Homo Erectus Research Articles

Why don’t we look to the west?

Paleontological data indicate that the beginning of Java Island’s human habitation took place at the Plio-Pleistocene boundary, around 2.4 Ma, along with uplift process and glacial-interglacial cycles. However, the oldest Homo erectus fossil was mainly found in the eastern part of Java Island where age-dating indicates that they were from ca. 1.5 Ma, especially along the riverbanks of Bengawan Solo and Brantas, such as Perning, Sangiran, Kedungbrubus, Ngandong, Ngawi, Trinil, and Sambungmacan.Recently, Pleistocene sites were discovered from the western part of Java, e.g., Rancah (Ciamis), Semedo (Tegal), and Bumiayu (Brebes) with their archeological, paleontological, and paleoanthropological potentials. This work will present the significance of the potential, especially paleoanthropological data from the new sites, and their implications to the Quaternary prehistory research strategies determination in the future.We present new geological, archeological, paleontological, and paleoanthropological evidence from those mentioned sites. The result shows that the distribution of Homo erectus were extended to the western part of Java, between 1.8-1.7 Ma, older than the oldest previous finding of Homo erectus from Perning and Sangiran. This finding suggests a new window of the human arrival on this island. So, why don’t we look to the west? Intensive research in the future should be addressed to the western part of Java Island.

Significance of Skull IX in Asian Paleoanthropology

Skull IX (Tjg-1993.05) was found in Sangiran, Central Java, in 1993, and is now stored at the Institute of Technology, Bandung. It is a remarkably complete fossil cranium of Homo erectus. It is one of the six H. erectus crania from the early Pleistocene of Java that preserves a nearly complete vault and is the third Javanese H. erectus crania with a substantial part of the facial skeleton preserved. After preliminary studies by late Prof. Sartono and other researchers, our team, led by Prof. Yahdi Zaim, had an opportunity to re-reconstruct and fully describe the specimen during 2008-2010. In this paper, I introduce how we corrected the distorted original reconstruction and summarize the specimen’s implications for the Homo erectus evolution in and outside Indonesia. Skull IX closely resembles Sangiran 17 and other Javanese H. erectus crania from the same stratigraphic level. The neurocranium of Skull IX is slightly smaller than the so-far-recorded smallest cranium from this zone, suggesting this individual was female. The face exhibits some “Asian” features, which highlights regional variation in Afro-Asian archaic members of Homo.

Hominin Femur finding from Banjarejo: Its morphological character and taxonomical position

Recently, Banjarejo became an important prehistoric site for Quaternary research because of its significant faunal and hominin remains. This study aims to describe a new finding of the hominin femur specimen from the site and its taxonomical position in human evolution. The specimen was identified by morphological and metric descriptions of the external feature of the femur. Then, comparative study to Homo erectus (n=2), Homo neanderthalensis (n=8), Homo heidelbergensis (n=1), prehistoric Homo sapiens (n=44), Australopithecus africanus (n=1), Paranthropus robustus (n=2), also non-human primate including Pongo (n=1), Macaca (n=1) and Gibbon (n=1) using bivariate and multivariate statistical analysis presented the specimen in the evolutionary perspective. This study shows the morphological and metric character of Banjarejo specimen located between Homo erectus and prehistoric Homo sapiens population sample. Further study should be addressed to investigate the cultural and chronological context of the hominin specimen.

Reconstruction and analysis of the DAN5/P1 and BSN12/P1 Gona Early Pleistocene Homo fossils

Two Early Pleistocene fossils from Gona, Ethiopia, were originally assigned to Homo erectus, and their differences in size and robusticity were attributed to either sexual dimorphism or anagenetic evolution. In the current study, we both revisit the taxonomic affinities of these fossils and assess whether morphological differences between them reflect temporal evolution or sexual variation. We generated virtual reconstructions of the mostly complete ∼1.55 Ma DAN5/P1 calvaria and the less complete 1.26 Ma BSN12/P1 fossil, allowing us to directly compare their anterior vault shapes using landmark-based shape analysis. The two fossils are similar in calvaria shape to H. erectus and also to other Early Pleistocene Homo species based on a geometric morphometric analysis of calvaria landmarks and semilandmarks. The DAN5/P1 fossil bears a particularly close affinity to the Georgian H. erectus fossils and to KNM-ER 1813 (H. habilis), probably reflecting allometric influences on vault shape. Combined with species-specific traits of the neurocranium (e.g., midline keeling, angular torus), we confirm that these fossils are likely early African H. erectus. We calculated regression-based estimates of endocranial volume for BSN12/P1 of 882–910 cm3 based on three virtual reconstructions. Although BSN12/P1 is markedly larger than DAN5/P1 (598 cm3), both fossils represent the smallest adult H. erectus known from their respective time periods in Africa. Some of the difference in endocranial volume between the two Gona fossils reflects broader species-level brain expansion from 1.77 to 0.01 Ma, confirmed here using a large sample (n = 38) of H. erectus. However, shape differences between these fossils did not reflect species-level changes to calvaria shape. Moreover, the analysis failed to recover a clear pattern of sexually patterned size or shape differences within H. erectus based on our current assessments of sex for individual fossils.

Comparative dental study between Homo antecessor and Chinese Homo erectus: Nonmetric features and geometric morphometrics

The Chinese Middle Pleistocene fossils from Hexian, Xichuan, Yiyuan, and Zhoukoudian have been generally classified as Homo erectus s.s. These hominins share some primitive features with other Homo specimens, but they also display unique cranial and dental traits. Thus, the Chinese Middle Pleistocene hominins share with other European and Asian hominin populations the so-called ‘Eurasian dental pattern’. The late Early Pleistocene hominins from Gran Dolina-TD6.2 (Spain), representing the species Homo antecessor, also exhibit the Eurasian dental pattern, which may suggest common roots. To assess phylogenetic affinities of these two taxa, we evaluated and compared nonmetric and metric dental features and interpreted morphological differences within a comparative hominin framework. We determined that the robust roots of the molars, the shelf-like protostylid, the dendrite-like pattern of the enamel-dentine junction surface of the upper fourth premolars and molars, the strongly folded dentine of the labial surface of the upper incisors, and the rare occurrence of a mid-trigonid crest in the lower molars, are all characteristic of Chinese H. erectus. With regard to H. antecessor, we observed the consistent expression of a continuous mid-trigonid crest, the absence of a cingulum in the upper canines, a complex root pattern of the lower premolars, and a rhomboidal occlusal contour and occlusal polygon and protrusion in the external outline of a large a bulging hypocone in the first and second upper molars. Using two-dimensional geometric morphometrics, we further demonstrated that H. antecessor falls outside the range of variation of Chinese H. erectus for occlusal crown outline shape, the orientation of occlusal grooves, and relative locations of anterior and posterior foveae in the P4s, P3s, M1s, M2s, and M2s. Given their geographic and temporal separation, the differences between these two species suggest their divergence occurred at some point in the Early Pleistocene, and thereafter they followed different evolutionary paths.

Existe-t-il un Paléolithique inférieur au Japon ?

All Lower Paleolithic sites discovered by Mr. Shinïchi FUJIMURA from 1981 to 2000 were falsified by himself. We wonder if Lower Paleolithic sites really exist (corresponding to a period of 30,000years ago in Japan) in the archipelago of Japan. We examined cultures of sites likely dating to the Early Upper Paleolithic and Lower Paleolithic discovered in the archipelago of Japan. While Japan was continuous with the continent in the Mindel glaciation (400,000years ago) or in the Riss glaciation (200,000years ago), Homo erectus or Homo heidelbergensis with the lithic industry of the Acheulean arrived in the archipelago of Japan. Thereafter, they survived on the islands of Japan cut the continent to the Upper Paleolithic while keeping the industry.

Le Paléolithique supérieur dans l’île d’Hokkaïdo

Since the discovery of the first Paleolithic site in Hokkaido, we consider that the time in the half more than centuries already passed, and float today. However, our research is still a way. By today, we have not discovered the trace about a Pithecantropines (Homo erectus) and a Paleanthropic man (Homo sapiens neanderthalensis) at all in this region. We know that a Neanthropic man (Homo sapiens sapiens) dominated all the continents except Antarctica in the final time of the Pleistocene era. And they passed along the dried shoal which appeared to the north in the same time from the Asian Continent to the Island of Hokkaido, and reached it to this place. The first attainment time is an age which exceeds 20,000-years before a little from the present. We are not checking a fact which exceeds it you to be much. However, the habitants of these beginnings left the trace of the activity of them in large numbers, so that he was wonderful to inside of this island. The studies-of-prehistory research in Hokkaido has little proof of the natural science field concerned on age determination or chronology. This has serious restriction. However, the tephra-chronology of volcanic ashes has played the big role uniquely. The eruption of the Eniwa volcano of the Hokkaido central part is especially a very important index. This volcano blows off ash fall (En-a) in large quantities, and covered all large ranges. This deposited volcanic-ashes layer plays a big role to discernment of an old industry and a recent industry on the cultural chronology. The obsidian from this island and nearby hard shale was used for many industries of this Upper Paleolithic cultures as main materials. Although obsidians in the island where the different four outcrops are known well. This rock was mined in large quantities, and was carried in various places. As well as many examples in the world, we divided this rock, and the human beings of the prehistoric age were demanded for it, and it was often utilized. In it, especially the outcrop of Shirataki was located in the northern part of Hokkaido, and was very important. Large quantities of obsidian from Shirataki outcrop is discovered in a large number of Paleolithic sites within a segment 700km in diameter at least. It covered the territory in Hokkaido and circulated further also except this island. This fact circulated as important trade materials on stone-tool fabrication among the human-beings groups from whom this island's every place differed. In this time, human beings’ activity in Hokkaido is a clear proof, which was not isolated at all. As for the studies of the prehistorians of Hokkaido distinguish, the exception put the technical process of eight different microblade productions. The technical process of some of them was limited only to the specific narrow area. Or it is proved that other technical processes are widely connected with the human-beings group of the Asian Continent on the contrary. At present, our research grasps the whole second-half Paleolithic culture, and has not yet established the year system edited by locally complete. The oldest industry of this area composed the typical stone tool of second-half Paleolithic culture from the beginning, and was accompanied by the manufacturing technique of a braid and microblade. The feature is the example in which the ruins of Kashiwadai-I were most excellent. These ruins correspond at time older than deposition of the Eniwa volcano tephra. It is located at the chill time in the second half of a Würm glacial period. The second continuing step is an industry of succession newer than this volcano tephra. It is raised as an example with Ôsatsu-16 typical ruins etc. The third phase is an industry of the end term of second-half Paleolithic culture. It is simultaneously located in the dawn of Jōmon culture. The feature of this cultural aspect is a transitional cultural stage which earthenware appears and is called what is called “Sub-Neolithic-Culture”. We can check having passed expansively, while the aspect of this last continues within a short period of time relatively.

How did language evolve in the lineage of higher primates?

Speech components emerged in the hominin lineage before the rise of modern human behavior and were already in place in monkey species. Evidence from genetics to archaeological records points to an accumulative increase of those computational properties required for modern language. At about 2.4 mya, the polytypical species Homo erectus sensu lato (s.l.) appeared with significant cortical growth indicated by neural migration factors and fossil skulls. The evidence suggests that early Homo erectus s.l. was equipped with a computational capacity for premodern language. The same species developed Acheulean toolmaking and showed signs of a symbolic and aesthetic mind at about half a mya. We conclude that the modern language capacity evolved at around 1 mya in the merging species late Homo erectus s.l. and pre-archaic Homo sapiens.

Late Middle Pleistocene Harbin cranium represents a new Homo species

Late Middle Pleistocene Harbin cranium represents a new Homo species

Massive cranium from Harbin in northeastern China establishes a new Middle Pleistocene human lineage

It has recently become clear that several human lineages coexisted with Homo sapiens during the late Middle and Late Pleistocene. Here, we report an archaic human fossil that throws new light on debates concerning the diversification of the Homo genus and the origin of H. sapiens. The fossil was recovered in Harbin city in northeastern China, with a minimum uranium-series age of 146 ka. This cranium is one of the best preserved Middle Pleistocene human fossils. Its massive size, with a large cranial capacity (∼1,420 mL) falling in the range of modern humans, is combined with a mosaic of primitive and derived characters. It differs from all the other named Homo species by presenting a combination of features, such as long and low cranial vault, a wide and low face, large and almost square orbits, gently curved but massively developed supraorbital torus, flat and low cheekbones with a shallow canine fossa, and a shallow palate with thick alveolar bone supporting very large molars. The excellent preservation of the Harbin cranium advances our understanding of several less-complete late Middle Pleistocene fossils from China, which have been interpreted as local evolutionary intermediates between the earlier species Homo erectus and later H. sapiens. Phylogenetic analyses based on parsimony criteria and Bayesian tip-dating suggest that the Harbin cranium and some other Middle Pleistocene human fossils from China, such as those from Dali and Xiahe, form a third East Asian lineage, which is a part of the sister group of the H. sapiens lineage. Our analyses of such morphologically distinctive archaic human lineages from Asia, Europe, and Africa suggest that the diversification of the Homo genus may have had a much deeper timescale than previously presumed. Sympatric isolation of small populations combined with stochastic long-distance dispersals is the best fitting biogeographical model for interpreting the evolution of the Homo genus.

Java Man and the Politics of Natural History

Abstract Natural history museums have long escaped postcolonial or decolonial scrutiny; their specimens were and are usually presented as part of the natural world, containing only biological or geological information. However, their collections, like those of other museums, are rooted in colonial practices and thinking. In this article, we sketch a political and decolonial biography of ‘Java Man’, the fossilized remains of a Homo erectus specimen, housed in Naturalis, the Natural History Museum, in the Netherlands. We describe the context of Dutch colonialism and the role of indigenous knowledge and activity in the discovery of Java Man. We also follow Java Man to the Netherlands, where it became a contested specimen and part of a discussion about repatriation. This article argues that the fossils of Java Man and their meanings are products of ‘creolized’ knowledge systems produced by Empire and sites of competing national and disciplinary histories and identities.

Editor's Column

Editor's Column

Asymmetries of Cerebellar Lobe in the Genus Homo

The endocast was paid great attention in the study of human brain evolution. However, compared to that of the cerebrum, the cerebellar lobe is poorly studied regarding its morphology, function, and evolutionary changes in the process of human evolution. In this study, we define the major axis and four measurements to inspect possible asymmetric patterns within the genus Homo. Results show that significant asymmetry is only observed for the cerebellar length in modern humans and is absent in Homo erectus and Neanderthals. The influence of occipital petalia is obscure due to the small sample size for H. erectus and Neanderthals, while it has a significant influence over the asymmetries of cerebellar height and horizontal orientation in modern humans. Although the length and height of the Neanderthal cerebellum are comparable to that of modern humans, its sagittal orientation is closer to that of H. erectus, which is wider than that of modern humans. The cerebellar morphological difference between Neanderthals and modern humans is suggested to be related to high cognitive activities, such as social factors and language ability.

Accessory cusp expression at the enamel-dentine junction of hominin mandibular molars.

Studies of hominin dental morphology frequently consider accessory cusps on the lower molars, in particular those on the distal margin of the tooth (C6 or distal accessory cusp) and the lingual margin of the tooth (C7 or lingual accessory cusp). They are often utilized in studies of hominin systematics, where their presence or absence is assessed at the outer enamel surface (OES). However, studies of the enamel-dentine junction (EDJ) suggest these traits may be more variable in development, morphology and position than previously thought. Building on these studies, we outline a scoring procedure for the EDJ expression of these accessory cusps that considers the relationship between these accessory cusps and the surrounding primary cusps. We apply this scoring system to a sample of Plio-Pleistocene hominin mandibular molars of Paranthropus robustus, Paranthropus boisei, Australopithecus afarensis, Australopithecus africanus, Homo sp., Homo habilis and Homo erectus from Africa and Asia (n = 132). We find that there are taxon-specific patterns in accessory cusp expression at the EDJ that are consistent with previous findings at the OES. For example, P. robustus M1s and M2s very often have a distal accessory cusp but no lingual accessory cusp, while H. habilis M1s and M2s show the opposite pattern. The EDJ also reveals a number of complicating factors; some apparent accessory cusps at the enamel surface are represented at the EDJ only by shouldering on the ridges associated with the main cusps, while other accessory cusps appear to have little or no EDJ expression at all. We also discuss the presence of double and triple accessory cusps, including the presence of a double lingual accessory cusp on the distal ridge of the metaconid in the type specimen of H. habilis (OH 7–M1) that is not clear at the OES due to occlusal wear. Overall, our observations, as well as our understanding of the developmental underpinnings of cusp patterning, suggest that we should be cautious in our comparisons of accessory cusps for taxonomic interpretations.

Body mass estimation from footprint size in hominins

Although many studies relating stature to foot length have been carried out, the relationship between foot size and body mass remains poorly understood. Here we investigate this relationship in 193 adult and 50 juvenile habitually unshod/minimally shod individuals from five different populations—Machiguenga, Daasanach, Pumé, Hadzabe, and Samoans—varying greatly in body size and shape. Body mass is highly correlated with foot size, and can be predicted from foot area (maximum length × breadth) in the combined sample with an average error of about 10%. However, comparisons among populations indicate that body shape, as represented by the body mass index (BMI), has a significant effect on foot size proportions, with higher BMI samples exhibiting relatively smaller feet. Thus, we also derive equations for estimating body mass from both foot size and BMI, with BMI in footprint samples taken as an average value for a taxon or population, estimated independently from skeletal remains. Techniques are also developed for estimating body mass in juveniles, who have relatively larger feet than adults, and for converting between foot and footprint size. Sample applications are given for five Pliocene through Holocene hominin footprint samples from Laetoli (Australopithecus afarensis), Ileret (probable Homo erectus), Happisburgh (possible Homo antecessor), Le Rozel (archaic Homo sapiens), and Barcin Höyük (H. sapiens). Body mass estimates for Homo footprint samples appear reasonable when compared to skeletal estimates for related samples. However, estimates for the Laetoli footprint sample using the new formulae appear to be too high when compared to skeletal estimates for A. afarensis. Based on the proportions of A.L. 288-1, this is apparently a result of relatively large feet in this taxon. A different method using a ratio between body mass and foot area in A.L. 288-1 provides estimates more concordant with skeletal estimates and should be used for A. afarensis.

Hominin homelands of East Java: Revised stratigraphy and landscape reconstructions for Plio-Pleistocene Trinil

Trinil (Java, Indonesia) yielded the type fossils of Homo erectus and the world’s oldest hominin-made engraving. As such, the site is of iconic relevance for paleoanthropology. However, our understanding of its larger geological context is unsatisfactory. Previous sedimentological studies are around 100 years old and their interpretations sometimes contradictory. Moreover, the existing stratigraphic framework is based on regional correlations, which obscure differences in local depositional dynamics. Therefore, a new and more local framework is urgently needed. We carried out a comprehensive geological study of the Trinil area. Using a Digital Elevation Model, we identified seven fluvial terraces. Terrace deposits were described and OSL-dated and fluvial behaviour was reconstructed. The terraces were correlated with terraces of the Kendeng Hills (e.g. the hominin-bearing Ngandong terrace) and date back to the past ∼350 ka. Thus far, most of the Trinil terraces and their deposits had remained unidentified, confounding sedimentological and stratigraphic interpretations. The exposed pre-terrace series has a thickness of ∼230 m. Together with the terraces, it forms a ∼3 Ma record of tectonism, volcanism, climate change and sea-level fluctuations. We subdivided the series into five new and/or revised stratigraphic units, representing different depositional environments: Kalibeng Formation, Padas Malang Formation, Batu Gajah Formation, Trinil Formation and Solo Formation. Special attention was paid to erosional contacts and weathering profiles, forming hiatuses in the depositional series, and offering insight into paleoclimate and base-level change. The Trinil Formation provides a new landscape context of Homo erectus. Between ∼550 and 350 ka, the area was part of a lake basin (Ngawi Lake Basin), separated from the marine base level by a volcanic barrier, under dry, seasonal conditions and a regular supply of volcanic ash. An expanding and retreating lake provided favourable living conditions for hominin populations. After 350 ka, this role was taken over by the perennial Solo River. Landscape reconstructions suggest that the Solo formed by headward erosion and stream piracy, re-connecting the Ngawi Lake Basin to the plains in the west. Our study offers a local framework, but its Pleistocene landscape record has regional significance. Most of all, it forms a much-needed basis for future, detailed studies on the build-up of the hominin site of Trinil, its fossil assemblages and numerical ages.

Gold Miners on the Trail of the Earliest Humans in Eastern Saharan Africa. Investigating the Acheulean and Middle Stone Age in Sudanese Nubia

Abstract This research note presents evidence for the oldest Middle Pleistocene Eastern Saharan human activity from the area referred to as the Eastern Desert Atbara River (EDAR), Sudan, which is currently threatened by gold mining. Preliminary results of multifaceted analyses indicate the activity of Homo sapiens during MIS 5 as well as Homo erectus during MIS 7–11 or earlier.

New hominin remains and revised context from the earliest Homo erectus locality in East Turkana, Kenya

The KNM-ER 2598 occipital is among the oldest fossils attributed to Homo erectus but questions have been raised about whether it may derive from a younger horizon. Here we report on efforts to relocate the KNM-ER 2598 locality and investigate its paleontological and geological context. Although located in a different East Turkana collection area (Area 13) than initially reported, the locality is stratigraphically positioned below the KBS Tuff and the outcrops show no evidence of deflation of a younger unit, supporting an age of >1.855 Ma. Newly recovered faunal material consists primarily of C4 grazers, further confirmed by enamel isotope data. A hominin proximal 3rd metatarsal and partial ilium were discovered <50 m from the reconstructed location where KNM-ER 2598 was originally found but these cannot be associated directly with the occipital. The postcrania are consistent with fossil Homo and may represent the earliest postcrania attributable to Homo erectus.

Shorter distal forelimbs benefit bipedal walking and running mechanics: Implications for hominin forelimb evolution.

Brachial index is a skeletal ratio that describes the relative length of the distal forelimb. Over the course of hominin evolution, a shift toward smaller brachial indices occurred. First, Pleistocene australopiths yield values between extant chimpanzees and humans, with further evolution in Pliocene Homo to the modern human range. We hypothesized that shorter distal forelimbs benefit walking and running performance, notably elbow and shoulder joint torques, and predicted that the benefit would be greater in running compared to walking. We tested our hypothesis in a modern human sample walking and running while carrying hand weights, which increase the inertia (mass and effective length) of the distal forelimb, simulating a larger brachial index. We found longer distal forelimbs and the added mass increased elbow muscle torque by 98% while walking and 70% in running, confirming our hypothesis that shorter distal forelimbs benefit walking and running performance. Shoulder muscle torque similarly increased in both gaits with the addition of hand weights due to elongation of the effective forelimb length. Normalized elbow torque, which accounted for the effect on shoulder torque caused by the experimental manipulation, increased by 16% while walking but 52% while running, indicating that shorter distal forelimbs provide a greater benefit for running by approximately three-fold. Selection for economical bipedal walking in Australopithecus and endurance running in Homo likely contributed to the shift toward relatively smaller distal forelimbs across hominin evolution, with modern human proportions attained in Pleistocene Homo erectus and retained in later species.

Assessing the status of the KNM-ER 42700 fossil using Homo erectus neurocranial development

Based on ontogenetic data of endocranial shape, it has been proposed that a younger than previously assumed developmental status of the 1.5-Myr-old KNM-ER 42700 calvaria could explain why the calvaria of this fossil does not conform to the shape of other Homo erectus individuals. Here, we investigate (ecto)neurocranial ontogeny in H. erectus and assess the proposed juvenile status of this fossil using recent Homo sapiens, chimpanzees (Pan troglodytes), and Neanderthals (Homo neanderthalensis) to model and discuss changes in neurocranial shape from the juvenile to adult stages. We show that all four species share common patterns of developmental shape change resulting in a relatively lower cranial vault and expanded supraorbital torus at later developmental stages. This finding suggests that ectoneurocranial data from extant hominids can be used to model the ontogenetic trajectory for H. erectus, for which only one well-preserved very young individual is known. However, our study also reveals differences in the magnitudes and, to a lesser extent, directions of the species-specific trajectories that add to the overall shared pattern of neurocranial shape changes. We demonstrate that the very young H. erectus juvenile from Mojokerto together with subadult and adult H. erectus individuals cannot be accommodated within the pattern of the postnatal neurocranial trajectory for humans. Instead, the chimpanzee pattern might be a better ‘fit’ for H. erectus despite their more distant phylogenetic relatedness. The data are also compatible with an ontogenetic shape trajectory that is in some regards intermediate between that of recent H. sapiens and chimpanzees, implying a unique trajectory for H. erectus that combines elements of both extant species. Based on this new knowledge, neurocranial shape supports the assessment that KNM-ER 42700 is a young juvenile H. erectus if H. erectus followed an ontogenetic shape trajectory that was more similar to chimpanzees than humans.