Icelandic Shelf Research Articles

Structure of the Bottom Boundary Current South of Iceland and Spreading of Deep Waters by Submesoscale Processes

AbstractThe northeastern part of the North Atlantic subpolar gyre is a key passage for the Atlantic Meridional Overturning Circulation upper cell. To this day, the precise pathway and intensity of bottom currents in this area is not clear. In this study, we make use of regional high resolution numerical modeling to suggest that the main bottom current flowing south of Iceland originates from both the Faroe‐Banks Channel and the Iceland‐Faroe Ridge and then flows along the topographic slope. When flowing over the rough topography, this bottom current generates a 200 m large bottom mixed layer. We further demonstrate that many submesoscale structures are generated at the southernmost tip of the Icelandic shelf, which subsequently spread water masses in the Iceland Basin. These findings have major implication for the understanding of the water masses transport in the North Atlantic, and also for the distribution of benthic species along the Icelandic shelf.

Environmental preferences and critical habitat for the velvet belly lanternshark (Etmopterus spinax) in Icelandic waters.

The velvet belly lanternshark (Etmopterus spinax) is a small, bioluminescent shark that is caught as bycatch in many deep-sea fisheries in the Atlantic Ocean. Using data from 10,597 seasonal research survey tows spanning 11 years, the distribution, relative abundance, life history, and environmental preferences of E. spinax in Icelandic waters was examined for the first time. E. spinax (n = 8774) were only captured in relatively deep offshore waters to the south and west of Iceland. Females grew to larger sizes than males and reached 50% sexual maturity at a total length of 50 cm. Females at a late stage of maturity and very small juveniles (<20 cm) were restricted to the central south Icelandic shelf, suggesting that this might be critical habitat for the reproduction of the species. Most of the sharks were captured at depths of 400-500 m, a relatively narrow depth range, and classified as a stenothermic warm-water species with habitat temperature restricted to about 6.3-8.0°C. Teleosts, crustaceans and cephalopods made up most of the diet. There was no indication of a decline in abundance over the time span of the survey. However, climate-induced warming of the deep ocean may shift the distribution of the species to more northerly waters within Iceland.

Diversity under a magnifier lens: the case of Typhlotanaidae (Crustacea: Tanaidacea) in the N Atlantic

Research focusing on ‘stout-bodied’ typhlotanaids collected from several sites around Iceland and adjacent N Atlantic region has resulted in the description of 15 species new to science, as well as the creation of eight new genera. Typhlotanais eximius Hansen, 1913 is redescribed and transferred to a new genus, while Typhlotanais crassus and Peraeospinosus adipatus are transferred to the genus Larsenotanais. The morphological and the molecular data were combined to consolidate and confirm the validity of the results obtained from both approaches. The polyphyletic nature of the Typhlotanaidae and its serious of its taxonomic diversity are emphasized, although molecular analysis reveals that the ‘stout-bodied’ Typhlotanaidae form monophyletic clade. Depth and temperature are identified as the main environmental parameters determining the distribution of this group of Typhlotanaidae. Several species are clearly associated with the shelf and upper bathyal of Iceland. The Greenland-Iceland-Faroe Ridge is shown to be a distinct zoogeographical barrier for typhlotanaids inhabiting the deeper slope and abyssal regions around Iceland.

Detection and distribution of the North Atlantic capelin (Mallotus villosus) using environmental DNA—comparison with data from the main fishery management survey

AbstractCapelin (Mallotus villosus) is both an important commercial and ecological resource of the North Atlantic subpolar region. Two decades ago, the stock distribution around Iceland drastically changed. During autumn, which corresponds to the main feeding period, the capelin stock was previously located between the North of Iceland and the Jan Mayen area. Since the beginning of 2000s, the feeding aggregation has been located at the east coast of Greenland, inducing slight changes in the timing and route of the capelin spawning migration along the Icelandic shelf, and therefore in the catches. Changes in the distribution of capelin around Iceland made it both more difficult and expensive to assess the distribution of the stock with current survey methods. Here, we compare environmental DNA (eDNA) data to the acoustic data collected during the autumn monitoring survey, which leads to a preliminary estimation of the stock size. eDNA samples were collected at five different depths and were analyzed both horizontally across latitudes and longitudes and vertically across depth profiles. We detected eDNA in most of the locations where acoustic data detected capelin. Generalized linear models suggested that eDNA concentrations can be used as a proxy for the detection and quantification of capelin. The horizontal distribution of eDNA observed during both years corresponds with the horizontal distribution of capelin registered with the acoustic approach, while the vertical distribution indicated both effects of oceanic currents and diel vertical migration on eDNA detection and quantification.

Destabilisation of the Subpolar North Atlantic prior to the Little Ice Age

The cooling transition into the Little Ice Age was the last notable shift in the climate system prior to anthropogenic global warming. It is hypothesised that sea-ice to ocean feedbacks sustained an initial cooling into the Little Ice Age by weakening the subpolar gyre circulation; a system that has been proposed to exhibit bistability. Empirical evidence for bistability within this transition has however been lacking. Using statistical indicators of resilience in three annually-resolved bivalve proxy records from the North Icelandic shelf, we show that the subpolar North Atlantic climate system destabilised during two episodes prior to the Little Ice Age. This loss of resilience indicates reduced attraction to one stable state, and a system vulnerable to an abrupt transition. The two episodes preceded wider subpolar North Atlantic change, consistent with subpolar gyre destabilisation and the approach of a tipping point, potentially heralding the transition to Little Ice Age conditions.

Diatom-reconstructed summer sea-surface temperatures and climatic events off North Iceland during the last deglaciation and Holocene

Lateglacial and Holocene summer sea-surface temperature (SST) variability in the northern North Atlantic was reconstructed based on diatom records from two sediment cores from the North Icelandic shelf. The temperature changes are discussed in relation to palaeoceanographic circulation patterns and past climatic changes. Modern diatom data from surface sediments from around Iceland and Southeast and West Greenland, with known modern environmental variables, were used as the basis for the quantitative reconstruction of summer SST. The results show that summer SSTs varied during both the Lateglacial and the Holocene, but the amplitude of SST variations during the Lateglacial was greater than that during the Holocene. No pronounced warm or cool events were recorded on the North Icelandic shelf during the GI-1e to GI-1a events (Bølling-Allerød) and the GS-1 event (Younger Dryas). The changes in SST were possibly caused by major changes in the ocean circulation pattern around Iceland, and by minor variations in the interaction between the cold and warm currents in the region. Comparison of the reconstructed summer SSTs with other marine records from the North Atlantic suggests an antiphase relationship between the oceanic climatic responses off North Iceland and in the eastern North Atlantic during the Lateglacial and the earliest part of the Holocene (the Preboreal).

Marine mammal consumption and fisheries removals in the Nordic and Barents Seas

AbstractIn this study, we assess prey consumption by the marine mammal community in the northeast Atlantic [including 21 taxa, across three regions: (I) the Icelandic shelf, Denmark Strait, and Iceland Sea (ICE); (II) the Greenland and Norwegian Seas (GN); and (III) the Barents Sea (BS)], and compare mammal requirements with removals by fisheries. To determine prey needs, estimates of energetic requirements were combined with diet and abundance information for parameterizing simple allometric scaling models, taking uncertainties into account through bootstrapping procedures. In total, marine mammals in the ICE, GN, and BS consumed 13.4 [Confidence Interval (CI): 5.6–25.0], 4.6 (CI: 1.9–8.6), and 7.1 (CI: 2.8–13.8) million tonnes of prey year–1. Fisheries removed 1.55, 1.45, and 1.16 million tonnes year–1 from these three areas, respectively. While fisheries generally operate at significantly higher trophic levels than marine mammals, we find that the potential for direct competition between marine mammals and fisheries is strongest in the GN and weakest in the BS. Furthermore, our results also demonstrate significant changes in mammal consumption compared to previous and more focused studies over the last decades. These changes likely reflect both ongoing population recoveries from historic whaling and the current rapid physical and biological changes of these high-latitude systems. We argue that changing distributions and abundances of mammals should be considered when establishing fisheries harvesting strategies, to ensure effective fisheries management and good conservation practices of top predators in such rapidly changing systems.

Evolution and Transformation of the North Icelandic Irminger Current Along the North Iceland Shelf

AbstractThe North Icelandic Irminger Current (NIIC) flowing northward through Denmark Strait is the main source of salt and heat to the north Iceland shelf. We quantify its along‐stream evolution using the first high‐resolution hydrographic/velocity survey north of Iceland that spans the entire shelf along with historical hydrographic measurements as well as data from satellites and surface drifters. The NIIC generally follows the shelf break. Portions of the flow recirculate near Denmark Strait and the Kolbeinsey Ridge. The current's volume transport diminishes northeast of Iceland before it merges with the Atlantic Water inflow east of Iceland. The hydrographic properties of the current are modified along its entire pathway, predominantly because of lateral mixing with cold, fresh offshore waters rather than air‐sea interaction. Progressing eastward, the NIIC cools and freshens by approximately 0.3°C and 0.02–0.03 g kg−1 per 100 km, respectively, in both summer and winter. Dense‐water formation on the shelf is limited, occurring only sporadically in the historical record. The hydrographic properties of this locally formed water match the lighter portion of the North Icelandic Jet (NIJ), which emerges northeast of Iceland and transports dense water toward Denmark Strait. In the region northeast of Iceland, the NIIC is prone to baroclinic instability. Enhanced eddy kinetic energy over the steep slope there suggests a dynamical link between eddies shed by the NIIC and the formation of the NIJ as previously hypothesized. Thus, while the NIIC rarely supplies the NIJ directly, it may be dynamically important for the overturning circulation in the Nordic Seas.

Dense Water Formation on the Icelandic Shelf and Its Contribution to the North Icelandic Jet

AbstractThe North Icelandic Jet (NIJ) is the densest component of the Denmark Strait Overflow Water, feeding the abyssal limb of the Atlantic Meridional Overturning Circulation. Here, by using observational and numerical model data, we explore the formation of overflow water on the Icelandic shelf, the mechanisms involved, and its potential contribution to the NIJ. The sparse observational data on the western Icelandic shelf for the month of February shows top‐to‐bottom mixing on the shelf, distinct and well separated from the dense water offshore, with densities larger than 27.8 kg/m3 in some years. Using a 1‐D mixing model and winter heat flux reanalysis, we suggest that waters with densities exceeding 27.8 kg/m3 are likely to be formed on the shelf in most years by the end of winter. High‐resolution numerical model data shows that the transformation of the Atlantic inflow along the northwest Icelandic shelf generates a dense plume whose waters feed into the NIJ. The bulk of the plume cascades down‐slope north of Iceland, funneled through deep cross‐shelf troughs, with some cascading occurring west of Iceland as well. During years of strong cascading events (2008, 2013, and 2016), the modeled dense plume potentially feeds up to 21% of the NIJ transport at the Siglunes and Kögur sections. Back‐tracked Lagrangian particle trajectories confirm that the western Icelandic shelf is a source of the NIJ. The dense plume transport and variability are found to be dependent on the total oceanic heat loss west of Iceland and along Denmark Strait.

Trends and centennial-scale variability of surface water temperatures in the North Atlantic during the Holocene

Two sediment cores retrieved off North Iceland (western Nordic Seas) and on the eastern flank of Reykjanes Ridge (Iceland Basin) were analyzed to generate high-resolution alkenone-derived sea surface temperature (SST) records to investigate North Atlantic Ocean circulation changes during the Holocene. Early Holocene SSTs off North Iceland were unstable (10 ± 1 °C) and 3 °C warmer than today reflecting active northward heat transport of the Atlantic Meridional Overturning Circulation (AMOC) interrupted by intermittent Polar Waters incursions onto the North Icelandic shelf. The Holocene thermal optimum occurred synchronously east of Reykjanes Ridge, with a mean value of 11.5 °C (±0.5 °C) similar to today, consistent with a sustained influence of AMOC. Both records indicate that the circulation across the North Atlantic intensified between 8000 and 7000 yr BP. Thereafter, SSTs in the two basin sites broadly depict opposing trends and centennial-scale oscillations and a notable cooling at ∼5300 yr BP that coincides with Bond 4 event and the temporary collapse of the deep-water circulation. From 2500 yr BP onwards, SSTs in the Iceland Basin and the western Nordic Seas diverge leading to a marked cooling/warming dipole resulting in a temperature difference today of 4.5 °C. We show that SST trends and centennial-scale variability reflect variations of the subpolar gyre (SPG) circulation linked to drifting ice events and convection changes in the Labrador and Nordic Seas.

Response of biological productivity to North Atlantic marine front migration during the Holocene

Abstract. Marine fronts delineate the boundary between distinct water masses and, through the advection of nutrients, are important facilitators of regional productivity and biodiversity. As the modern climate continues to change, the migration of frontal zones is evident, but a lack of information about their status prior to instrumental records hinders future projections. Here, we combine data from lipid biomarkers (archaeal isoprenoid glycerol dibiphytanyl glycerol tetraethers and algal highly branched isoprenoids) with planktic and benthic foraminifera assemblages to detail the biological response of the marine Arctic and polar front migrations on the North Iceland Shelf (NIS) over the last 8 kyr. This multi-proxy approach enables us to quantify the thermal structure relating to Arctic and polar front migration and test how this influences the corresponding changes in local pelagic productivity. Our data show that following an interval of Atlantic water influence, the Arctic front and its associated high pelagic productivity migrated southeastward to the NIS by ∼6.1 ka. Following a subsequent trend in regional cooling, Polar Water from the East Greenland Current and the associated polar front spread onto the NIS by ∼3.8 ka, greatly diminishing local algal productivity through the Little Ice Age. Within the last century, the Arctic and polar fronts have moved northward back to their current positions relative to the NIS and helped stimulate the productivity that partially supports Iceland's economy. Our Holocene records from the NIS provide analogues for how the current frontal configuration and the productivity that it supports may change as global temperatures continue to rise.

A multi-decadal record of oceanographic changes of the past ~165 years (1850-2015 AD) from Northwest of Iceland.

Extending oceanographic data beyond the instrumental period is highly needed to better characterize and understand multi-decadal to centennial natural ocean variability. Here, a stable isotope record at unprecedented temporal resolution (1 to 2 years) from a new marine core retrieved off western North Iceland is presented. We aim to better constrain the variability of subsurface, Atlantic-derived Subpolar Mode Water (SPMW), using near surface-dwelling planktic foraminifera and Arctic Intermediate Water (AIW) mass changes using benthic foraminifera over the last ~165 years. The reconstruction overlaps in time with instrumental observations and a direct comparison reveals that the δ18O record of Neogloboquadrina pachyderma is reliably representing temperature fluctuations in the SPMWs. Trends in the N. pachyderma δ13C record match the measured phosphate concentration in the upper 200 m on the North Icelandic Shelf well. Near surface-dwelling foraminifera trace anthropogenic CO2 in the Iceland Sea by ~ 1950 ± 8, however, a reduced amplitude shift in the Marine Suess effect is identified. We argue that this is caused by a contemporary ongoing increase in marine primary productivity in the upper ocean due to enhanced Greenland’s freshwater discharge that has contributed to a nutrient-driven fertilization since the 1940s/50s (Perner et al., 2019). Multi-decadal variability is detected. We find that the 16-year periodicity evident in SPMW and AIWs based on the δ18O of N. pachyderma and M. barleeanum is a signal of SST anomalies propagated into the Nordic Seas via the Atlantic inflow branches around Iceland. Spectral analyses of the planktic foraminiferal δ13C signal indicate intermittent 30-year cycles that are likely reflecting the ocean response to atmospheric variability, presumably the East Atlantic Pattern. A long-term trend in benthic δ18O suggests that Atlantic-derived waters are expanding their core within the water column from the subsurface into deeper intermediate depths towards the present day. This is a result of increased transport by the North Icelandic Irminger Current to the North Iceland Shelf over the historical era.

When logbooks show the path: Analyzing the route and timing of capelin (Mallotus villosus) migration over a quarter century using catch data

Migration routes of exploited marine resources are often investigated using tracking methods such as satellite tags or Data Storage Tags (DSTs). However, this type of research crucially depends on the tagging and recapture effort, and is often unfeasible for small migratory pelagic fish such as capelin because of size constraints. We investigated the spawning migration route of capelin along the shelf break from the north and east of Iceland to spawning grounds located south and west of Iceland using catch data. Fishing vessels usually follow the front of the capelin migration as soon as it appears in fishable concentrations at the beginning of the year. This provides a good opportunity to assess the reliability of the catch data to assess the migration path of this species. The results show that capelin fishery starts in the first week of January along the Icelandic shelf and gradually moves towards the southern area in a clockwise manner, which is reflected in the fleet behavior. In addition, by analyzing catches for two different time periods encompassing more than two decades, we were able to detect slight changes in the timing and route of the capelin migration. We argue that in the case of small migratory pelagic fish, logbook data could be used to assess migration route and timing and its changes over time.

A 120 000-year record of sea ice in the North Atlantic?

Abstract. Although it has been demonstrated that the speed and magnitude of the recent Arctic sea ice decline is unprecedented for the past 1450 years, few records are available to provide a paleoclimate context for Arctic sea ice extent. Bromine enrichment in ice cores has been suggested to indicate the extent of newly formed sea ice areas. Despite the similarities among sea ice indicators and ice core bromine enrichment records, uncertainties still exist regarding the quantitative linkages between bromine reactive chemistry and the first-year sea ice surfaces. Here we present a 120 000-year record of bromine enrichment from the RECAP (REnland ice CAP) ice core, coastal east Greenland, and interpret it as a record of first-year sea ice. We compare it to existing sea ice records from marine cores and tentatively reconstruct past sea ice conditions in the North Atlantic as far north as the Fram Strait (50–85∘ N). Our interpretation implies that during the last deglaciation, the transition from multi-year to first-year sea ice started at ∼17.5 ka, synchronously with sea ice reductions observed in the eastern Nordic Seas and with the increase in North Atlantic ocean temperature. First-year sea ice reached its maximum at 12.4–11.8 ka during the Younger Dryas, after which open-water conditions started to dominate, consistent with sea ice records from the eastern Nordic Seas and the North Icelandic shelf. Our results show that over the last 120 000 years, multi-year sea ice extent was greatest during Marine Isotope Stage (MIS) 2 and possibly during MIS 4, with more extended first-year sea ice during MIS 3 and MIS 5. Sea ice extent during the Holocene (MIS 1) has been less than at any time in the last 120 000 years.

Marker tephra in Haukadalsvatn lake sediment: A key to the Holocene tephra stratigraphy of northwest Iceland

Emerging lake sediment records are rapidly improving Iceland‘s Holocene tephra stratigraphy and chronology. Here, we expand upon these recent developments with a securely-dated tephrochronological record from the lake Haukadalsvatn in West Iceland. We capitalize on the existing paleomagnetic secular variation (PSV) age model, which uses 54 ties points to synchronize Haukadalsvatn‘s high-resolution lake sediment record to a well-dated PSV master chronology from North Iceland Shelf marine core MD99-2269. Major element composition permits the identification of 38 tephra layers over the last 10500 years. Of these tephra, 15 can be traced to tephra found on the Vestfirðir peninsula and represent a series of well-dated marker tephra for West and Northwest Iceland that feature precise geochemical fingerprints. Based on these premises, and in contrast to recent assertations, there is no firm evidence for the presence of the presumed Middle Holocene „Hekla Ö tephra” in West or Northwest Iceland. Despite their close geographical locations, comparison of Haukadalsvatn‘s tephra layer frequency record to that of Northwest Iceland‘s reveals different trends. This regional disparity is likely the result of predominately small eruptions that featured narrower ash plumes. We argue that Haukadalsvatn serves as an important template for the tephrochronology of West Iceland and for future studies investigating the tephra records in more distal regions of the northwest North Atlantic, such as Greenland.

Sea Ice Control on Winter Subsurface Temperatures of the North Iceland Shelf During the Little Ice Age: A TEX86 Calibration Case Study

AbstractHolocene paleoceanographic reconstructions along the North Iceland Shelf have employed a variety of sea surface temperature and sea ice proxies. However, these surface proxies tend to have a seasonal bias toward spring/summer and thus only provide a discrete snapshot of surface conditions during one season. Furthermore, sea surface temperature proxies can be influenced by additional confounding variables resulting in markedly different Holocene temperature reconstructions. Here, we expand Iceland's marine paleoclimate toolkit with TEX86L: a temperature proxy based on the distribution of archaeal glycerol dibiphytanyl glycerol tetraether (GDGT) lipids. We develop a local Icelandic calibration from 21 surface sediment samples covering a wide environmental gradient across Iceland's insular shelves. Locally calibrated GDGT results demonstrate that (1) TEX86L reflects winter subsurface (0–200 m) temperatures on the North Iceland Shelf and (2) our calibration produces more realistic temperature estimates with substantially lower uncertainty (S.E. ±4 °C) over global calibrations. We then apply this new calibration to a high‐resolution marine sediment core (last millennium) collected from the central NIS with age control constrained by 14C‐dated mollusks. To test the veracity of the GDGT subsurface temperatures, we analyze quartz and calcite wt% and a series of highly branched isoprenoid alkenes, including the sea ice biomarker IP25, from the same core. The sediment records demonstrate that the development of thick sea ice during the Little Ice Age warmed the subsurface due to winter insulation. Importantly, this observation reflects a seasonal component of the sea ice/ocean feedback to be considered for the nonlinear cooling of the Little Ice Age in and around Iceland.

Tracking Holocene drift-ice limits on the northwest–southwest Iceland shelf: Comparing proxy data with observation and historical evidence

ABSTRACTWe detail variations in the weight percent (wt%) of quartz, a proxy for drift ice, in fifteen marine sediment cores from the northwest, west, and southwest Iceland shelf throughout the past 10 cal ka BP. We present the first map of iceberg distributions in Iceland waters between 1983 and 2011 and a new compilation of sea-ice records in the century from 850 to 1950 CE. The wt% of quartz, determined by quantitative X-ray diffraction (qXRD) analysis, is used to evaluate changes in the importation of drift ice. Small wt% of quartz were added to milled basalt (0% quartz), and to a mixture of non-clay and clay minerals; the qXRD method replicated 0 percent quartz, while measured 1–3 percent quartz always resulted in a “presence” estimate. The outer sites in the northwest sector lie close to the average position of the sea-ice margin between 1870 and 1920 CE; the southwest shelf sites lie south of this limit. Transects of cores along the Húnaflói and Djúpáll troughs indicate that the traces of drift ice decrease rapidly landward from the outer sites. The cores from the west/southwest of Iceland have limited amounts of quartz, generally possibly limited incursions of drift ice.

A Numerical Study of Interannual Variability in the North Icelandic Irminger Current

AbstractThe North Icelandic Irminger Current (NIIC) is an important component of the Atlantic Water (AW) inflow to the Nordic Seas. In this study, both observations and a high‐resolution (1/12°) numerical model are used to investigate the seasonal to interannual variability of the NIIC and its forcing mechanisms. The model‐simulated velocity and hydrographic fields compare well with the available observations. The water mass over the entire north Icelandic shelf exhibits strong seasonal variations in both temperature and salinity, and such variations are closely tied to the AW seasonality in the NIIC. In addition to seasonal variability, there is considerable variation on interannual time scales, including a prominent event in 2003 when the AW volume transport increased by about 0.5 Sv. To identify and examine key forcing mechanisms for this event, we analyzed outputs from two additional numerical experiments: using only the seasonal climatology for buoyancy flux (the momentum case) and using only the seasonal climatology for wind stress (the buoyancy case). It is found that changes in the wind stress are predominantly responsible for the interannual variations in the AW volume transport, AW fraction in the NIIC water, and salinity. Temperature changes on the shelf, however, are equally attributable to the buoyancy flux and wind forcing. Correlational analyses indicate that the AW volume transport is most sensitive to the wind stress southwest of Iceland.

Climate change opens new frontiers for marine species in the Arctic: Current trends and future invasion risks.

Climate change and increased anthropogenic activities are expected to elevate the potential of introducing nonindigenous species (NIS) into the Arctic. Yet, the knowledge base needed to identify gaps and priorities for NIS research and management is limited. Here, we reviewed primary introduction events to each ecoregion of the marine Arctic realm to identify temporal and spatial patterns, likely source regions of NIS, and the putative introduction pathways. We included 54 introduction events representing 34 unique NIS. The rate of NIS discovery ranged from zero to four species per year between 1960 and 2015. The Iceland Shelf had the greatest number of introduction events (n = 14), followed by the Barents Sea (n = 11), and the Norwegian Sea (n = 11). Sixteen of the 54 introduction records had no known origins. The majority of those with known source regions were attributed to the Northeast Atlantic and the Northwest Pacific, 19 and 14 records, respectively. Some introduction events were attributed to multiple possible pathways. For these introductions, vessels transferred the greatest number of aquatic NIS (39%) to the Arctic, followed by natural spread (30%) and aquaculture activities (25%). Similar trends were found for introductions attributed to a single pathway. The phyla Arthropoda and Ochrophyta had the highest number of recorded introduction events, with 19 and 12 records, respectively. Recommendations including vector management, horizon scanning, early detection, rapid response, and a pan‐Arctic biodiversity inventory are considered in this paper. Our study provides a comprehensive record of primary introductions of NIS for marine environments in the circumpolar Arctic and identifies knowledge gaps and opportunities for NIS research and management. Ecosystems worldwide will face dramatic changes in the coming decades due to global change. Our findings contribute to the knowledge base needed to address two aspects of global change—invasive species and climate change.

Two New Species of Recent and Upper Holocene Coccolith-agglutinated Foraminifera from the North Icelandic Shelf, North Atlantic

AbstractTwo new coccolith-agglutinated foraminiferal species, Pseudobolivina islandica n. sp. and Haplophragmoides atlanticus n. sp., are described from Recent and Upper Holocene sediments at Húnaflóaáll on the North Icelandic shelf. Specimens of the two species have been found only in the small (63–106 µm) fraction using the wet-counting method. This method allows preservation of these fragile tests, which are predominantly composed of coccoliths, mainly of the species Coccolithus pelagicus (Wallich, 1877).