The fauna of sewage filter beds is dominated by certain species of flies and worms, which play an important part in sewage purification processes as they feed on the bacterial, algal and fungal matter which develops in the bed. This growth, if unchecked, would soon block the filter and make it useless for purification. Experience has shown that a filter with large grade medium supplied with a moderate amount of average town sewage builds up a fauna with several successful forms but, under other conditions, the fauna tends to be limited to very few species. It has been suggested that the strength of the sewage is the chief cause of limitation (Lloyd 1945), whilst the method of delivery, the medium used and the way in which the bed is constructed may all have some measure of responsibility (Lloyd, Graham & Reynoldson 1940). On the other hand, it has been held that the loading of the filter is the overriding factor (Tomlinson & Stride 1945). The present studies were made at the Bradford Corporation's works at Esholt and North Bierley. The beds here provide a variety of conditions, both small and large medium with strong sewage and small medium with weak sewage. They were chosen as contrasting with the Knostrop works at Leeds where the medium is large and the sewage relatively weak. The fauna of Knostrop has already been described in detail (Lloyd 1935, 1937). The sewage passing to the Esholt works contains a large proportion of industrial wastes and is characterized by its high biochemical oxygen demand and its low pH. The beds are rectangular in shape, flush with the ground and 6 ft in depth. They are fed from travelling distributors at a rate of 75 gal per sq. yd per day. Hard coal of 3 in. grade is employed as the filtering medium in most of the beds but, in recent years, five acres of this has been replaced by sandstone and river pebbles of 2-21 in. grade. The fungus Fusarium aqueductum (Ascomycetes) is found on both types of medium and is evidently suited to the acid conditions (Butcher 1932). The growth is especially heavy during the winter months when there is a danger of its blocking the filters. There is also a growth of the alga Phormidium (Cyanophyceae) on the pebble beds but not on the coal. In late autumn and winter, this forms a thick leathery growth but
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