Neanderthal Groups Research Articles

Between two worlds: Cova Eirós and the Middle-Upper Palaeolithic transition in NW Iberia

Iberia, a natural cul-de-sac peninsula, plays a major role in the study of the Neanderthals demise and its eventual relationship with the spread of Anatomically Modern Humans (AMH) in Europe. The site of Cova Eirós (Galicia, Spain), located in NW Iberia, contains Middle and Upper Palaeolithic levels, based on the cultural remains recovered at the site. No human remains directly associated with those levels were discovered yet. The available radiocarbon dates from the levels 2 (c. 35 ka cal BP, Early Upper Paleolithic) and 3 (c. 41 ka cal BP, Late Middle Paleolithic), point to a late survival of Neanderthal groups in North Iberia and to a relative quick arrival of the AMH, c. 35-36 ka cal BP, with respect to other territories of the Iberian Peninsula. The archaeological record shows clear differences between the Middle and the Upper Palaeolithic occupations, regarding raw-material acquisition, lithic technology and subsistence strategies. The location of Cova Eirós in the westernmost margin of the Cantabrian Rim and in the Atlantic Façade, makes this site a key place to understand the spread of the first AMH and the progressive demise of Neanderthal populations.

A neanderthal hunting camp in the central system of the Iberian Peninsula: A zooarchaeological and taphonomic analysis of the Navalmaíllo Rock Shelter (Pinilla del Valle, Spain)

The interior of the Iberian Peninsula has few Middle Palaeolithic sites, especially when compared to other areas of the Mediterranean Basin and the northern Spanish region. Few in number too are the zooarchaeological and taphonomic studies that throw light on the relationships between Neanderthal groups, their environment, and the use they made of it. The present work examines, both zooarchaeologically and taphonomically, the faunal remains of levels F and D of the Navalmaíllo Rock Shelter (Pinilla del Valle, Madrid, Spain) - the largest collection of such remains ever studied from the Iberian interior. The results allow this site to be interpreted as a Neanderthal hunting camp where occupations were short-term. Neanderthal people were the main agents that accumulated the site's faunal remains - largely those of large bovids and to a lesser extent medium-sized cervids. The activity of carnivores was also identified, but these animals mostly left behind the remains of small prey or fed upon carcasses abandoned at the camp by human hunters.

A road to nowhere? The non-transitional sequence at El Esquilleu (Cantabria, Spain)

El Esquilleu cave has one of the most complete Middle Paleolithic stratigraphies of northern Iberia with a complete chronological framework almost continuous from the beginning of MIS3. The complete analysis of the materials including the last section of the sequence corresponding to the last chronological interval of the occupation in the region shows clear evolutionary tendencies by important changes in the composition of the lithic assemblage. Its study confirms a continuous occupation without any gaps during the Mousterian until recent dates but without any Upper Paleolithic presence. This sequence, referenced to the Mousterian settlement of the Picos de Europa mountains, testifies to the existence of a Mousterian presence in northern Iberia with no clear relation to other Cantabrian areas. Consequently, its occupation without a break until the crucial phase of the end of the Middle Paleolithic suggests a non-transitional process in the history of local Neanderthal groups.

Magnetic Susceptibility Properties of Loess From the Willendorf Archaeological Site: Implications for the Syn/Post-Depositional Interpretation of Magnetic Fabric

At the Willendorf site Upper Paleolithic archeological layers associated to early Aurignacian cultures were found. The environmental conditions of the associated society, potentially co-existing with Neanderthal groups, is still not fully understood. Here, we report on environmental magnetic analyses including anisotropy of the magnetic susceptibility (AMS) carried out on loessic aeolian sediments at the Willendorf site. Data on lineation, foliation and the degree of anisotropy were used to assess depositional and post-depositional magnetic fabric properties and to deduce site-specific environmental processes. Overall, the loess is of aeolian origin and shows magnetic enhancement and magnetic fabric properties similar to those of other European loess geoarchives, but the magnetic mineralogy differs from many ‘dry’ loess sites, insofar as it shows a higher susceptibility during heating. We infer an enhanced neoformation of magnetite during heating due to the presence of organic matter. While at face value the AMS properties are indicative of pure aeolian loess consistent with previous studies, imbrication suggests post-depositional slope movement toward the Danube, which obscures inferences on palaeo-wind direction. It is well possible that these post-depositional magnetic fabric alterations occur at other localities with similar geomorphological settings.

Erratum to “Palaeoecological context for the extinction of the Neanderthals: A small mammal study of Stratigraphic Unit V of the El Salt site, Alcoi, eastern Spain” [Palaeogeography, Palaeoclimatology, Palaeoecology 530 (2019) 163–175

Abstract El Salt is an important reference site for understanding the extinction of Neanderthal populations in the eastern Iberian Peninsula during MIS 3. In this paper, we describe the small mammal assemblage from Stratigraphic Unit V, the youngest unit with evidence of human presence, based on nearly 1300 specimens. A total of seven rodents (Microtus arvalis, Microtus duodecimcostatus, Microtus cabrerae, Sciurus vulgaris, Arvicola sapidus, Eliomys quercinus and Apodemus sylvaticus), three insectivores (Talpa occidentalis, Crocidura sp., Sorex sp.) and one lagomorph (Oryctolagus cf. cuniculus) were identified. Palaeocological analyses point to drier conditions in this part of the stratigraphic sequence, supporting the hypothesis that an aridification scenario may have played a role in the extinction of the Neanderthal groups inhabiting this region of the Iberian Peninsula.

A new look at the Middle Paleolithic lithic industry of the Teshik-Tash Cave, Uzbekistan, West Central Asia

Teshik-Tash Cave, excavated in 1938 and 1939, is a key archaeological site for our understanding of the Middle Paleolithic population dynamics of Central Asia. Its privileged status is due to the discovery of well-preserved Neanderthal skeletal remains in association with rich lithic artifacts. However, despite repeated studies of the skeletal remains, the lithic industry has not received sufficient re-analysis with up-to-date information since its first publication more than a half century ago. The present paper reports our first results based on the collection stored at the State Museum of History of Uzbekistan, Tashkent. We show that a combination of hierarchical and nonhierarchical core reduction strategies, the former of which includes a particular type of Levallois technology, characterizes the Teshik-Tash industry. Typologically, the common manufacturing of side scrapers, associated with a small number of heavy-duty tools, are diagnostic. It is also notable that triangular points often common in the Middle Paleolithic of Southwest Asia are rare. This industry is regarded as an important Neanderthal cultural tradition of West Central Asia, whose precise spatio-temporal placement needs to be determined through comparisons with other assemblages from controlled excavations. In the meantime, its disparity from the Sibiryachikha Mousterian industry left by the Neanderthals of East Central Asia is incontestable. This finding helps us understand the complicated origin and development of Neanderthal groups in Central Asia.

Neanderthal plant use and stone tool function investigated through non-pollen palynomorphs analyses and pollen washes in the Abri du Maras, South-East France

This pioneering research performed at the Abri du Maras (Ardèche, South-East France) explores the potential of non-pollen palynomorphs (NPP) to provide new insights into the relationships between Neanderthals and plant resources. Remains of endophytic fungi found during the analysis are key indicators of the local presence of plants, which could have potentially been used by Neanderthal groups. The recording of high concentrations of Chaetomium-type (HdV-7A) in archaeological contexts from localized areas within the shelter suggests the local presence of decayed plants near an in situ fireplace. This could point to the use of grasses or other types of plants from aquatic or marshy areas such as reed and rushes for a surface preparation (bedding?) near the fireplace. Our research has also applied for the first time a combined study of archaeological sediments and pollen washes of artefacts following a microspatial approach in the study of Middle Palaeolithic archaeological contexts. One artefact (1079), identified as a sidescrapper, provided positive results, with the recording of high concentrations of Diporotheca webbiae-type (HdV-143). We could identify the host plant of the fungal signature documented on the tool surface as alder roots by comparing archaeobotanical results with NPP analyses in modern surface samples. Following this, we have proposed the use of this Middle Palaeolithic sidescrapper for processing this plant by Neanderthal groups. Marsh and riparian ecosystems were thus attractive for Neanderthals occupying the Abri du Maras. This study clearly shows that NPP must be considered as key taphonomical, environmental and cultural indicators in archaeobotanical analyses. It also underlines the contribution of combined studies of archaeological sediments and pollen washes of artefacts following a microspatial approach to a better understanding of Middle Palaeolithic lithic tools and plant use relationships.

High-resolution Neanderthal settlements in mediterranean Iberian Peninsula: A matter of altitude?

Neanderthals are widely known to be a resilient human species that successfully faced constant and strong environmental fluctuations modifying the landscapes they inhabited and the availability of their potential resources. It has been traditionally assumed that environmental features could strongly affect human behaviour due to the stretch relationship between their potential prey and the availability of habitats. Environmental changes would produce a high variability in the settlement patterns of the Neanderthal groups. However, the preponderant influence of environmental conditions on these human groups has recently been strongly questioned. This is especially true for the Iberian Peninsula, where latitudinal position and orographic features supported the persistence of environmental conditions that allowed the permanence of a wide range of biotopes and animal and vegetal resources. In the present study, we reconstruct the Neanderthal settlement patterns (i.e. duration and season) from four archaeological sites located in the Mediterranean area of the Iberian Peninsula: Arbreda Cave (Serinyà, Girona), Teixoneres Cave (Moià, Barcelona), Abric Romaní (Capellades, Barcelona), and El Salt (Alcoy, Alicante). We focus on identifying whether they show a high variability in settlement patterns and check for the existence of local influences, such as the altitude. To reach these objectives, we designed a multi-proxy approach combining tooth wear and dental cementum analysis of the main Neanderthal preys: Cervus elaphus, Equus ferus, Equus hydruntinus, Bos primigenius and Capra pyrenaica. Our results first suggest that local features (i.e. orography, altitude, and environment) appear to be predominant factors determining the main feeding behaviour of the ungulates hunted by Neanderthals. Additionally, seasonal environmental variations seem to have influenced horse migratory behaviour, involving altitudinal movements in search of high-nutritive pastures. Neanderthal survival strategies and settlement patterns seem to have been less affected by environmental particularities and more linked to selective prey procurement based on the seasonal resource availability. Therefore, the duration and seasonality of their settlement patterns and their mobility throughout the landscape indicate they had a high level of knowledge on the territory they inhabited, as well as on the behaviour and availability of their potential preys.

Atapuerca Neanderthal landscape at Fuente Mudarra site in Burgos, Spain, during Marine Isotope Stages 5–3

AbstractFuente Mudarra is on a gentle slope on the left bank of the Pico River, near Sierra de Atapuerca, Burgos, Spain. A 12−m2test pit was dug at this large open-air site between 2012 and 2017. Several upper Pleistocene archaeological levels were documented. Results from Fuente Mudarra confirm that Neanderthal groups, little represented at cave sites, occupied Sierra de Atapuerca from the end of the middle Pleistocene and during the upper Pleistocene. The site also provides insight into Neanderthal spatial organization in the Atapuerca area and whether they used the caves in an occasional, non-habitual way like the open-air sites.

Use and selection of bone fragments in the north of the Iberian Peninsula during the Middle Palaeolithic: bone retouchers from level 4 of Prado Vargas (Burgos, Spain)

Bone specimens showing a large amount of markings that include scratching, cutting, and fracture provide us with a clear idea of the use of animals from an archaeological site for butchery-related work. Nevertheless, there is one type of damage that stands out from the rest—the stigmas associated with retouch work. There is a great deal of evidence concerning the use of bones by Neanderthal groups as retouchers. In particular, we present a sample of 65 retouchers from the level 4 of Prado Vargas (Burgos, Spain). This level has yielded an important assemblage of retouchers; however, very few retouched tools have been retrieved, a fact that has led us to consider their peculiarity. Through this investigation, we seek to explore the question as to how the inhabitants of Prado Vargas used bone retouchers during the Middle Palaeolithic.

Techno-functional and 3D shape analysis applied for investigating the variability of backed tools in the Late Middle Paleolithic of Central Europe.

In the Late Middle Paleolithic of Central Europe, two main cultural complexes have been distinguished: the Micoquian or Keilmessergruppe (KMG), and the Mousterian. Their differences mainly consist in the frequence of some retouched tools and the presence of bifacial technology. When these industries coexist, one element of discussion is the application of different concepts to manufacture tools with the same techno-functionality. This is particularly true for backed artifacts, such as Keilmesser (backed, asymmetrical bifacially-shaped knives) opposed to flake-tools equipped with a natural or knapped back. We conducted a techno-functional analysis of the backed tools from the G-Layer-Complex of Sesselfelsgrotte, one of the main Late Middle Paleolithic sequences in Central Europe, characterized by a combination of KMG and Mousterian aspects. In order to better understand the morpho-metrical data, 3D scans were used for recording technical features and performing semi-automatic geometric morphometrics. Results indicate that the techno-functional schemes of Keilmesser show a moderate variability and often overlap with the schemes of other typological groups. Within bifacial backed knives, a process of imitation of unifacial flake tools' functionaly was recognized particularly in the cutting edge manufacturing. Keilmesser proved to be the long-life, versatile version of backed flake-tools, also due to the recurrent valence as both tool and core. This is why Keilmesser represent an ideal strategic blank when a mobile and multi-functional tool is needed. Based on these data, it is assumed that the relationship between Mousterian and KMG is deeply rooted and the emergence of KMG aspects could be related to constrained situations characterizing the long cold stages of the Early Weichselian. A higher regional mobility caused by the comparably low predictability of resources characterized the subsistence tactics of Neanderthal groups especially at the borders of their overall distribution. For this reason, Keilmesser could have represented an ecological answer before possibly becoming a marker of cultural identity.

Neanderthal faunal exploitation and settlement dynamics at the Abri du Maras, level 5 (south-eastern France)

Over the past two decades, taphonomic and zooarchaeological studies have focused on Neanderthal settlement patterns and subsistence strategies. The south-eastern margins of the Massif Central constitute one of the regions with the most abundant archaeological evidence of Neanderthal occupations in France. The faunal record of level 5 of Abri du Maras is a unique source of information for analysing Neanderthal behaviour at the end of the MIS 5. The assemblage is divided into three levels 5.1, 5.2 and 5.3, which correspond to the three main phases of human occupation of the shelter in level 5. Through the taphonomic and zooarchaeological analysis of the faunal remains, we define the characteristics of human occupation developed in the rock shelter. In this paper, we show that Neanderthal groups used the Abri du Maras as a residential campsite for long-term occupation events during a whole season, mainly summer. The intensive exploitation of the faunal resources is typical of a residential campsite, and the selective seasonal hunting strategy of medium and large ungulates, centred on prime adults, indicates different Neanderthal subsistence patterns to the upper levels of the sequence. Here, our results document the chronological changes in occupation patterns in one of the reference sites in south-eastern France and address questions related to cultural choices and environmental constraints.

Petrographic and SEM‐EDX characterization of Mousterian white/beige chert tools from the Navalmaíllo rock shelter (Madrid, Spain)

AbstractStudying lithic raw material sourcing, processing and distribution is helpful when trying to reconstruct the territory, ecology, and cultural practices of Neanderthal groups. The use of multiple methods in such analyses allows for more refined characterizations to be made, helping to distinguish between materials better than any single method. Although 85% of the raw materials making up the Mousterian assemblage at the Navalmaíllo Rock Shelter (Pinilla del Valle, Madrid Region of Spain) correspond to the available local geological resources, 10% is made of white/beige chert, which is not registered in regional geological cartography. Petrographic and scanning electron microscopy with energy dispersive X‐ray spectroscopy analyses were performed to determine the origin of this white/beige chert and thus shed light on the procurement strategies of the Neanderthal groups that occupied the rock shelter. The results show this chert to correspond to three different types of rock: quartz–kaolinite rocks from dykes, cretaceous marine cherts, and quartz fillings of cavities/fractures. These findings are in accordance with the geological features and formations present in the Lozoya valley, as recorded during a geoarchaeological survey, and indicate that the Neanderthals occupying the center of the Iberian Peninsula possessed a detailed knowledge of the landscape that allowed them to exploit its resources during MIS5a–early MIS4.

Neanderthals in a highly diverse, mediterranean-Eurosiberian forest ecotone: The pleistocene pollen record of Teixoneres Cave, northeastern Spain

A palynological study of the archaeological layers from the Neanderthal site of Teixoneres Cave, located in Northeastern Spain, is presented. Vegetation dynamics for the MIS 7-MIS 2 period are described, revealing the long-term resilience of mixed oak-pine forests throughout cold phases and a high diversity of woody taxa, including conifers, mesophytes, Mediterranean, and xerothermics. Unexpected features of the Teixoneres sequence include the relative abundances of evergreen Quercus, deciduous Quercus + suber and Juniperus, the continuous occurrences of Corylus, Castanea, Betula, Fraxinus, Buxus, Olea, Populus, and Salix, and the presence of Abies, Taxus, Cedrus, Acer, Alnus, Celtis, Juglans, Fagus, Ulmus, Calicotome, Ceratonia, Cistus, Ephedra fragilis, Myrtus, Pistacia, Phillyrea, Rhamnus and Viburnum. To our knowledge, the existence of a forest refugium during the coldest and most arid stages of the upper Pleistocene has not previously been documented across this region. The high values of oak pollen during depositional episodes III, II and I of Teixoneres do not have an equivalent in any pollen sequence of the northern Iberian Peninsula during the MIS 4-MIS 2 interval. Our palynological analyses reveals a specific forest environment within the Mediterranean-Eurosiberian ecotone for the human occupation of Teixoneres Cave. These forested environments would have undoubtedly required highly adaptive subsistence strategies and expertise to ensure the long-term survival of Neanderthal groups in the region.

Environmental conditions at the Last Interglacial (Eemian) site Neumark‐Nord 2, Germany inferred from stable isotope analysis of freshwater mollusc opercula

Mollusc biogenic carbonates are valuable records of past environmental conditions. In particular, carbonate oxygen (δ18O) and carbon (δ13C) stable isotopes can be used to reconstruct different physical and chemical parameters, according to the different genera used (marine, freshwater or terrestrial). The Last Interglacial (early Eemian) palaeolake of Neumark‐Nord 2 (NN2), Germany provides an excellent example of a Neanderthal archaeological site with abundant freshwater carbonate remains. As in other European contexts, one of the most abundant species is Bithynia tentaculata. In order to provide a robust regional baseline for the interpretation of the archaeological data, this study includes a calibration phase on modern B. tentaculata opercula. The results indicate that these calcitic structures are likely to be subjected to a growth slowdown/cessation during summer, which influences their geochemistry, reflecting mainly the water properties of the rest of the year. This modern calibration, together with the existing palaeoenvironmental reconstructions developed for NN2 (e.g. pollen data), represents a valuable opportunity to establish B. tentaculata opercula as reliable environmental proxies applicable to several other freshwater contexts. The isotope data of the NN2 opercula, in agreement with the pollen record, indicate that the major archaeological horizon was formed during a rather wet period and potentially in a semi‐forested environment. However, human occupation occurred also during drier phases at the site and within a wide temperature range, indicating the absence of restricted environmental preferences by the local Neanderthal groups.

Spatial analysis of the late Middle Palaeolithic open-air site of Bout-des-Vergnes (Bergerac, Dordogne) based on lithic technology and refitting

The late Mousterian open-air site of Bout-des-Verges was the subject of large-scale rescue excavations between 2012 and 2103 that yielded evidence for the spatial organisation of particular techno-typological elements amongst the 900 lithics recovered from the site. Three knapping concentrations in the southwest sector of the site produced two-thirds of the overall lithic assemblage, the remaining pieces were found scattered across the excavated area. Our techno-economic analysis of the Bout-des-Verges assemblage revealed clear differences in terms of raw material economy and artefact transport patterns between the dominant discoidal component and the larger, imported bifaces and Levallois flakes. Refitted reduction sequences and our spatial analysis confirm the three lithic concentrations to be contemporaneous and, by extension, to form part of a single Mousterian occupation with multiple technological components. The majority of discoidal products produced on the site were exported for future use, while the imported bifaces and Levallois scrapers with longer use-lives were transported multiple times by Neanderthal groups.

Krapina atlases suggest a high prevalence of anatomical variations in the first cervical vertebra of Neanderthals.

The first cervical vertebra, atlas, and its anatomical variants have been widely studied in Homo sapiens. However, in Neanderthals, the presence of anatomical variants of the atlas has been very little studied until very recently. Only the Neanderthal group from the El Sidrón site (Spain) has been analysed with regard to the anatomical variants of the atlas. A high prevalence of anatomical variants has been described in this sample, which points to low genetic diversity in this Neanderthal group. Even so, the high prevalence of anatomical variations detected in El Sidrón Neanderthal atlases needs to be confirmed by analysing more Neanderthal remains. In this context, we analysed the possible presence of anatomical variants in the three Neanderthal atlases recovered from the Krapina site (Croatia) within the Neanderthal lineage. Two of the three Krapina atlases presented anatomical variations. One atlas (Krapina 98) had an unclosed transverse foramen and the other (Krapina 99) presented a non-fused anterior atlas arch. Moreover, an extended review of the bibliography also showed these anatomical variations in other Middle and Upper Pleistocene hominins, leading us to hypothesise that anatomical variations of the atlas had a higher prevalence in extinct hominins than in modern humans.

Nouveau site du Paléolithique Moyen dans le Caucase Central (résultats préliminaires des études complexes)

The newest studies of the Caucasus Paleolithic indicate movements of human groups between the northern and southern, and western and eastern parts of this region. In the Baksan river valley in the north-central Caucasus, there is known a large source of obsidian, which was transported to sites in the north-western Caucasus already in the Middle Paleolithic. However, no stratified Middle Paleolithic site was known in the source area until 2016 when we discovered here the first cave site of Saradj-Chuko cave. The article presents the first results of a comprehensive research of this site. The Saradj-Chuko cave is located just 6–7km from the important obsidian source that represents the only source of obsidian, which was highly valued in the past, in the Northern Caucasus, and about 70km from the Elbrus volcanic peak. A comprehensive study in Saradj-Chuko cave began in 2017. A small (∼8 sq. m) excavation was dug next to the 2016 test pit. We report a detailed description of the materials excavated from the main Middle Paleolithic occupational Layer 6B. The study of the lithic assemblage allows us to make a preliminary conclusion that Layer 6B represents an active habitation level where Neanderthals were engaged in intensive flaking of obsidian and usage of tools made predominantly from obsidian for butchering and consumption of hunting game. The laminar character of the stone industry distinguishes the Mousterian industry of the north-central Caucasus and probably the entire Terek river basin from the Eastern Micoquian industry that is represented in factually all Middle Paleolithic sites in the north-western Caucasus (Kuban river basin), as well as makes the Mousterian of north-central Caucasus similar to the Mousterian industries known in the north-eastern Caucasus and the Southern Caucasus. We suggest cultural connections and contacts among Neanderthal groups from these regions during the Middle Paleolithic period.

The study of the lower limb entheses in the Neanderthal sample from El Sidrón (Asturias, Spain): How much musculoskeletal variability did Neanderthals accumulate?

Entheses have rarely been systematically studied in the field of human evolution. However, the investigation of their morphological variability (e.g., robusticity) could provide new insight into their evolutionary significance in the European Neanderthal populations. The aim of this work is to study the entheses and joint features of the lower limbs of El Sidrón Neanderthals (Spain; 49 ka), using standardized scoring methods developed on modern samples. Paleobiology, growth, and development of both juveniles and adults from El Sidrón are studied and compared with those of Krapina Neanderthals (Croatia, 130 ka) and extant humans. The morphological patterns of the gluteus maximus and vastus intermedius entheses in El Sidrón, Krapina, and modern humans differ from one another. Both Neanderthal groups show a definite enthesis design for the gluteus maximus, with little intrapopulation variability with respect to modern humans, who are characterized by a wider range of morphological variability. The gluteus maximus enthesis in the El Sidrón sample shows the osseous features of fibrous entheses, as in modern humans, whereas the Krapina sample shows the aspects of fibrocartilaginous ones. The morphology and anatomical pattern of this enthesis has already been established during growth in all three human groups. One of two and three of five adult femurs from El Sidrón and from Krapina, respectively, show the imprint of the vastus intermedius, which is absent among juveniles from those Neanderthal samples and in modern samples. The scant intrapopulation and the high interpopulation variability in the two Neanderthal samples is likely due to a long-term history of small, isolated populations with high levels of inbreeding, who also lived in different ecological conditions. The comparison of different anatomical entheseal patterns (fibrous vs. fibrocartilaginous) in the Neanderthals and modern humans provides additional elements in the discussion of their functional and genetic origin.

Last Neanderthal occupations at Central Iberia: the lithic industry of Jarama VI rock shelter (Valdesotos, Guadalajara, Spain)

The work undertaken at the Jarama VI site (Valdesotos, Guadalajara, Spain) in the 1990s resulted in the recovery of thousands of archeological remains from the three Pleistocene sedimentary units of this cavity. Prior to the systematic analysis of the lithic material and the reception of new geochronological data, it had been suggested that the upper unit of Jarama VI could correspond to the Early Upper Paleolithic, while the other two units could be related to Neanderthal occupations. We now present the results of a comprehensive analysis and review of the materials, enabling us to clarify that all three units at Jarama VI belong to the Middle Paleolithic and, hence, to the last known Neanderthals in this central area of the Iberian Peninsula, a relatively high altitude inland zone within the framework of mountainous landscapes—parameters which seem to be characteristic of these last groups of Neanderthals.