Higher Mean Annual Precipitation Research Articles

The empirical basis for modelling glacial erosion rates

Glaciers are highly effective agents of erosion that have profoundly shaped Earth’s surface, but there is uncertainty about how glacial erosion should be parameterised in landscape evolution models. Glacial erosion rate is usually modelled as a function of glacier sliding velocity, but the empirical basis for this relationship is weak. In turn, climate is assumed to control sliding velocity and hence erosion, but this too lacks empirical scrutiny. Here, we present statistically robust relationships between erosion rates, sliding velocities, and climate from a global compilation of 38 glaciers. We show that sliding is positively and significantly correlated with erosion, and derive a relationship for use in erosion models. Our dataset further demonstrates that the most rapid erosion is achieved at temperate glaciers with high mean annual precipitation, which serve to promote rapid sliding. Precipitation has received little attention in glacial erosion studies, but our data illustrate its importance.

Partial river flow recovery with forest age is rare in the decades following establishment.

Forest regeneration and expansion are occurring in many countries, with 80 million ha established from 2000 to 2012 under the Bonn accord and 17.5 million ha established from 1990 to 2005 according to the Food and Agriculture Organisation. Multiple reviews have linked increasing forest cover with reduced river flow and potentially detrimental effects downstream. Previous reviews have investigated trends in river flow response over time, but the influence of forest age remains uncertain. Partial river flow recovery (towards non‐forested conditions) has been reported in decades following forest establishment, but the role of climate in driving these trends has not been explored. Here, we evaluate river flow trends in 43 studies following forest establishment, which provide sufficient information to distinguish the effects of ageing forests from variable climate. Our meta‐analysis supports previous findings showing that forestation reduces annual river flow (by 23% after 5 years and 38% after 25 years) with greater reductions in catchments with higher mean annual precipitation, larger increases in forest cover, and which were idle, rather than agricultural land, prior to forestation. The impact of forests on river flow is sensitive to annual precipitation and potential evapotranspiration, but responses are highly variable. Forests affect river flow less when annual precipitation is low, and sensitivity to precipitation decreases as catchment aridity increases. The majority of catchments demonstrated persistent river flow declines after forest establishment. However, nine catchments showed partial flow recovery after an initial decrease, with peak flow reductions at an average age of 15 and across a range of tree species. The mean rate of recovery was 34 mm/year over 5 years. Partial flow recovery with forest age cannot be commonly expected, however, and forestation programmes should take into account that changes to annual river flow are likely to persist for up to five decades.

Climate, carbon content, and soil texture control the independent formation and persistence of particulate and mineral-associated organic matter in soil

Understanding the mechanisms controlling the formation and persistence of soil organic matter (SOM) is important for managing soil health and sustainable food production. The formation of SOM and the degree to which it is protected from decomposition are important for determining the long-term persistence of SOM. We used soils collected in a 13C-labelled litter decomposition study established at agricultural sites in Canada to understand the formation and persistence of newly-formed SOM. The ten agricultural sites spanned a wide range of soil carbon contents, texture, and climatic conditions. We fractionated the soil to isolate water extractable organic matter (WEOM), free light POM (fPOM), sand-sized and occluded particulate organic matter (oPOM), and silt and clay sized particles, referred to as mineral-associated organic matter (MAOM). Quantitative isotope tracing was used to determine the litter-derived C in all fractions. We performed these analyses early (six months after incubation) and later (five years after incubation) in the decomposition process to evaluate factors that control the formation and persistence of POM and MAOM. After six months litter-derived C was found in all fractions, but after five years it had declined in all fractions except the MAOM. Formation of MAOM was related to high mean annual precipitation and low sand content, whereas occluded POM formation was related to high soil C content. Persistence of MAOM and POM during the incubation were associated with low soil temperature and high soil C content. There was no consistent indication that formation of MAOM occurred from the decomposition of POM, suggesting that MAOM and POM are formed by two separate pathways. This has important implications for SOC models, which assume that plant-derived C passes through a sequence of pools, becoming more stable along the way.

Trace element and isotope geochemistry of macroelongatoolithid eggs as an indicator of palaeoenvironmental reconstruction from the Late Cretaceous Xixia Basin, China

The trace elements and isotopic compositions of macroelongatoolithid eggshells provide insights into the palaeoenvironment of these huge eggs preservation. We analysed the geochemical characteristics of newly discovered macroelongatoolithid eggs from the early Cretaceous Zhaoying Formation in the Xixia Basin, Henan Province, China. X-ray diffraction results reveal that the minerals of these eggs are predominately composed of calcite. The major elements in eggshells are Ca, Mg, K, Na, Al, Fe and Ti, and trace elements mainly include Sr, Ba, Mn, La, As, Sc, V, Cr, Co, Ni, Cu, Pb, Zn, Cd, U, and Ir. The high Sr and Ir concentrations could be caused by food materials and drinking water of dinosaurs. δ13C values of eggshells varying between −7.19‰ and −5.08‰ (average −5.69‰) indicate the C3 plants for dinosaurs with relatively high atmospheric CO2 concentration and low mean annual precipitation (MAP); further researches show that the carbon isotopic composition of atmospheric CO2 was around −4.11‰ to −2.19‰, and the MAP was lower than 10 mm/yr. δ18O values varying between −5.72‰ and −3.28‰ (average −4.53‰) show the temperature of nesting area were about 23 °C and the drinking water was dominated by leaf water with few meteoric water. The palaeoenvironment in which these huge eggs were preserved could be interpreted as continental deposit with extremely arid climate in the early Late Cretaceous Xixia Basin based on the high Fe contents of eggshells, the low palaeoclimate index of surrounding rocks, and the extremely low MAP shown by oxygen isotope.

Controls on granitic weathering fronts in contrasting climates

Granitic weathering profiles display highly diverse morphologies, reflecting the complex relationships between climate and weathering rates. Some profiles exhibit abrupt transitions from fresh bedrock to highly weathered material over short (<1 m) distances, while others exhibit only limited weathering extending 10s of meters into the bedrock. Although granitic weathering processes have been well studied, the controls on profile morphology and weathering rates within granitic, and many other lithologies remain poorly understood; these are likely influenced by a range of both intrinsic and extrinsic factors, which in turn will have crucial implications for understanding, for example, climate-weathering feedbacks. In this study we present multi-scale elemental and mineralogical analyses of a >30 m granitic weathering profile from the cool, temperate, Lysina catchment in the NW Czech Republic, from which we calculated mass transfer, weathering indices, and mineral specific weathering rates. The Lysina profile exhibits limited weathering extending >30 m into fractured bedrock, dominated by albite weathering at a rate of 9.3 × 10−17 mol m2 s−1.To identify environmental and geological controls on weathering front morphology and chemical weathering rates, Lysina was compared to previously published granitic weathering profiles from around the world. Weathering front morphology and weathering rates were calculated for the additional sites from published data and were correlated to mean annual precipitation (MAP), mean annual temperature (MAT), and erosion rates, with MAP having the strongest relationship. Higher MAP likely promotes lower saturation indices in pore waters, allowing weathering reactions to occur further from equilibrium. Comparison of erosion rates amongst the granitic catchments revealed an inconsistent effect on chemical weathering rates, but high erosion rates may promote weathering by reducing the thickness of the regolith and exposing the bedrock to reactive fluids. Mean annual temperatures appear to only have significant impacts on weathering fronts in environments with high precipitation and high erosion rates. Fractured bedrock profiles (Lysina and Río Icacos) have higher weathering intensities, than the other sites studied here. High connected porosity in fractured rocks enhances water movement allowing more efficient removal of weathering products, thus reducing thermodynamic saturation, increasing weathering rates, and producing sharper weathering gradients. These findings indicate that CO2 drawdown on geological timescales is also likely to be governed by precipitation rates, as well as temperature, and that much of the climate-significant weathering may occur within very narrow zones of the Earth's surface.

Survey of cattle and pasture management practices on focal pastures in Alberta

A survey of Alberta beef producers was conducted at sites overlapping with a province-wide network of permanent biodiversity monitoring plots to characterize focal pastures and their management, including estimates of stocking rates. Overall, greater stocking rates were reported in the boreal compared with the parkland and grassland natural regions, coinciding with an increased reliance on tame forage on relatively small land areas of largely deeded land. Higher stocking rates were also associated with earlier starting dates of grazing in the season, higher mean annual precipitation, and lower mean annual temperature.

Responses of soil organic carbon and nitrogen to land-use changes in a semiarid region of northwest China

Currently, we lack a clear understanding of how climate influences the responses of soil organic carbon (OC) and nitrogen (N) to land-use changes in semiarid regions. Thus, to clarify the mechanisms involved, we measured the soil OC and total N (TN) concentrations and stocks in the 0–100 cm soil layer in adjacent grassland, farmland, and woodland at five sites on the Loess Plateau in China, which varied in terms of the mean annual precipitation (MAP) and mean annual temperature (MAT). The farmland or woodland was converted from grassland at each site. Conversion of grassland to farmland significantly reduced the soil OC and TN, with greater decreases in sites with relatively lower MAP and MAT (–0.60 Mg OC ha−1 yr−1 and –0.22 Mg TN ha−1 yr−1, respectively) than sites with relatively higher MAP and MAT (–0.35 Mg OC ha−1 yr−1 and –0.04 Mg TN ha−1 yr−1). Conversion of grassland to woodland increased OC and TN at sites with higher MAP and MAT, but decreased them at sites with lower MAP and MAT. OC and TN increased to a depth of 100 cm with MAP and MAT after this land-use change. OC and TN decreased at relatively drier and colder sites, but increased at relatively wetter and warmer sites. These findings provide insights into the effects of land-use changes on the soil OC and TN under different climate conditions, thereby facilitating the sustainable management of grasslands in semiarid regions.

Global soil–climate–biome diagram: linking surface soil properties to climate and biota

Abstract. Surface soils interact strongly with both climate and biota and provide fundamental ecosystem services that maintain food, climate and human security. However, the quantitative linkages between soil properties, climate and biota remain unclear at the global scale. By compiling a comprehensive global soil database, we mapped eight major soil properties (bulk density; clay, silt, and sand fractions; soil pH; soil organic carbon, SOC, density; soil total nitrogen, STN, density; and soil C:N mass ratios) in the surface soil layer (0–30 cm), based on machine learning algorithms, and demonstrated the quantitative linkages between surface soil properties, climate and biota at the global scale, which we call the global soil–climate–biome diagram. In the diagram, bulk density increased significantly with higher mean annual temperature (MAT) and lower mean annual precipitation (MAP); soil clay fraction increased significantly with higher MAT and MAP; soil pH decreased with higher MAP and lower MAT and the “critical MAP”, which means the corresponding MAP at a soil pH of =7.0 (a shift from alkaline to acidic soil), decreased with lower MAT. SOC density and STN density were both jointly affected by MAT and MAP, showing an increase at lower MAT and a saturation towards higher MAP. Surface soil physical and chemical properties also showed remarkable variation across biomes. The soil–climate–biome diagram suggests shifts in soil properties under global climate and land cover change.

Geochemical characteristics of newly discovered Elongatoolithidae eggs from the Upper Cretaceous of Jiangxi Province, southern China: Palaeoenvironmental and palaeoclimatic inferences

We analyzed identified oospecies and associated geochemical characteristics of newly discovered dinosaur eggs from the Cretaceous Hekou Formation in Yudu District, Jiangxi Province, China. Results reveal that the Yudu eggs can be identified as representatives of the known oospecies Macroolithus rugustus within Elongatoolithidae based on external shape, size, ornamentation, and internal microstructure between the columnar and cone layers. The major element found in Macroolithus rugustus shells is Ca, while trace elements mainly include Sr, Ba, As, Cr, Mn, Pb, Cd, La, and Ir, characterised by Ir and Sr abnormalities, as well as the toxic elements As and Pb. Enrichment of trace elements in shells might be due to the diet of dinosaurs and subsequent deposition in eggs, consistent with global Ir anomalies and the regional enrichment of trace elements. δ13C values for Macroolithus rugustus ranging between −8.81‰ and −7.65‰ are indicative of C3 plants with relatively high CO2 concentration and low mean annual precipitation (MAP), while δ18O values ranging between −8.51‰ and 3.14‰ indicate relatively low humidity and the alteration of water sources of female dinosaur. The Sr and Ca contents as well as δ18O values of eggshells could be associated with the drinking water and food materials together with dissolved bone apatite. In light of the red-brown colour of surrounding rocks, high Fe composition revealed by macroelement analysis, and the presence of a C3 vegetation with low MAP, the palaeoenvironment of Yudu dinosaur nesting area was most likely to be fluvial and the palaeoclimate was semi-arid to arid, which is conducive to the preservation of eggs.

Veronico beccabungae-Mimuletum guttati, a new plant community in Slovakia

Vegetation with high coverage by the alien species &lt;em&gt;Mimulus guttatus&lt;/em&gt; was studied in the hilly regions of central Slovakia in 2015 and 2016. The floristic composition of these stands was recorded in ten phytosociological relevés corresponding to the &lt;em&gt;Veronico beccabungae-Mimuletum guttati&lt;/em&gt; (alliance &lt;em&gt;Glycerio-Sparganion&lt;/em&gt;) association. This association was reported for the first time in Slovakia in this study. This association was found to be closed or almost closed (mean coverage value of herb layer = 92%) and formed relatively species-rich stands (15 species per relevé), usually in narrow and small patches along the upper parts of streams and their spring areas in uplands at altitudes from 561 to 1,048 m. Localities with the &lt;em&gt;Veronico beccabungae-Mimuletum guttati&lt;/em&gt; association were characterized by typical mountain climates, with both relatively low mean annual air temperature (5.5°C) and high mean annual precipitation (885 mm). While water temperature (~13°C) of these habitats varied considerably among streams (7.4–19.9°C), their herbaceous vegetation preferred neutral to slightly alkaline water (pH = 6.1–7.5) with low amounts of soluble mineral matter (~72.6 μS cm&lt;sup&gt;−1&lt;/sup&gt;). This kind of vegetation was most often developed on sites with coarser sediments (stone, gravel, and sand) formed from crystalline bedrock. A comparison of vegetation data of the &lt;em&gt;Veronico beccabungae-Mimuletum guttati&lt;/em&gt; association across Central Europe demonstrated considerable floristic variability among regions.

The effects of increased snow depth on plant and microbial biomass and community composition along a precipitation gradient in temperate steppes

Shift in precipitation regime could greatly alter plant and microbial activity, and thus the contemporary and future ecosystem dynamics in grasslands. We investigated how changes in snow depth affect plants, microbes and their relationships after 10 consecutive years of snow treatments in different steppes. We selected 8 snow fences along a mean annual precipitation (MAP) gradient from 225 to 375 mm in Inner Mongolia. For each snow fence, study plots were set up at 7 transects with different levels of snow depth. We found that ecosystem properties, including soil moisture, the biomass and nitrogen (N) pools of microbes and plants, the fungi: bacteria ratio and the grass: forb ratio, increased with increasing snow depth at the drier sites with lower MAP, but not at the wetter sites with higher MAP. At any given site, the sensitivity of these ecosystem properties to changes in snow depth was determined by the slopes of these variables against snow depth. The results showed that the sensitivity of these ecosystem properties to changes in snow depth decreased linearly with increase in MAP levels. In addition, we also found that increased snow depth shifted the relationship between microbial and plant biomass from positive to negative. Our work reveals the importance of snow water in regulating plant and microbial processes in temperate steppes, especially under lower MAP conditions. The greater plant and microbial biomass and the shift of community toward greater fungi: bacteria and grass: forb ratio imply that increased snowmelt input alleviated water limitation in temperate steppes and altered plant and microbial communities. Our study helps to better predict that how changes in winter precipitation could affect the biomass and composition of plants and soil microbes in grasslands.

Response of Soils and Soil Ecosystems to the Pennsylvanian–Permian Climate Transition in the Upper Fluvial Plain of the Dunkard Basin, Southeastern Ohio, USA

Direct exposure of paleosols to the atmosphere during formation make them ideal for reconstructing paleoclimate. Paleosol and ichnofossil properties are dependently linked making it important to study them in tandem, to avoid errors in interpretation. Small scale studies (&lt;1 km) yield high resolution data that can be used to assess allogenic processes through comparison of spatial and temporal trends. This study used field and laboratory analyses to gather data from Late Pennsylvanian to Early Permian Upper Monongahela and Lower Dunkard group paleosols on Ohio Route 33 in Meigs County, OH, USA. The physical and geochemical properties of the paleosols from the field sites indicate that channel migration was the primary control on paleosol formation in the study area, however, a clear climate signal was observed. The change in paleosol type and calculated mean annual precipitation (MAP) values indicate that the climate became more strongly seasonal and drier over the course of the Pennsylvanian–Permian transition with a temporary excursion to a more ever-wet climate with higher MAP, marked by the occurrence of the Waynesburg Coal at the Pennsylvanian–Permian boundary.

Signatures of local adaptation along environmental gradients in a range-expanding damselfly (Ischnura elegans).

Insect distributions are shifting rapidly in response to climate change and are undergoing rapid evolutionary change. We investigate the molecular signatures underlying local adaptation in the range-expanding damselfly, Ischnura elegans. Using a landscape genomic approach combined with generalized dissimilarity modelling (GDM), we detect selection signatures on loci via allelic frequency change along environmental gradients. We analyse 13,612 single nucleotide polymorphisms (SNPs), derived from restriction site-associated DNA sequencing (RADseq), in 426 individuals from 25 sites spanning the I.elegans distribution in Sweden, including its expanding northern range edge. Environmental association analysis (EAA) and the magnitude of allele frequency change along the range expansion gradient revealed significant signatures of selection in relation to high maximum summer temperature, high mean annual precipitation and low wind speeds at the range edge. SNP annotations with significant signatures of selection revealed gene functions associated with ongoing range expansion, including heat shock proteins (HSP40 and HSP70), ion transport (V-ATPase) and visual processes (long-wavelength-sensitive opsin), which have implications for thermal stress response, salinity tolerance and mate discrimination, respectively. We also identified environmental thresholds where climate-mediated selection is likely to be strong, and indicate that I.elegans is rapidly adapting to the climatic environment during its ongoing range expansion. Our findings empirically validate an integrative approach for detecting spatially explicit signatures of local adaptation along environmental gradients.

The role of landscape heterogeneity in regulating plant functional diversity under different precipitation and grazing regimes in semi-arid savannas

1. Savanna systems exhibit a high plant functional diversity. While aridity and livestock grazing intensity have been widely discussed as drivers of savanna vegetation composition, physical soil properties have received less attention. Since savannas can show local differences in soil properties, these might act as environmental filters and affect plant diversity and ecosystem functioning at the patch scale. However, research on the link between savanna vegetation diversity and ecosystem function is widely missing.2. In this study, we aim at understanding the impact of local heterogeneity in soil conditions on plant diversity and on ecosystem functions. For this, we used the ecohydrological savanna model EcoHyD. The model simulates the fate of multiple plant functional types and their interactions with local biotic and abiotic conditions. We applied the model to a set of different landscapes under a wide range of livestock grazing and precipitation scenarios to assess the impact of local heterogeneity in soil conditions on the composition and diversity of plant functional types and on ecosystem functions.3. Comparisons between homogeneous and heterogeneous landscapes revealed that landscape soil heterogeneity allowed for a higher functional diversity of vegetation under conditions of high competition, i.e. scenarios of low grazing stress. However, landscape heterogeneity did not have this effect under low grazing stress in combination with high mean annual precipitation. Further, landscape heterogeneity led to a higher community biomass, especially for lower rainfall conditions, but also dependent on grazing stress. Total transpiration of the plant community decreased in heterogeneous landscapes under arid conditions.4. This study highlights that local soil conditions interact with precipitation and grazing in driving savanna vegetation. It clearly shows that vegetation diversity and resulting ecosystem functioning can be driven by landscape heterogeneity. We therefore suggest that future research on ecosystem functioning of savanna systems should focus on the links between local environmental conditions via plant functional diversity to ecosystem functioning.

Stable isotopes in large herbivore tooth enamel capture a mid-Miocene precipitation spike in the interior Pacific Northwest

Stable carbon and oxygen isotopes of fossil ungulate tooth enamel were used to investigate the paleoclimate of the Pacific Northwest from the mid-Oligocene through Pleistocene, particularly targeting the mid-Miocene Climatic Optimum (MMCO). We specifically test whether precipitation, as inferred from tooth enamel δ13C values relative to the δ13C values of atmospheric CO2 (Δ13C), increased with increasing partial pressure of atmospheric CO2 (pCO2). Analysis emphasized water-dependent equids, rhinos, and proboscideans, but several artiodactyls and Castor were also analyzed as available. Values of Δ13C>+19.5‰ in the lower John Day (~27 and ~18Ma) and Mascall (15–16Ma) localities imply relatively high mean annual precipitation (MAP=ca. 550–800mm/yr). Values of Δ13C<+18.5‰ at ~18Ma and at four levels between 15 and 3Ma imply low MAP (≤250mm/yr), similar to modern climate. High vs. low MAP generally correlates with high vs. low pCO2, implicating pCO2 as a probable driver of MAP in the Pacific Northwest. A climate oscillation model best explains data, such that warm-wet conditions during high pCO2 events alternated with cool-dry conditions during low pCO2 events on timescales of 100kyr. Because of poor time constraints associated with many of the specimens, changes in pCO2 are mainly inferred from marine isotopic data, rather than direct comparison between terrestrial pCO2 reconstructions and MAP. High δ18O values in specimens from the John Day (+21.8±1.0‰; VSMOW) and Mascall (+21.3±0.8‰) Formations could reflect lower elevations for the upwind Cascade Range prior to ~7Ma, or proximity to the coast compared to younger, more inland sites (δ18O=+17.7±0.9 to +19.6±1.1‰). Unusually high δ18O values of Dromomeryx sp. from Red Basin (+27.4±0.2‰) probably reflect drought tolerance. General circulation models commonly reproduce temperature but underestimate precipitation during the MMCO in the Pacific Northwest, suggesting that refinements are needed to predict more accurately how western North American precipitation will respond to increases in pCO2 during the next century.

The Early Pleistocene herpetofauna of Rivoli Veronese (Northern Italy) as evidence for humid and forested glacial phases in the Gelasian of Southern Alps

Among the Early Pleistocene Italian fossil herpetofaunas, Rivoli Veronese is remarkable in having yielded the youngest known allocaudates, represented by Albanerpeton pannonicum, together with remains of other amphibians and reptiles. The assemblage includes at least 15 other taxa, including two caudates (Speleomantes sp., Ichthyosaura alpestris), four anurans (Bombina sp., Bufo bufo, Hyla gr. H. arborea, Rana sp.), five lizards (Lacerta gr. L. viridis, cf. Zootoca vivipara, a third, small-sized lacertid, Anguis gr. A. fragilis, cf. Pseudopus sp.) and four snakes (Coronella sp., Zamenis longissimus, Natrix sp., Vipera gr. V. aspis). The finding of Speleomantes is particularly interesting as it is one of only a handful of fossil occurrences of this genus, being only the second one outside of its extant range. Excluding the extinct Albanerpeton and Speleomantes, which has a very narrow environmental range, the remainder of the assemblage suggests a Mean Annual Temperature rather similar or slightly colder than the present one in Rivoli Veronese (11.6°C versus 12.4°C), with much higher Mean Annual Precipitation (1322mm versus 834mm). The palaeoenvironment would have included either permanent or temporary water bodies, located in a forested area. More open landscapes might have also been present in the local vicinity. The palaeoherpetofauna of Rivoli Veronese supports the hypothesis of Albanerpeton favouring a moist environment, and confirms the presence of a humid forested landscape on the northern side of the Po Plain during the cold phases of the Gelasian, as already suggested by the pollen record for its southern margin.

Effects of climate, habitat and land use on the cover and diversity of the savanna herbaceous layer in Burkina Faso, West Africa

Herbaceous plants account for more than three-fourths of the total biomass in savanna ecosystems. However, which environmental factors mainly drive the biodiversity of herbaceous species is still under debate. In this study, we investigated the influence of climate, habitat and land use on species richness and cover in the West African savanna ecosystems of Burkina Faso. For a broader understanding, we analysed responses of the most important taxonomical groups of the herbaceous layer (Poaceae, Cyperaceae, Fabaceae s.l. and juvenile woody species) to the above mentioned environmental factors. We found light limitation by trees or shrubs and nutrient availability to be the variables that were mainly able to explain the differences in species richness, whereas precipitation and soil water-holding capacity, in general, drive species cover. However, specifically which environmental parameters influenced species richness and cover differed between taxonomical groups. Species richness of Cyperaceae species depended on water availability, while high Cyperaceae cover was found in soils with high water-holding capacity. Cyperaceae species are able to tolerate stagnant water. By contrast, Poaceae species cover was higher in dry habitats, and species richness declined with higher mean annual precipitation. Fabaceae species richness and cover were both higher in communal areas and sandy soils. This study revealed that both species richness and species cover, important aspects to describe the community composition, should be examined, as they are driven by sometimes different environmental conditions. Differentiating between the most important taxonomical groups of the savanna ecosystem increases the understanding in the important ecological processes that effect the herbaceous vegetation composition.

The size, distribution, and mobility of landslides caused by the 2015 Mw7.8 Gorkha earthquake, Nepal

Coseismic landslides pose immediate and prolonged hazards to mountainous communities, and provide a rare opportunity to study the effect of large earthquakes on erosion and sediment budgets. By mapping landslides using high-resolution satellite imagery, we find that the 25 April 2015 Mw7.8 Gorkha earthquake and aftershock sequence produced at least 25,000 landslides throughout the steep Himalayan Mountains in central Nepal. Despite early reports claiming lower than expected landslide activity, our results show that the total number, area, and volume of landslides associated with the Gorkha event are consistent with expectations, when compared to prior landslide-triggering earthquakes around the world. The extent of landsliding mimics the extent of fault rupture along the east-west trace of the Main Himalayan Thrust and increases eastward following the progression of rupture. In this event, maximum modeled Peak Ground Acceleration (PGA) and the steepest topographic slopes of the High Himalaya are not spatially coincident, so it is not surprising that landslide density correlates neither with PGA nor steepest slopes on their own. Instead, we find that the highest landslide density is located at the confluence of steep slopes, high mean annual precipitation, and proximity to the deepest part of the fault rupture from which 0.5–2Hz seismic energy originated. We suggest that landslide density was determined by a combination of earthquake source characteristics, slope distributions, and the influence of precipitation on rock strength via weathering and changes in vegetation cover. Determining the relative contribution of each factor will require further modeling and better constrained seismic parameters, both of which are likely to be developed in the coming few years as post-event studies evolve. Landslide mobility, in terms of the ratio of runout distance to fall height, is comparable to small volume landslides in other settings, and landslide volume-runout scaling is consistent with compilations of data on larger slope failures. In general, the size ratios of landslide source area to full landslide area are smaller than global averages, and hillslope length seems to largely control runout distance, which we propose reflects a topographic control on landslide mobility in this setting. We find that landslide size dictates runout distance and that more than half of the landslide debris was deposited in direct connection with stream channels. Connectivity, which is defined as the spatial proximity of landslides to fluvial channels, is greatest for larger landslides in the high-relief part of the High Himalaya. Although these failures are less abundant than those at lower elevations, they may have a disproportionate impact on sediment dynamics and cascading hazards, such as landslide reactivation by monsoon rainfall and landslide dams that lead to outburst floods. The overall high fluvial connectivity of coseismic landsliding in the Gorkha event suggests coupling between the earthquake cycle and sediment/geochemical budgets of fluvial systems in the Himalaya.

Variation in Essential Oil Yield and Composition of Dorema aucheri Boiss., an Endemic Medicinal Plant Collected from Wild Populations in Natural Habitats.

In the present research, variability in essential oil (EO) composition of five Dorema aucheri populations collected from natural habitats in different regions of Iran, were investigated. The EO content of populations varied from 0.28 to 0.68%. According to gas chromatography/mass spectrometry analysis, β-caryophyllene (7.17-35.73%), thymol (23.45-29.64%), β-gurjunene (2.58-5.89%), carvacrol (1.32-2.67%) and cuparene (1.97-2.98%) were the major components. Hierarchical cluster, principal component and canonical correspondence analyses classified the studied populations into three groups based on major EO components. The environmental parameters of the collected sites were also evaluated. According to the results, it might be suggested that sandy soils with high mean annual precipitation were major environmental factors influencing the amount of β-caryophyllene, while thymol, cuparene and caryophyllen oxide increased in silty and clay soils. Finally, the population collected in high altitudes and clay soils had higher amount of β-gurjunene.

Late Holocene evolution of glaciers in the southeastern Alps

ABSTRACTThe Julian Alps (in the southeastern European Alps, Italy and Slovenia) represent an important case study area for the study of small and very small maritime glaciers. High mean annual precipitation results in great snow accumulation during the winter, permitting the presence of ice bodies with the lowest Equilibrium Line Altitudes in the Alps. During the Little Ice Age (LIA) 19 small glaciers (<1 km2) existed, covering a total area of 2.4 km2. By 2012, the glacierized area had shrunk by 84% and only isolated glacierets and ice patches survived, each having a total area less than 0.5 km2. We present here a geomorphological and palaeoglaciological map of 8 sections of the Julian Alps related to the late Holocene distribution of glaciers, at a scale of 1:6000. Glacier topography during the LIA maximum was reconstructed on the basis of well-expressed geomorphological features together with historical archive data. The present-day distribution of ice bodies was inferred from orthophotos and 1 m resolution digital terrain models derived from airborne laser scanning. The past and present areal extent and surface morphology of glaciers permits calculation of volume loss since the LIA, which is estimated at 96%.