HI Deficiency Research Articles

Synthetic lethal mutants in Escherichia coli define pathways necessary for survival with RNase H deficiency.

Ribonucleotides frequently contaminate DNA and, if not removed, cause genomic instability. Consequently, all organisms are equipped with RNase H enzymes to remove RNA-DNA hybrids (RDHs). Escherichia coli lacking RNase HI (rnhA) and RNase HII (rnhB) enzymes, the ∆rnhA ∆rnhB double mutant, accumulates RDHs in its DNA. These RDHs can convert into RNA-containing DNA lesions (R-lesions) of unclear nature that compromise genomic stability. The ∆rnhAB double mutant has severe phenotypes, like growth inhibition, replication stress, sensitivity to ultraviolet radiation, SOS induction, increased chromosomal fragmentation, and defects in nucleoid organization. In this study, we found that RNase HI deficiency also alters wild-type levels of DNA supercoiling. Despite these severe chromosomal complications, ∆rnhAB double mutant survives, suggesting that dedicated pathways operate to avoid or repair R-lesions. To identify these pathways, we systematically searched for mutants synthetic lethal (colethal) with the rnhAB defect using an unbiased color screen and a candidate gene approach. We identified both novel and previously reported rnhAB-colethal and -coinhibited mutants, characterized them, and sorted them into avoidance or repair pathways. These mutants operate in various parts of nucleic acid metabolism, including replication fork progression, R-loop prevention and removal, nucleoid organization, tRNA modification, recombinational repair, and chromosome-dimer resolution, demonstrating the pleiotropic nature of RNase H deficiency. IMPORTANCE Ribonucleotides (rNs) are structurally very similar to deoxyribonucleotides. Consequently, rN contamination of DNA is common and pervasive across all domains of life. Failure to remove rNs from DNA has severe consequences, and all organisms are equipped with RNase H enzymes to remove RNA-DNA hybrids. RNase H deficiency leads to complications in bacteria, yeast, and mouse, and diseases like progressive external ophthalmoplegia (mitochondrial defects in RNASEH1) and Aicardi-Goutières syndrome (defects in RNASEH2) in humans. Escherichia coli ∆rnhAB mutant, deficient in RNases H, has severe chromosomal complications. Despite substantial problems, nearly half of the mutant population survives. We have identified novel and previously confirmed pathways in various parts of nucleic acid metabolism that ensure survival with RNase H deficiency.

Star Formation Activity of Galaxies Undergoing Ram Pressure Stripping in the Virgo Cluster

We study galaxies undergoing ram pressure stripping in the Virgo cluster to examine whether we can identify any discernible trend in their star formation activity. We first use 48 galaxies undergoing different stages of stripping based on HI morphology, HI deficiency, and relative extent to the stellar disk, from the VIVA survey. We then employ a new scheme for galaxy classification which combines HI mass fractions and locations in projected phase space, resulting in a new sample of 365 galaxies. We utilize a variety of star formation tracers, which include g - r, WISE [3.4] - [12] colors, and starburstiness that are defined by stellar mass and star formation rates to compare the star formation activity of galaxies at different stripping stages. We find no clear evidence for enhancement in the integrated star formation activity of galaxies undergoing early to active stripping. We are instead able to capture the overall quenching of star formation activity with increasing degree of ram pressure stripping, in agreement with previous studies. Our results suggest that if there is any ram pressure stripping induced enhancement, it is at best locally modest, and galaxies undergoing enhancement make up a small fraction of the total sample. Our results also indicate that it is possible to trace galaxies at different stages of stripping with the combination of HI gas content and location in projected phase space, which can be extended to other galaxy clusters that lack high-resolution HI imaging.

Angular momentum regulates H igas content and H icentral hole size in the discs of spirals

The neutral atomic hydrogen (HI) content of spiral galaxies has been observed to vary with environment, with more HI-deficient spirals residing in high density environments. This can be attributed to environmental effects such as ram pressure stripping and tidal interactions, which remove HI from the discs of galaxies. However, some spirals in low-density environments have also been observed to have relatively low HI mass fractions. The low densities of the Intergalactic Medium and lack of nearby galaxies in such environments make ram pressure stripping and tidal interactions unlikely candidates of gas removal. What then could be making these spirals HI deficient? Obreschkow et al. introduced a parameter-free model for the neutral atomic gas fraction ($f_{atm}$), in a symmetric equilibrium disc as a function of the global atomic stability parameter ($q$), which depends on specific angular momentum. In order to examine if this model accounts for HI-deficient galaxies in low-density environments, we have used the $M_{HI} ~-$ M$_{R}$ scaling relation to select six HI-deficient spiral galaxies and observed them with the ATCA. By measuring their $f_{atm}$ and $q$ values we find that the galaxies owe their observed HI deficiencies to low specific angular momenta. Additionally, we also find that the central HI hole sizes of our sample galaxies are related to their $q$ values, following the prediction of Obreschkow et al. This result brings to light the importance of angular momentum in understanding the physics of the interstellar medium in the discs of galaxies and consequently their evolution.

Morphological properties of galaxies in different Local Volume environments

We consider an all-sky sample of 1029 Local Volume (LV) galaxies situated within a distance of 11 Mpc. Their majority have precise distances, estimates of hydrogen mass fraction and star-formation rate derived from far-ultraviolet or Halpha fluxes. To describe an environment, we attribute two dimensionless values: the density contrast created by the most significant neighbour and the local density contrast produced by all neighbours within a separation of 1 Mpc. The hydrogen mass fraction exhibits a weak effect of HI deficiency being the most pronounced for dwarf irregular galaxies. The specific star-formation rate (sSFR) is more sensitive to the environment than the hydrogen mass fraction. Almost all (99 per cent) LV galaxies have their sSFR below -9.4 dex (1/yr). We notice that irregular dwarfs as well as late-type bulgeless galaxies are capable to reproduce their stellar mass with the observed sSFR over the cosmic time. Thus, the transformation of gas into stars in dIrs and spiral disks is rather sluggish unlike that in E, S0, dSph galaxies, whose star-formation history has been stormy and short. Scatter of SFR(Halpha)-to-SFR(FUV) ratio increases from Sc, Sd, Sm galaxies towards BCD, Im, Ir types that favours the idea of bursty star formations in low-mass galaxies. However, this bursty activity is caused rather by internal processes than by an external tidal action. A fraction of quenched E, S0, dSph galaxies increases from ~5 per cent in the field up to ~50 per cent in the densest regions.

Erratum: Choirs, H i galaxy groups: catalogue and detection of star-forming dwarf group members

The paper 'Choirs, HI galaxy groups: catalogue and detection of star-forming dwarf group members' was published in MNRAS 433(1), 543 559 (Sweet et al. 2013). Two methods for calculating the H I deficiency parameter were presented in Section 3.7. For method (1) there was an error in the transformation used to scale our Survey for Ionization in Neutral Gas Galaxies (Meurer et al. 2006) R-band (AB) magnitudes (MagRAB) to the SuperCosmos magnitudes (MagRSC) on which Denes, Kilborn &Koribalski (2014) was based. The transformation should be MagRSC = -8.52 + 0.63MagRAB. We note that Denes et al. ( 2014) subsequently presented an updated, slightly different R-band scaling relation, which does not affect our results. In Fig. 1 we show the replacement H I deficiency figure. Methods (1) (upper panel) and (2) (lower panel, scaling from H a luminosity and R-band surface brightness) are now in closer agreement. Our main previous finding is unchanged: the Choir groups are not significantly HI-deficient in either method for calculating HI deficiency. HIPASS J1059-09 is now HI deficient in both methods, though not at a significant level. As before, HIPASS J1403-06 was found to be marginally HI deficient by method (2) and not method (1), and is likely dominated by high H a equivalent width starbursting galaxies. We thank Robert Dzud zar for discovering the error.

The EAGLE simulations: atomic hydrogen associated with galaxies

We examine the properties of atomic hydrogen (HI) associated with galaxies in the EAGLE simulations of galaxy formation. EAGLE's feedback parameters were calibrated to reproduce the stellar mass function and galaxy sizes at $z=0.1$, and we assess whether this calibration also yields realistic HI properties. We estimate the self-shielding density with a fitting function calibrated using radiation transport simulations, and correct for molecular hydrogen with empirical or theoretical relations. The `standard-resolution' simulations systematically underestimate HI column densities, leading to an HI deficiency in low-mass ($M_\star < 10^{10}M_\odot$) galaxies and poor reproduction of the observed HI mass function. These shortcomings are largely absent from EAGLE simulations featuring a factor of 8 (2) better mass (spatial) resolution, within which the HI mass of galaxies evolves more mildly from $z=1$ to $0$ than in the standard-resolution simulations. The largest-volume simulation reproduces the observed clustering of HI systems, and its dependence on HI-richness. At fixed $M_\star$, galaxies acquire more HI in simulations with stronger feedback, as they become associated with more massive haloes and higher infall rates. They acquire less HI in simulations with a greater star formation efficiency, since the star formation and feedback necessary to balance the infall rate is produced by smaller gas reservoirs. The simulations indicate that the HI of present-day galaxies was acquired primarily by the smooth accretion of ionized, intergalactic gas at $z\simeq1$, which later self-shields, and that only a small fraction is contributed by the reincorporation of gas previously heated strongly by feedback. HI reservoirs are highly dynamic: over $40$ percent of HI associated with $z=0.1$ galaxies is converted to stars or ejected by $z=0$.

H i in group interactions: HCG 44

Extending deep observations of the neutral atomic hydrogen (HI) to the environment around galaxy groups can reveal a complex history of group interactions which is invisible to studies that focus on the stellar component. Hickson Compact Group 44 (HCG 44) is a nearby example and we have combined HI data from the Karoo Array Telescope, Westerbork Synthesis Radio Telescope, and Arecibo Legacy Fast ALFA survey, in order to achieve high column density sensitivity (N_HI < 2x10^18 cm^-2) to the neutral gas over a large field-of-view beyond the compact group itself. We find the giant HI tail north of HCG 44 contains 1.1x10^9 M_Sun of gas and extends 450 kpc from the compact group: twice as much mass and 33% further than previously detected. However, the additional gas is still unable to account for the known HI deficiency of HCG 44. The tail likely formed through a strong tidal interaction and HI clouds in the tail have survived for 1 Gyr or more after being stripped. This has important implications for understanding the survival of neutral clouds in the intragroup and circumgroup medium, and we discuss their survival in the context of simulations of cold gas in hot halos. HCG 44 is one of a growing number of galaxy groups found to have more extended HI in the intragroup and circumgroup medium than previously measured. Our results provide constraints for simulations on the properties of galaxy group halos, and reveal a glimpse of what will be seen by future powerful HI telescopes and surveys.

Star-forming dwarf galaxies in the Virgo cluster: the link between molecular gas, atomic gas, and dust

We present $^{12}$CO(1-0) and $^{12}$CO(2-1) observations of a sample of 20 star-forming dwarfs selected from the Herschel Virgo Cluster Survey, with oxygen abundances ranging from 12 + log(O/H) ~ 8.1 to 8.8. CO emission is observed in ten galaxies and marginally detected in another one. CO fluxes correlate with the FIR 250 $\mu$m emission, and the dwarfs follow the same linear relation that holds for more massive spiral galaxies extended to a wider dynamical range. We compare different methods to estimate H2 molecular masses, namely a metallicity-dependent CO-to-H2 conversion factor and one dependent on H-band luminosity. The molecular-to-stellar mass ratio remains nearly constant at stellar masses <~ 10$^9$ M$_{\odot}$, contrary to the atomic hydrogen fraction, M$_{HI}$/M$_*$, which increases inversely with M$_*$. The flattening of the M$_{H_2}$/M$_*$ ratio at low stellar masses does not seem to be related to the effects of the cluster environment because it occurs for both HI-deficient and HI-normal dwarfs. The molecular-to-atomic ratio is more tightly correlated with stellar surface density than metallicity, confirming that the interstellar gas pressure plays a key role in determining the balance between the two gaseous components of the interstellar medium. Virgo dwarfs follow the same linear trend between molecular gas mass and star formation rate as more massive spirals, but gas depletion timescales, $\tau_{dep}$, are not constant and range between 100 Myr and 6 Gyr. The interaction with the Virgo cluster environment is removing the atomic gas and dust components of the dwarfs, but the molecular gas appears to be less affected at the current stage of evolution within the cluster. However, the correlation between HI deficiency and the molecular gas depletion time suggests that the lack of gas replenishment from the outer regions of the disc is lowering the star formation activity.

BUDHIES – III: the fate of H i and the quenching of galaxies in evolving environments

In a hierarchical Universe clusters grow via the accretion of galaxies from the field, groups and even other clusters. As this happens, galaxies can lose their gas reservoirs via different mechanisms, eventually quenching their star-formation. We explore the diverse environmental histories of galaxies through a multi-wavelength study of the combined effect of ram-pressure stripping and group "processing" in Abell 963, a massive growing cluster at $z=0.2$ from the Blind Ultra Deep HI Environmental Survey (BUDHIES). We incorporate hundreds of new optical redshifts (giving a total of 566 cluster members), as well as Subaru and XMM-Newton data from LoCuSS, to identify substructures and evaluate galaxy morphology, star-formation activity, and HI content (via HI deficiencies and stacking) out to $3\times R_{200}$. We find that Abell 963 is being fed by at least 7 groups, that contribute to the large number of passive galaxies outside the cluster core. More massive groups have a higher fraction of passive and HI-poor galaxies, while low-mass groups host younger (often interacting) galaxies. For cluster galaxies not associated with groups we corroborate our previous finding that HI gas (if any) is significantly stripped via ram-pressure during their first passage through the intra-cluster medium, and find mild evidence for a starburst associated with this event. In addition, we find an overabundance of morphologically peculiar and/or star-forming galaxies near the cluster core. We speculate that these arise as groups pass through the cluster (post-processing). Our study highlights the importance of environmental quenching and the complexity added by evolving environments.

Disentangling the intragroup HI in Compact Groups of galaxies by means of X3D visualization

AbstractAs an extreme kind of environment, Hickson Compact groups (HCGs) have shown to be very complex systems. HI-VLA observations revealed an intrincated network of HI tails and bridges, tracing pre-processing through extreme tidal interactions. We found HCGs to show a large HI deficiency supporting an evolutionary sequence where gas-rich groups transform via tidal interactions and ISM (interstellar medium) stripping into gas-poor systems. We detected as well a diffuse HI component in the groups, increasing with evolutionary phase, although with uncertain distribution. The complex net of detected HI as observed with the VLA seems hence so puzzling as the missing one. In this talk we revisit the existing VLA information on the HI distribution and kinematics of HCGs by means of X3D visualization. X3D constitutes a powerful tool to extract the most from HI data cubes and a mean of simplifying and easing the access to data visualization and publication via three-dimensional (3-D) diagrams.

On the neutral gas content of nine new Milky Way satellite galaxy candidates

We use a new, improved version of the HI Parkes All-Sky Survey to search for HI emission from nine new, ultra-faint Milky Way satellite galaxy candidates recently discovered in data from the Dark Energy Survey. None of the candidates is detected in HI, implying upper limits for their HI masses of typically several hundred to a few thousand solar masses. The resulting upper limits on M_HI / L_V and M_HI / M_star suggest that at least some of the new galaxy candidates are HI deficient. This finding is consistent with the general HI deficiency of satellite galaxies located within the Milky Way's virial radius and supports the hypothesis that gas is being removed from satellites by tidal and ram-pressure forces during perigalactic passages. In addition, some of the objects may be embedded in, and interacting with, the extended neutral and ionised gas filaments of the Magellanic Stream.

Replication of the Escherichia coli chromosome in RNase HI‐deficient cells: multiple initiation regions and fork dynamics

DNA replication in Escherichia coli is normally initiated at a single origin, oriC, dependent on initiation protein DnaA. However, replication can be initiated elsewhere on the chromosome at multiple ectopic oriK sites. Genetic evidence indicates that initiation from oriK depends on RNA-DNA hybrids (R-loops), which are normally removed by enzymes such as RNase HI to prevent oriK from misfiring during normal growth. Initiation from oriK sites occurs in RNase HI-deficient mutants, and possibly in wild-type cells under certain unusual conditions. Despite previous work, the locations of oriK and their impact on genome stability remain unclear. We combined 2D gel electrophoresis and whole genome approaches to map genome-wide oriK locations. The DNA copy number profiles of various RNase HI-deficient strains contained multiple peaks, often in consistent locations, identifying candidate oriK sites. Removal of RNase HI protein also leads to global alterations of replication fork migration patterns, often opposite to normal replication directions, and presumably eukaryote-like replication fork merging. Our results have implications for genome stability, offering a new understanding of how RNase HI deficiency results in R-loop-mediated transcription-replication conflict, as well as inappropriate replication stalling or blockage at Ter sites outside of the terminus trap region and at ribosomal operons.

Hα3: an Hαimaging survey of HI selected galaxies from ALFALFA

We present the analysis of Halpha3, an Halpha imaging survey of 409 galaxies selected from the HI Arecibo ALFALFA Survey in the Local Supercluster, including the Virgo cluster. We explore the relations between the stellar mass, the HI mass and the current, massive SFR of nearby galaxies in the Virgo cluster and we compare them with those of isolated galaxies in the Local Supercluster, disentangling the role of the environment in shaping the star formation properties of galaxies at the present cosmological epoch. We investigate the relationships between atomic neutral gas and newly formed stars in different environments, across many morphological types, and over a wide range of stellar masses adopting an updated calibration of the HI deficiency parameter. Studying the mean properties of late-type galaxies in the Local Supercluster, we find that galaxies in increasing local galaxy density conditions (or decreasing projected angular separation from M87) show a significant decrease in the HI content and in the mean specific star formation rate, along with a progressive reddening of their stellar populations. The gradual quenching of the star formation occurs outside-in, consistently with the predictions of the ram pressure model. Once considered as a whole, the Virgo cluster is effective in removing neutral hydrogen from galaxies, and this perturbation is strong enough to appreciably reduce the SFR of its entire galaxy population. An estimate of the present infall rate of 300-400 galaxies per Gyr in the Virgo cluster is obtained from the number of existing HI-rich late-type systems, assuming 200-300 Myr as the time scale for HI ablation. If the infall process has been acting at constant rate this would imply that the Virgo cluster has formed approximately 2 Gyr ago, consistently with the idea that Virgo is in a young state of dynamical evolution.

CO in late-type galaxies within the central region of Abell 1367

We present CO (J = 1 - 0) and CO (J = 2 - 1) spectra for 19 bright, late-type galaxies (spirals) in the central region of the galaxy cluster Abell 1367 (z = 0.02) from observations made with the IRAM 30 - m telescope. All 19 spirals were observed at the position of their optical center and for a subset, at multiple positions. For each spiral the integrated CO (J = 1 - 0) intensity from the central pointing, in few cases supplemented with intensities from offset pointings, was used to estimate its molecular hydrogen mass and H_2 deficiency. Accepting the considerable uncertainties involved in determining H_2 deficiencies, spirals previously identified by us to have redder colours and higher HI deficiencies as a result of environmental influence, were found to be more H_2 deficient compared to members of the sample in less advanced evolutionary states. For eight of the observed spirals multiple pointing observations were made to investigate the distribution of their molecular gas. For these spirals we fitted Gaussians to the CO intensities projected in a line across the galaxy. In two cases, CGCG 097-079 and CGCG 097-102(N), the offset between the CO and optical intensity maxima was significantly larger than the pointing uncertainty and the FWHMs of the fits were significantly greater than those of the other spirals, irrespective of optical size. Both signatures are indicators of an abnormal molecular gas distribution. In the case of CGCG 097-079, which is considered an archetype for ram pressure stripping, our observations indicate the CO intensity maximum lies ~ 15.6 +/- 8.5 arcsec (6 kpc) NW of the optical centre at the same projected position as the HI intensity maximum.

Molecular gas content and SFR in Hickson compact groups: enhanced or deficient?

We study the effect of the extreme environment in Hickson Compact groups (HCGs) on the molecular gas mass, \mhtwo, and the star formation rate (SFR) of galaxies as a function of atomic hydrogen (HI) content and evolutionary phase of the group. We have selected a redshift limited (D$<$100 Mpc) sample of 88 galaxies in 20 HCGs with available atomic hydrogen (HI) VLA maps, covering a wide range of HI deficiencies and evolutionary phases of the groups, and containing at least one spiral galaxy. We derived the far-infrared (FIR) luminosity (\lfir) from IRAS data and used it as a tracer of the SFR. We calculated the HI mass, \lfir and \mhtwo deficiencies. The mean deficiencies of \lfir and \mhtwo of spiral galaxies in HCGs are close to 0, indicating that their average SFR and molecular gas content are similar to those of isolated galaxies. However, there are indications of an excess in \mhtwo\($\sim$ 50%) in spiral galaxies in HCGs which can be interpreted as either an enhanced molecular gas content or as a higher concentration of \mhtwo towards the center in comparison to galaxies in lower density environments. In contrast, the mean \mhi of spiral galaxies in HCGs is only 12% of the expected value. The specific star formation rate (sSFR= SFR/stellar mass) tends to be lower for galaxies with a higher \mhtwo\ or \mhi\ deficiency. This trend is not seen for the star formation efficiency (SFE=SFR/\mhtwo). We found tentative indications for an enhancement of \mhtwo\ in spiral galaxies in HCGs in an early evolutionary phase and a decrease in later phases. We suggest that this might be due to an enhancement of the conversion from atomic to molecular gas due to on-going tidal interactions in an early evolutionary phase, followed by HI stripping and a decrease of the molecular gas content because of lack of replenishment.

Atomic hydrogen deficiency in galaxies of the virgo and coma clusters: A new estimation method

We propose a new method for estimating the HI deficiency in galaxies. The method is based on a semi-empirical relationship between the total mass of HI and specific angular momentum of isolated galaxies. The atomic-hydrogen deficiency is estimated for nearby spiral galaxies and for spiral galaxies in the Virgo and Coma clusters. The mean HI deficiencies determined for these samples using our method are similar to those obtained with conventional methods, although there are considerable differences in some cases. The HI deficiency in nearby galaxies does not depend on their degree of isolation, and there is no systematic discrepancy between their HI and “normal” masses. Significant HI deficiencies are observed in the Virgo and Coma clusters, out to distances of 1.5 and 3–4 Mpc from the cluster centers, respectively. At such distances, the ram pressure is too small to sweep a considerable amount of gas from the galactic disks. Either these galaxies have passed through the dense cluster center, or their gas deficiency is due to the fact that the halo had stopped accreting onto the disk when the galaxy entered the cluster.

The effect of the environment on the H i scaling relations

We use a volume-, magnitude-limited sample of nearby galaxies to investigate the effect of the environment on the HI scaling relations. We confirm that the HI-to-stellar mass ratio anti correlates with stellar mass, stellar mass surface density and NUV-r colour across the whole range of parameters covered by our sample (10^9 <M*<10^11 Msol, 7.5 <mu*<9.5 Msol kpc^-2, 2<NUV-r<6 mag). These scaling relations are also followed by galaxies in the Virgo cluster, although they are significantly offset towards lower gas content. Interestingly, the difference between field and cluster galaxies gradually decreases moving towards massive, bulge-dominated systems. By comparing our data with the predictions of chemo-spectrophotometric models of galaxy evolution, we show that starvation alone cannot explain the low gas content of Virgo spirals and that only ram-pressure stripping is able to reproduce our findings. Finally, motivated by previous studies, we investigate the use of a plane obtained from the relations between the HI-to-stellar mass ratio, stellar mass surface density and NUV-r colour as a proxy for the HI deficiency parameter. We show that the distance from the `HI gas fraction plane' can be used as an alternative estimate for the HI deficiency, but only if carefully calibrated on pre-defined samples of `unperturbed' systems.

Quantifying the importance of ram-pressure stripping in a galaxy group at 100 Mpc

We examine two members of the NGC 4065 group of galaxies: a bent-double (a.k.a. wide angle tail) radio source and an HI deficient spiral galaxy. Models of the X-ray emitting intragroup gas and the bent-double radio source, NGC 4061, are used to probe the density of intergalactic gas in this group. HI observations reveal an asymmetric, truncated distribution of neutral gas in spiral galaxy, UGC 07049, and the accompanying radio continuum emission reveals strong star formation. We examine the effectiveness of ram pressure stripping as a gas removal mechanism and find that it alone cannot account for the HI deficiency that is observed in UGC 07049 unless this galaxy has passed through the core of the group with a velocity of ~800 km/s. A combination of tidal and ram pressure stripping are necessary to produce the HI deficiency and asymmetry in this galaxy.

STAR FORMATION IN PARTIALLY GAS-DEPLETED SPIRAL GALAXIES

Broadband B and R and H-alpha images have been obtained with the 4.1-m SOAR telescope atop Cerro Pachon, Chile for 29 spiral galaxies in the Pegasus I galaxy cluster and for 18 spirals in non-cluster environments. Pegasus I is a spiral-rich cluster with a low density intracluster medium and a low galaxy velocity dispersion. When combined with neutral hydrogen (HI) data obtained with the Arecibo 305-m radiotelescope, acquired by Levy et al. (2007) and by Springob et al. (2005a), we study the star formation rates in disk galaxies as a function of their HI deficiency. To quantify HI deficiency, we use the usual logarithmic deficiency parameter, DEF. The specific star formation rate (SSFR) is quantified by the logarithmic flux ratio of H-alpha flux to R band flux, and thus roughly characterizes the logarithmic SFR per unit stellar mass. We find a clear correlation between the global SFR per unit stellar mass and DEF, such that the SFR is lower in more HI-deficient galaxies. This correlation appears to extend from the most gas-rich to the most gas-poor galaxies. We also find a correlation between the central SFR per unit mass relative to the global values, in the sense that the more HI-deficient galaxies have a higher central SFR per unit mass relative to their global SFR values than do gas-rich galaxies. In fact, approximately half of the HI-depleted galaxies have highly elevated SSFRs in their central regions, indicative of a transient evolutionary state. In addition, we find a correlation between gas-depletion and the size of the H-alpha disk (relative to the R band disk); HI-poor galaxies have truncated disks. Moreover, aside from the elevated central SSFR in many gas-poor spirals, the SSFR is otherwise lower in the H-alpha disks of gas-poor galaxies than in gas-rich spirals.

Parkes H I observations of galaxies behind the southern Milky Way

As part of our programme to map the large-scale distribution of galaxies behind the southern Milky Way, we observed 314 optically-selected, partially-obscured galaxies in the Zone of Avoidance (ZOA) in the Crux and Great Attractor (GA) regions. The observations were conducted with the Parkes 64m radio telescope, in a single-pixel pointed mode, reaching an rms noise level of typically 2-6 mJy over the velocity search range of 400<v<10500km/s. A total of 162 galaxies were detected. This can be explained by the prominence of the GA overdensity in the survey regions, which leads to a relatively higher fraction of nearby galaxies. It is also evident from the quite narrow velocity distribution (largely confined to 3000-6000km/s) and deviates significantly from the expectation of a uniform galaxy distribution for the given sensitivity and velocity range. No systematic differences were found between detections and non-detections, in terms of latitude, foreground extinction, or environment, except for the very central part of the rich Norma cluster, where hardly any galaxies were detected. A detailed investigation of the HI content of the galaxies reveals strong HI deficiency at the core of the Norma cluster (within about a 0.4 Abell radius), similar to what has been found in the Coma cluster. The redshifts obtained by this observing technique result in a substantial reduction of the so-called redshift ZOA. This is obvious when analysing the large-scale structure of the new HI data in combination with data from optical ZOA redshift surveys. The lower latitude detections provide further evidence of the extension of the Norma Wall, across the ZOA, in particular its bending towards the Cen-Crux clusters above the Galactic plane at slightly higher redshift, rather than a straight continuation towards the Centaurus clusters.