Summary In Selliera radicans 5 sepals originate in a spiral sequence (Fig. 1). Alternating with them 5 petals arise simultaneously. From the beginning of petal development sympetaly is obvious (Fig. 3). The 5 episepalous stamens are initiated rapidly in a spiral sequence (Figs. 4,5). The gynoecium consists of 2 median carpels, which develop from an elliptic ring primordium situated on the margin of a funnel-shaped floral apex (Fig. 7). Parallel to the formation of a massive transversal septum, on which the ovules arise basipetally (Figs. 34, 35, 37), the ovary becomes increasingly inferior. In a relatively late stage of flower development a cup-like structure, the “indusium”, which is characteristic of Goodeniaceae (and Brunoniaceae) , develops near the tip of the style, enveloping the young stigma (Figs. 12—14). Through the late growth first of the filaments and then of the style — a sequence of events also known from Compositae, Campanulaceae and Lobeliaceae — the indusium takes up the pollen after anther dehiscense (Fig. 17). The pollen grains of each anther, stuck together by pollenkitt, slide into the indusium while the style elongates. When the indusium reaches the tips of the anthers, which are bent towards the centre of the flower, the pollen is pressed into it. The indusium then closes (Fig. 20). In open flowers the corolla forms a lower lip, whereas the style with its cupular indusium functions as an upper lip which is bent downwards (Fig. 19). Pollen is pressed out of the indusium by growth of the stigma (transition from the male to the female phase of the flower). In several characters of floral ontogeny Selliera resembles Brunoniaceae, Compositae and Campanulaceae. Such characters are for example the early sympetaly and the special mode of petal development (relatively early stop of apical growth, lengthening through a diffuse intercalary meristem below the tips, Fig. 36). With Compositae Selliera shares the tendency towards spiral inception in the androecium and the growth of the receptacle beneath the androecial primordia (Fig. 32, arrow)