from 1965, 1981, and 1995. This region of the southern Great Plains is fragmented by agricultural activity, and in recent years many remnant native grasslands have experienced extensive invasion by woody juniper (Juniperus virginiana L.). Concurrently, many cropland areas are being planted into perennial forage grasses and converted to intensively managed introduced grasslands as part of the U.S. Conservation Reserve Program (CRP). Our objectives were to document land cover and landscape pattern changes in the region relative to the expansion of juniper and CRP activity. We then examined how local landscape dominance by either anthropogenic or woody vegetation patches affected landscape pattern indices. Land cover changes from 1965 to 1995 included substantial increases in juniper woodlands and mixed woodlands that resulted from juniper encroachment into deciduous woodlands. Introduced grasslands also increased in many areas as a result of CRP implementation. Changes in landscape pattern generally reflected the influx of juniper into many areas. Landscapes dominated by woody vegetation had significantly more patches, smaller patches and patch core areas, more total edge, and higher patch diversity than landscapes dominated by anthropogenic cover types. Results indicate that expanding juniper is exacerbating the fragmentation process initiated by previous human activity, and represents a serious threat to the continued integrity and conservation of remaining southern Great Plains grasslands.

Juniper Encroachment into Aspen in the Northwest Great Basin

In the northwest Great Basin, western juniper (Juniperus occidentalis subsp. occidentalis Hook.) is encroaching into aspen (Populus tremuloides Michx.) communities. There is a concern that aspen communities in this region are in a state of decline, but their status has not been documented. This study determined the timing, extent, and some of the effects of this expansion. Ninety-one aspen stands were sampled for density, canopy cover, age, stand structure, and recruitment of western juniper and aspen. Soils and tree litter beneath aspen and western juniper were collected to analyze the effects of western juniper on soils. Additionally, 2 large aspen complexes in southeast Oregon were intensively aged to determine disturbance (fire) frequencies. Western juniper encroachment peaked between 1900 and 1939 with 77% of all juniper trees sampled having been established during this period. Three-fourths of aspen stands sampled have established populations of western juniper. Twelve percent of aspen stands sampled were completely replaced by western juniper and another 23% dominated by western juniper. Average density of western juniper in aspen sites was 1,573 trees ha(-1). Seventy percent of aspen stands sampled had zero recruitment of new aspen. Aspen stands averaged 98 years old. There was an inverse correlation between aspen canopy cover and western juniper canopy cover. Soils influenced by western juniper had a higher C:N ratio, pH, salts, lime, and sulfate, and lower amounts of magnesium, iron, copper, and manganese. Aspen litter had a lower C:N ratio than western juniper litter. Two major aspen complexes sampled had even-age, 2-tiered even-age, and multiple-age aspen trees. The absence of presettlement juniper within all sampled aspen stands suggests fire was the primary stand-replacing disturbance in these northwest Great Basin aspen communities. The lack of fire coupled with aspen stand decadence and low recruitment levels will allow for the continued encroachment and replacement of aspen communities by western juniper in the northwest Great Basin. DOI:10.2458/azu_jrm_v54i6_wall