Forest Reserve Area Research Articles

Species Evenness and Diversity of Soil Invertebrates at Different Agricultural Lands and a Forest Reserve at Kota Belud, Sabah

Increasing agricultural activities can lead to changes in soil ecosystems, potentially impacting soil invertebrate communities as they are highly responsive to soil disturbances. The composition of soil invertebrates within each agricultural habitat provides insights into how these communities respond to their environmental conditions. This study aimed to investigate the evenness and diversity of soil invertebrates in agricultural areas of Kota Belud, Sabah. Soil samples were collected from three different agricultural sites, a rubber plantation, an oil palm plantation, and a paddy field, as well as a forest reserve as a control site. Pitfall traps and Berlese-Tullgren funnels were used to collect soil invertebrates. A total of 180 soil samples and 474 individuals of soil invertebrates were collected and analyzed. Physicochemical analyses, including moisture content, soil texture, pH, soil nutrient content, and organic matter, were conducted on all soil samples to assess their influence on the composition of soil invertebrates. Results indicated that Julida, Coleoptera, Gastropoda and Araneae were frequently associated with high organic matter, pH, moisture content and phosphorus. On the other hand, Haplotaxida appeared to be more sensitive to potassium levels. Notably, the Blattodea, Orthoptera, Hymenoptera, Isopoda, Polydesmida and Diptera, were found to be abundant in areas with lower organic matter and pH in the control site, i.e., forest reserve area. These findings underscore the significance of soil layer activities in influencing the presence and survival of soil invertebrates. Given their vital role in sustaining life, prioritizing the enhancement of soil invertebrate populations is crucial, particularly in Sabah, a prominent food crop-producing state in Malaysia.

Farmer-Agropastoralist Conflict Escalation in Morogoro Region, Tanzania: A Structural and Process Analysis

In Africa, access to natural resources such as land, water, forest, and wildlife reserve areas is a key factor for socio-economic development. The scarcity and degradation of these resources in rural environments threaten human security, leading to conflicts. Conflicts between agropastoralists and farmers are increasing in Tanzania. This research deployed the Social Conflict Theory that embeds structural and process elements to address the question of why and how the conflict between farmers and agropastoralists has escalated into deadly violence in the Morogoro region, Tanzania. Qualitative and ethnographic research methods were used. Results demonstrate that the way farmers lose their land by illegal means (e.g., bribes, force and coercion, and deception and stealth) is a critical factor that contributes more to land resource conflicts. We argue that land resource-related conflicts in Tanzania cannot be explained solely by a single driver (e.g., limited land resources), but by a combination of drivers including the illegal mechanisms used to control access to land resources. Conflicts between farmers and agropastoralists in the Morogoro region escalated to violence following the general patterns and transformation dynamics (i.e., process variables) described in Social Conflict Theory. These process variables involve tactics shifting from light to heavy, goals shifting from specific to general, and involvement shifting from few to many. The analysis of conflicts by focusing on process variables (conflict dynamics) enabled the identification of new factors (culture and age of participant in the conflict) that helped explain why some conflicts between farmers and agropastoralists escalate to deadly violence. The study recommends the formation of a loose coalition (e.g., Elders’ Tribunal) which includes equal representation of members from the farmer and agropastoralist communities which may help solve the current conundrum caused by top-down administrative procedures and practices, which often leads to outcomes that are ineffective and unsatisfactory to all parties

Moth Diversity in the Mahamaya Reserve Forest, Kokrajhar, Assam

The present study was conducted in the Mahamaya Reserve Forest area of Kokrajhar, Assam for one year from 2021 to 2022 to record the moths found in the area. A total of 129 species were identified, belonging to 20 families and 114 genera. The most predominant are the Erebidae and Crambidae, with 13 species being identified as new records from this region of Assam. The diversity indices revealed moderate dominance (D 0.19) of moths in the study sites. Simpson diversity with 0.8, Shannon diversity presented 2.1 moderate diversity. Margalef index value with 4.012 at par and Berger-Parker index 0.3 indicated a good diversity of moths in the region.

The Relationship of Community Livelihoods in and Around Forest Reserve Areas to Forest Resources in Humbang Hasundutan District, North Sumatra Province

The Relationship of Community Livelihoods in and Around Forest Reserve Areas to Forest Resources in Humbang Hasundutan District, North Sumatra Province

Inventory study of ethno-medicinal herbs, shrubs and climbers from Terakanahalli reserve forest area of Sirsi taluka, Karnataka, India

Enlisting of plant species from the different regions is a continuous and important process in understanding the changes occurring in floral composition over a period of time. Present study enumerated a total of 60 medicinal species of herbs, shrubs and climbers belonging to 34 families from Teraknahalli Reserve Forest area. The plants belong to diverse families with Apocynaceae, Fabaceae, Phyllanthaceae, Rubiaceae, and Solanceae were top five families contributing a sum of 23 species. This shows the richness in species diversity of the present study area. The documented plants have diverse medicinal usage. Total 25 medicinal species were regarded under different Rare Endemic Threatened categories by FRLHT, Bengaluru and IUCN. The study area is under great pressure because of human interference. The reserve forest needs proper management and conservation practices to improve the floral and faunal diversity of the area.

Conservation or deforestation: Households attitudinal analysis of forest resources for sustainable development of the Tain (II) Forest Reserve, Ghana

Forest conservation has recently become a global necessity, capturing considerable attention from scholars and policymakers. However, rampant deforestation remains a significant catalyst for the recent exacerbated climatic repercussions in forest regions. This study examined the perspectives regarding forest conservation and assessed their relationship with household income conditions in Ghana. Primary data were gleaned from two hundred fifty households within five communities around Ghana's Tain (II) forest reserve using a participatory rural assessment approach and interviews with structured questionnaires. The survey considered (i) socio-demographic drivers, (ii) basic family information, and (iii) forest conservation characteristics as hypothetical incentives to promote forest conservation and poverty alleviation. The findings revealed varied perspectives on conservational attitudes in the forest fringe communities. Marital status, education, employment, home size, and other factors influenced opinions in forest conversations. The study suggested the need for tailored ecological compensation and agroforestry practices to reduce deforestation in the Tain (II) Forest reserve area.

Distribution and ranging area of elephant in Simpan Panti Forest, Kota Tinggi, Johor, Malaysia

Abstract. Hassan NHN, Abdullah NM, Norazlimi NA, Saaban S, Yazi MFA. 2023. Distribution and ranging area of elephant in Simpan Panti Forest, Kota Tinggi, Johor, Malaysia. Biodiversitas 24: 5731-5738. Home range estimation is one of the most used animal ecology studies using a Geographic Information System (GIS). One adult female Asian elephant (Elephas maximus Linnaeus, 1758) was fitted with a telemetry satellite collar African Wildlife Tracking (AWT) to determine its home range size distribution through its ranging behaviour and to identify the relationship between the GPS collar location record and the area covered. The home range estimation was determined using GPS collar location data from a satellite telemetry system and analyzed by using the Minimum Convex Polygon methods. GPS collar location data were obtained from the Department of Wildlife and National Parks (DWNP). The movement of elephant data was overlayed with the land use data of the Simpan Panti Forest (Hutan Simpan Panti), Kota Tinggi District, Johor, Malaysia before being exported to ArcGIS Pro software. The ArcGIS Pro software can estimate this Asian elephant home range sizes by using the MCP method for twelve months, from 1 June 2019 until 31 May 2020. The total home range size is 447.17 km². Home range size is estimated to be 31.77 to 303.86 km² in one year. The size of this home range has presumably been impacted by extensive human activities such as highways and settlement areas. So, good and strong management is needed to control this conflict for humans to live in harmony in one area and protect the remaining forest reserve area from expanding to agricultural activity and human development areas.

A preliminary assessment of faunal mortality on a road through Mannur Reserve Forest, Kanchipuram, Tamil Nadu, India

Many studies have shown that anthropological acts driven by human demands have led to the extinction of numerous plant and animal species, or have put them in danger of becoming extinct. Deforestation is a serious occurrence that damages the ecosystem permanently, reduces biodiversity, and interferes with an entire ecological niche's capacity to function. Evidence on the causes and impacts of vehicle mobility in a forest reserve area is provided by the present research. Animal mortality was shown to be higher during the lockdown with traffic relaxation and somewhat lower during the lockdown without relaxation. The present study shows how unregulated automobile access into the Mannur Protected Forest results in significant faunal fatalities. The state has to be persuaded to exclusively adopt Good Environmental Practices (GEP) while constructing roads through the forest, and the appropriate authorities must keep an eye on the movement of automobiles through the remote forest zone.

The potential of Avicennia alba blume in phytoremediation of heavy metal Pb and Cu in the area east coast mangrove forest Reserve Jambi, Indonesia

The Batanghari River is the fourth longest river in Indonesia, and along the course of the river, there are many community activities, such as fishing, coal-carrying, and industrial activities. These various activities can cause contamination of heavy metals lead (Pb) and copper (Cu). This pollution can be overcome by using phytoremediation techniques using Avicennia alba Blume, which grows in the East Coast Mangrove Forest Reserve Area (CAHBPT), Jambi. This study aims to determine the potential and mechanism of phytoremediation of heavy metals Pb and Cu by A. alba at CAHBPT, Tanjung Solok, Tanjung Jabung Timur, Jambi. Sampling was done by a purposive sampling method. This study showed that the accumulation level of Pb metal in sediments was 10.08-13.42 mg/kg, and Cu metal was 8.61-22.92 mg/kg. The potential of A. alba in reducing the impact of Pb and Cu metal pollution is classified as an excluder marked BCF <1, even though the concentration of heavy metals in sediments is still below the EPA quality standard threshold for sediments. A. alba that grows in CAHBPT, Jambi, has a Pb metal translocation mechanism in the form of phytoextraction which is characterized by a TF value >1, and for Cu metal with a phytostabilization mechanism which is characterized by a TF <1.

An Extended Unit Restriction Model with Environmental Considerations for Forest Harvesting

This paper addresses a forest harvesting problem with adjacency constraints, including additional environmental constraints to protect wildlife habitats and minimize infrastructure deployment costs. To this end, we propose an integer programming model to include those considerations during the optimization of the harvest regime of a Mexican forest. The model considered was based on the Unit Restriction Model, a benchmark approach that merges the management units before the optimization process. The resulting model, namely the Green Unit Restriction Model (GURM) and the benchmark model (URM) from the literature were tested with the forest Las Bayas, using information obtained from the SiPlaFor project from Universidad Juárez. The proposed model was solvable in all tested instances. Furthermore, a sensitivity analysis study over a core data set of test instances was carried out on the different parameters of the GURM model to determine optimal configurations for the specific case study. Several environmental measures were assessed in our experimental work. The parameters evaluated were the distance value between pairs of units harvested in the same period, the distance value between those considered natural reserve units, the timber volume to be harvested, the green-up period, and the minimum forest reserve area. An interesting observation from the experiments was that the maximum area inversely affected the URM and GURM models; larger regions resulted in a reduced number of management units in the URM model, thus reducing the computational time to solve the instance of the problem, but in this case, at the expense of a reduced profit. One of the interesting findings was that, in all experiments under all different factors, harvesting every 5 or 6 years yields better profits than harvesting every 10 or 12 years. The current standard in the Mexican system is to harvest every 10 years.

Bird community structure and habitat association in Owabi Wildlife Sanctuary, Ashanti Region (Ghana)

Context Globally, an estimated 1.3% of the bird species have gone extinct over the past millennia, largely owing to habitat loss. Aims This study investigated bird–habitat associations and assemblages in the Owabi Ramsar wetland. Methods The study was conducted over 5 months (May–September 2019) in four habitat types (agricultural land, built-up, forest reserve, and open-water area). Data were collected in 84 plots across four habitats (i.e. built-up area = 25, forest reserve = 25, agricultural land = 25 and open-water area = 9 plots), using a point-count technique. The Gambin model, non-metric multidimensional scaling, Chao-1, and Hill numbers models were used to evaluate differences in bird diversity and composition among the habitat types. Key results In all, 1260 individual birds, belonging to 81 species, were encountered. Many of the species occurred in the forest habitat (n = 46, 56.8%. Species from the agricultural land (n = 37) and built-up (n = 30) were mainly habitat generalists that used these two habitats as their shared feeding station. Bird–habitat specialisation grouping equally showed the forest specialists to be the highest (n = 23, 38.4%), whereas the open water had the least number of habitat specialisation associations (e.g. white-faced whistling duck, n = 20, 24.7%). Five species were widely distributed in all four habitats (e.g. bronze-mannikins and white-throated bee-eaters), indicating their broad habitat preferences and ability to adapt to varied conditions. The forest reserve tended to be the most diverse, which was likely mediated by factors such as nesting microhabitats, varied food availability, and human-led activities. Conclusions This study highlighted bird dietary structure and associated habitat type and bird-habitat specialisation in four different land-use types at Owabi Ramsar wetland. Implications Given the increasing level of disturbance, there is the likelihood that the population of forest-dependent and open water-dependent birds will be lost or will have to change or modify their behaviour to be similar to that of open-tolerant or open-country birds if managers of the Owabi Wildlife Sanctuary fail to implement the recommended management interventions highlighted.

Relationship of the Seasonal Vegetation Indices against the NDVI and LST in the Region of Kamuku Game Reserve and Kwiambana National Park, Nigeria

While it is anticipated that there would be some similarities amongst spectral Vegetation Indices (VIs) because the majority of the indices use the red and NIR bands, it is also expected that there would be some variances. The NDVI, derived earlier by Rouse (1973), and is the commonly used VI, there have meagre understanding of the relationship between the NDVI and another VIs. Similarly, investigations on the correlation between LST and other VIs (other than NDVI) in both dry and raining seasons have not been adequately explored. This motivated the study to determine the seasonal correlation of some spectral VIs against the NDVI and LST over the forest reserve area. The study investigated two categories of VIs: slope-based and distance-based. It derived spectral VIs from Landsat 8 images for dry (January) and raining (August) seasons; and estimated LST from MODIS. The findings showed that the ARVI, GNDVI and TVI not only showed resemblance in appearance with the NDVI in both seasons, but also had a high coefficient of correlation: ARVI = 0.973, 0.964; GNDVI = 0.919, 0.879; TVI = 0.779, 0.716. Based on this finding, the ARVI, GNDVI and TVI can be used to supplant the NDVI for biomass related studies in the study area. The study further revealed that the LST-VIs relationship was negative for both dry and rainy seasons, except for the distance-based VIs (DVI, SAVI, MSAVI) that specifically had a positive correlation with the LST. The LST was strongly correlated with the GNDVI, TVI, NDVI, ARVI (0.664 ? r ? 0.598). However, the strength of the correlation for the LST-VIs in the raining season was very weak (0.003 ? r ? 0.245). The study concluded that the correlation of the LST versus the ARVI, GNDVI, NDVI, and TVI can be used for climate related studies.

Human-Wildlife Conflict Management in Garhwal Himalayan Landscape: Present scenario, Issues, Challenges and Strategies

Human-wildlife conflict is an important issue in Uttarakhand for many decades. Severe loss incurred in terms of livestock predation, human injury, and death along with the crop raiding. Due to anthropogenic pressures on the wildlife habitats, large groups of wild animals are forced to occupy the multiuse landscapes outside the reserve forest areas. About 645 incidents have been recorded with a total of 740 casualties of livestock predation along with 126 incidents of human injured and 36 incident of human death in Narendranagar forest division between the periods of 2000 - 2020. Maximum attacks by wild animals were recorded when livestock was freely grazing without herder within the multi-use land. Leopard and elephant were mainly involved in human death whereas elephant and monkey in crop raiding. Present study also suggests supervised grazing, awareness of leopard prone areas, removing of unnecessary canopy of plants to reduce the conflicts cases in the study area. Further, the areas demand culturally and sustainable acceptable strategies to mitigate the damages and a comprehensive Human-Wildlife management plan should be formulated and implemented in each ranges of Narendranagar forest division.

Stratification Properties of Peninsular-Malaysian Peat

Peat is known for its importance as carbon storage. Peatlands in Malaysia especially forest reserve areas are generally lack of natural resources due to poor soil nutrients. Hence, many of peatlands in Malaysia have been converted to plantations and industrialized area. Peninsular-Malaysian peat properties are less studied in terms of its physical and chemical properties. A total of 38 peat samples from 5 depths (surface, 50cm, 100cm, 150cm and 200cm) from 10 locations around Peninsular-Malaysia were characterized for its physicochemical properties: moisture, organic matter, pH, carbon, hydrogen, nitrogen, humic acid content and its FTIR properties. Results showed that moisture content ranges from 60.92% to 94.22% and moisture increases with increasing depth. Organic matter percentage was very high, having an average of 95% (range 74-99%) with most samples having organic matter more than 90%. Peninsular Malaysian peat is quite acidic, with pH average of 3.56 owing to its humic and fulvic acid contents. The min % humic acid (HA) is 32.06% and the highest is 62.44%. There is a slight increase in HA content as the depth increases. C:N ratio increasing with depth, indicating very slow decomposition rate. FTIR results indicated the presence of polysaccharides, fat, wax, lipids and lignins content which increases as the depth increases, especially the aromatics such as humic acids and lignins. These distinct characteristics are described to provide a better understanding of the significance of sustaining peat swamp forest for long-term management.

Drivers of human-megaherbivore interactions in the Eastern and Western Ghats of southern India

The global effort to protect megaherbivore populations is largely dependent on how human-wildlife conflict is identified, prioritized, and remedied. We examined the socio-ecological and landscape-scale factors determining spatial patterns of human-megaherbivore (Asian elephant Elephas maximus and gaur Bos gaurus) interactions across sixteen Forest Divisions in Tamil Nadu, India. Using a systematic grid-based design, we conducted questionnaire-based surveys of 1460 households at the human-wildlife interface adjacent to Protected Areas, Reserve Forest and Fringe Areas. We specifically collected information on elephant and gaur conflict incidents (e.g., human death/injuries, property damage, and crop-raiding), cropland type, extent of crop area and area lost to crop-raiding, from each household. We found that human-elephant conflict increased with percentage of crop cover, diversity of major and minor crops grown, proximity to water source, flat terrain, and lower rates of precipitation. Human-gaur conflict was greatest with a high diversity of major crops, proximity to water source, moderate precipitation, and more undulating terrain. We identified ca. 7900 km2 hotspot area of contiguous high-intensity elephant conflict. For gaur, we identified high-frequency conflict hotspot areas covering ca. 625 km2, which were patchily distributed, highly localised, and attributed mostly to the recent changing land-use patterns. Our findings will help policymakers and park managers in developing landscape-scale human-wildlife conflict mitigation plans in the identified conflict hotspots.

An Annotated Checklist of Ethno Medicinal Tree Species in Terakanahalli Reserve Forest area of Sirsi range, Karnataka, (India)

Sirsi town is situated in the midst of Northern Central Western Ghats of Karnataka State. The forests adjoining to the town are known for its natural diversified floral wealth. These plants are used by local people as herbal medicine. Present study aims to prepare a check list of existing tree species in Terakanahalli reserved forest area situated adjacent to Sirsi town. The list was presented according to the Angiosperm Phylogeny Group IV (APG-IV) system of classification. Study enumerated a total of 63 tree species belonging 32 families from Terakanahalli reserve forest area. Enlisted tree species belongs to diverse families particularly Leguminosae, Anacardiaceae, Moraceae and Combretaceae. Ethno medicinal uses from Uttara Kannada district were also indicated for the listed tree species. Globally, Santalum album and Garcinia indica are categorised as vulnerable species with declining population by IUCN. The study area is under pressure because of anthropogenic activity and it needs proper management and conservation practices for sustainability.

A STUDY ON THE MUD PUDDLING ACTIVITY OF BUTTERFLIES IN ALAGARKOVIL HILLS, MADURAI, TAMIL NADU, INDIA

Butterflies are one of the most remarkable creatures of nature, which exhibits an interesting behavior called puddling. The term puddling is used to describe the supplementary feeding of minerals and salts from mud puddles, excrement and carrion. The mud puddling activity of lepidopterans have great significance in influencing the nutritional and reproductive status of the insects. This study aims to record the mud puddling activity of butterflies in Alagarkovil hills, a reserve forest area with rich biodiversity and high anthropogenic activity; and test the soil sample for various minerals and salts to understand the importance of mud puddling in the reproductive behavior of butterflies. The survey of butterflies was done on six sites in the Alagarkovil hills, from July 2017- January 2018 on sunny days, between 0800-1600 hrs, thrice a week using the transect method. A total of 111 species belonging to six families were recorded during the study period, of which the Lycaenidae, Nymphalidae and Pieridae families were mostly found to exhibit mud puddling. In agreement with previous findings, greater numbers of male butterflies were found to participate in mud puddling. Among the six study sites, the mud puddling activity was predominant in the site with copious puddles. The soil analysis of the mud puddling sites showed the presence of salts like sodium and potassium, metal ions like manganese, iron and copper, macro elements like phosphorus, nitrogen and carbon. The percentage of sodium and potassium were found to be higher, which were reported to be the chief elements consumed during mud puddling. The alkaline pH of the soil was found to favor the puddling activity. All these abiotic factors play a significant role in the nutritional and reproductive biology of the butterflies, which can be monitored to assess the community status and diversity of butterflies in an area. Conservation of mud puddling sites could help restore the biodiversity index of the butterflies.

ASSESSMENT OF THE SUSTAINABILITY OF FORESTRY AND STATE FOREST MANAGEMENT

This article, deals the momentum of the country's forest reserve area, the distribution of forest reserve lands, the dynamics of forest reserve lands by regions and the percentage share of forestry institutions by territory. The current period of economic development is characterized by the escalation of the environmental crisis, the creation of new post-industrial civilization bases, the concept of sustainable development, the change of socio-economic relations in the direction of the creation of various property forms and their implementation, and the strengthening of integration processes. International agreements have declared the need for the world community to move towards sustainable development. The purpose of the sustainable development strategy is to effectively solve the social and economic tasks of the current and future generations, as well as the preservation of natural resource potential. Forest is an important strategic resource of Kazakhstan. It plays a big role in human life. As the main component of the regulation of the natural environment and its processes, the forest ensures the vital activity of humanity, and it also occupies a special place in the socio-economic development of society. Observance of the principle of continuous use of forests in a non-depleting manner by all participants in forest relations is considered the basis for preserving and increasing the ecological and resource potential of forests. Also, reproduction of renewable forest resources is the use of resources in such a way as to maintain the stability of the forest as an ecological system

Assessment of land use change in the Thuma forest reserve region of Malawi, Africa

A study was conducted in Thuma area in central Malawi to quantify contemporary land cover and to explore the degree of land use change in the Thuma forest reserve area of Malawi by analysing and comparing satellite-derived land cover maps from 1997, 2007 and 2017. The study was carried out using Remote Sensing and Geographic Information System (GIS), focusing on analysis of Landsat 5 ETM and Landsat 8 ORI/TIRS satellite images. The classification was conducted for the following distinct classes; closed forest, open forest, shrubland, savanna grassland, agriculture fields, and water. The analysis revealed that closed forest diminished from 19% in 1997 to 10% in 2007 to 6% in 2017. Open forest reduced from 30% to 21% from 1997 to 2007 but increased to 22% in 2017. Agriculture area almost doubled from 37% in 1997 to 64% in 2017. The actual area from 1997 to 2017, shows that closed forest has reduced from 7,000 ha to 3,000 ha while open forest from 12,900 ha to 7800 ha. Savanna grassland has doubled from 5,900 ha to 13,000 ha. However, future studies should use modern satellites such as Sentinel and Landsat 9 for improved quantification of changes. The findings show that even the protected forest reserve (previously dominated by closed forest) is not fully protected from deforestation by local communities. Government and other stakeholders should devise measures to meet the needs of the surrounding communities and the ecological/biophysical needs of the reserves. Based on this study, issues of re-demarcation of the forest reserve and the accessed area should also be explored. This study serves as a reference for the management of Thuma Forest Reserve as a refuge for natural tree species, rivers that harbour endemic fish species (Opsaridium microlepis and Opsaridium microcephalis) and the sustainable management of endangered elephants in the reserve.

Contributions of some non-timber forest products to household income of rural dwellers in Onigambari Forest Reserve, Oyo State, Nigeria

The communities surrounding the forest reserve get their daily income from the sales of some of the non-timber forest products (NTFPs) as a means of survival for them and their house hold without proper documentation. The study described the socio-economic characteristics of the rural dwellers in the study area and determined the income made from some of these NTFPs. Data was generated through the use of structured questionnaires administered to 120 randomly selected rural household heads. Data obtained were described using frequency counts, percentages and mean scores; the relationship between dependent variables and independent variables of the study was tested using binomial test and logistic regression. Finding reveal that, non-timber forest products gathering in Onigambari was male dominated (72.3%), 78.1% were married and 38.1% had formal education. Contribution of income from non-timberforest products to total income of respondents in the State was 59.46%. It was concluded that the non-timber forest products contribute significantly to rural household income at Onigambari forest Reserve Area of Oyo State, Nigeria