Forager Societies Research Articles

Wild fruits, forager children and secret treats in pre-Columbian South Florida

The archaeology of childhood has shown that research is possible on even the most ephemeral aspects of ancient human lives if we ask the correct questions of the material record. Children are now visible members of ancient societies and their impact on the physical environment in which they lived can be identified. This paper looks at food remains possibly recovered by children as a lens into one of the secret places and spaces of childhood. Fruit is often gathered by children in forager societies, in small peer groups that incorporate play, learning and socialisation into food provisioning. Using palaeobotanical and environmental data from the hunter-fisher-gatherer Tequesta society situated at the mouth of the Miami River along Biscayne Bay in south Florida, this paper suggests the human biological predilection for sugar facilitated a uniquely powerful relationship between plants, children and fruit trees. Fruit would have satisfied a powerful biological craving for sugar in this marine-resources focused culture. The relationship between children, fruit and wild food gathering shaped the foodscape of ancestral south Florida and the children who lived there. The lens of plant studies allows us to better appreciate the active role plants had in shaping human behaviour and landscape experiences.

Freshwater exploitation at Ajvide - Pitted ware culture fishing practises investigated through laser ablation facilitated strontium isotope analyses

The importance of marine resources for the Neolithic hunter-fisher-gathers of the Pitted Ware Culture of Gotland, Sweden, is well documented through zooarchaeological analyses and diet studies of human remains. Terrestrial areas were important for living and supplementing the diet but the extent of the terrestrial territories and regions of land use for different groups is largely unknown. The presence of euryhaline species in recovered zooarchaeological assemblages indicates that freshwater fishing or fishing in the brackish estuaries of the Baltic Sea was part of the subsistence practises. To explore if the inland freshwaters of Gotland were used and, if exploited, where they were located, 18 teeth from euryhaline fish from the Pitted Ware Culture site Ajvide on Gotland were selected. The 87Sr/86Sr isotope ratios in the fish teeth were analysed using laser ablation-multi collector-inductively coupled plasma-mass spectrometry and correlated with an updated bioavailable baseline of Gotlandic water sources. Through this approach, the habitational origin of the fish was shown to primarily stem from at least six freshwater sources located in the west-central area of Gotland, in close relation to the site, with a few individuals originating from within the Baltic Sea. The study highlights the significance of ichthyoarchaeological analysis in understanding the territorial practice of past foraging societies and recommends further studies on euryhaline species to expand our knowledge of fish habitat, human resource utilization and land use.

Ostrich eggshell beads from Little Muck Shelter, southern Africa: First impressions and regional perspectives

Hunting and gathering communities in southern Africa produced ostrich eggshell (OES) beads for personal ornamentation, trade and exchange, and various forms of symbolism. OES beads convey information related to not only technological processes, but also social histories, making them useful tools for investigating these processes in the archaeological record. In the middle Limpopo Valley, hunter-gatherers produced beads from periods that predated the arrival of farmer societies, before ca. AD 150, until the decline of the Mapungubwe capital, AD 1300. Their analysis may therefore lead to insights into local economies, craft activities, trade and exchange, and social roles. However, no study in the middle Limpopo Valley has sufficiently investigated these beads and their status within forager society. In this paper we study the manufacture of OES beads from one of the excavated forager-occupied shelters in the valley, Little Muck. This study is the first of its kind from a hunter-gatherer context in the region. It shows that Little Muck’s beads were standardised but produced in varying frequencies across temporal periods, and made following different production strategies. Use-wear analysis hints that some of the beads were pigmented and possibly altered using heat treatment. As an initial foray into a forager bead assemblage of the region, this study demonstrates the value such an approach may yield, and it aids in guiding future attempts. It also compares Little Muck’s bead assemblage with those from other sites across a much wider region where such studies have taken place. The findings demonstrate similar patterns to other parts of southern and eastern Africa, but also illustrate local shifts in bead production that follow changes in local socio-political dynamics.

Feral ecologies of the human deep past: multispecies archaeology and palaeo‐synanthropy

AbstractThis article articulates recent advances in palaeo‐ecology with the goals and ambitions of multispecies archaeology. It centres the synanthropic nexus as a key context for the study of early human‐animal relationships and argues that its evolution yields important yet currently overlooked dynamics shaping the structure of the archaeological record. I first show how the dominant heuristic of wild versus domesticated nonhuman animals obfuscates the inherent variability and creativity of past animal agency. I then illustrate how such agency is caught up with human behaviour and ecosystem impacts from relatively early on in human history, navigating historically shifting configurations of autonomy and control. Drawing on deep‐historical examples of synanthropic niches co‐assembled by human foragers and nonhuman animals, I argue that such interspecies configurations require careful attention to concepts of liminality and ferality and challenge species‐level approaches. I finally highlight potential human behavioural, material, and cosmological consequences of the synanthropic nexus, including integrative foraging patterns and notions of the giving animal, which play an important role in many Indigenous and ethnographic forager societies and illustrate the value of attending to this nexus as a focus of comparative multispecies research.

The Prevalence of Behavioural Symptoms and Psychiatric Disorders in Hadza Children

The worldwide pooled prevalence of psychiatric disorders in children is 13.4%. Studying the prevalence of childhood psychiatric disorders across radically different economic systems and social structures could indicate universal factors leading to their development. The prevalence of childhood psychiatric disorders in a mixed-subsistence foraging society has not been studied. The Strengths and Difficulties Questionnaire and the Development and Well-Being Assessment were used to compare the prevalence of behavioural symptoms and psychiatric disorders in Hadza children aged 5–16 years (n = 113) to a nationally representative sample from England (n = 18,029) using a cross-sectional study design. Emotional problems, conduct problems and hyperactivity were lower in the Hadza children. Prosocial behaviour and peer problems were higher in Hadza children. 3.6% of Hadza children met the criteria for a psychiatric disorder compared to 11.8% of English children. All psychiatric disorders in Hadza children were co-morbid with autism spectrum disorder. No child from the Hadza group met the criteria for an emotional, behaviour or eating disorder. Further work should study the factors which lead to the different prevalence of psychiatric disorders in Hadza children.

Human-beaver cohabitation in the Early and Mid-Holocene of Northern Europe: Re-visiting Mesolithic material culture and ecology through a multispecies lens

The Eurasian beaver ( Castor fiber) was an important member of Early and Mid-Holocene landscapes and animal communities in Northern Europe. Previous zooarchaeological research has established the alimentary roles of beavers for Mesolithic societies and their importance for fur procurement. In this paper, we develop an integrated biocultural approach to human-beaver interactions, examining the position of humans and beavers in Mesolithic and Early Neolithic multispecies systems. We contextualize beaver landscape agency in hydroactive environments with human behaviour, synthesizing currently available data on mammalian assemblages, ichtyofauna and beaver-related material culture across Northern Europe. This cross-cultural, diachronic analysis reveals previously overlooked facilitations of human behaviour by beaver practices and ecological legacies. We show that long-term trajectories of human-beaver cohabitation differed between northern European regions. While in Southern Scandinavia, human-beaver intersections witnessed major re-organizations during the Mid-Holocene, beavers retained a key role for human societies across Northeastern Europe throughout much of the Holocene and played an important part at the Mesolithic-Neolithic transition in the Dutch wetlands. Divergent pathways are also evidenced by Mesolithic beaver-related material culture, highlighting the cultural keystone status of Castor fiber in higher latitude European landscapes. We argue that this keystone status is grounded in the supply of human hunting, fishing, and gathering affordances by the animals, pointing to the generative commensality between Mesolithic foragers and their beaver neighbours. Taken together, our findings demonstrate the importance of the beaver in the making of Early and Mid-Holocene forager societies in Northern Europe and illustrate the fruitfulness of deploying an integrated multispecies approach.

‘The Stars Know Where He Is’: World-Making, Wayfaring, and Navigational Theory in Southern African |Xam Forager Folklore

Forager societies have had a profound influence on the natural world as modernity has come to know it, engineering environments over the course of millennia through consistent interventions in the lifeways of the plant and animal species around them. Leveraging the human capacity for processing social information, these societies preserved and transmitted their understanding of these processes through folklore. Taking the example of the |Xam foragers of nineteenth-century southern Africa, this article examines the narratives related to navigation and spatial education as a means of articulating value in a landscape, and ordering the lives of those who passed through it. Amongst a society for whom absolute references were rare, the |Xam communicated in stories that overlapped personal life history, deep mythology, and momentary politics, preserving an understanding of how to track the sun, moon, stars, and many social others besides, reckoning with an ever-shifting definition of what ‘time’ meant for ‘place’.

Evidence for an emotional adaptive function of dreams: a cross-cultural study

The function of dreams is a longstanding scientific research question. Simulation theories of dream function, which are based on the premise that dreams represent evolutionary past selective pressures and fitness improvement through modified states of consciousness, have yet to be tested in cross-cultural populations that include small-scale forager societies. Here, we analyze dream content with cross-cultural comparisons between the BaYaka (Rep. of Congo) and Hadza (Tanzania) foraging groups and Global North populations, to test the hypothesis that dreams in forager groups serve a more effective emotion regulation function due to their strong social norms and high interpersonal support. Using a linear mixed effects model we analyzed 896 dreams from 234 individuals across these populations, recorded using dream diaries. Dream texts were processed into four psychosocial constructs using the Linguistic Inquiry and Word Count (LIWC-22) dictionary. The BaYaka displayed greater community-oriented dream content. Both the BaYaka and Hadza exhibited heightened threat dream content, while, at the same time, the Hadza demonstrated low negative emotions in their dreams. The Global North Nightmare Disorder group had increased negative emotion content, and the Canadian student sample during the COVID-19 pandemic displayed the highest anxiety dream content. In conclusion, this study supports the notion that dreams in non-clinical populations can effectively regulate emotions by linking potential threats with non-fearful contexts, reducing anxiety and negative emotions through emotional release or catharsis. Overall, this work contributes to our understanding of the evolutionary significance of this altered state of consciousness.

Mesolithic Scandinavian foraging patterns and hunting grounds targeted through laser ablation derived 87Sr/86Sr ratios at the Early-Mid Holocene site of Huseby Klev on the west coast of Sweden

Mobility is one of the most fundamental aspects of a foraging society. Since prehistoric mobility is often difficult to identify in the archaeological record, our understanding is largely based on comparison with ethnographic communities. In recent years the application of 87Sr/86Sr isotope analysis has, however, greatly broadened our knowledge of mobility in the past. Despite this, few studies have been undertaken on faunal remains to explore their mobility patterns and infer human exploitation patterns with more precision. In this contribution we sampled 28 mammal teeth from three different occupation phases at the Early to Mid-Holocene coastal site of Huseby Klev, Sweden. We first established the local baseline for seven geographical areas in the region surrounding Huseby Klev. Then, by applying laser ablation-multi collector-inductively coupled plasma-mass spectrometry to the selected teeth, we identified the likely origins of a range of terrestrial and marine fauna, and the possible human mobility patterns required in their exploitation. Overall, our results demonstrate that the prehistoric communities inhabiting Huseby Klev undertook both short- and long-distance forays for the exploitation of particular species. By relating inferences on hunting grounds, derived from strontium isotope data, to zooarchaeological evidence from the site and ethnographic human mobility patterns, we establish and discuss the prerequisites for landscape utilization. Lastly, we demonstrate that glacial meltwater may have temporally affected the local oceanic Sr ratios – suggesting significantly increased soil and bedrock weathering may influence the Sr values in aquatic ecosystems and, consequently, should be considered in such regions and at times of melting glaciers. By applying the method to additional sites and assemblages in the future, our understanding of prehistoric mobility will be greatly enhanced.

Socioecology shapes child and adolescent time allocation in twelve hunter-gatherer and mixed-subsistence forager societies

A key issue distinguishing prominent evolutionary models of human life history is whether prolonged childhood evolved to facilitate learning in a skill- and strength-intensive foraging niche requiring high levels of cooperation. Considering the diversity of environments humans inhabit, children’s activities should also reflect local social and ecological opportunities and constraints. To better understand our species’ developmental plasticity, the present paper compiled a time allocation dataset for children and adolescents from twelve hunter-gatherer and mixed-subsistence forager societies (n = 690; 3–18 years; 52% girls). We investigated how environmental factors, local ecological risk, and men and women’s relative energetic contributions were associated with cross-cultural variation in child and adolescent time allocation to childcare, food production, domestic work, and play. Annual precipitation, annual mean temperature, and net primary productivity were not strongly associated with child and adolescent activity budgets. Increased risk of encounters with dangerous animals and dehydration negatively predicted time allocation to childcare and domestic work, but not food production. Gender differences in child and adolescent activity budgets were stronger in societies where men made greater direct contributions to food production than women. We interpret these findings as suggesting that children and their caregivers adjust their activities to facilitate the early acquisition of knowledge which helps children safely cooperate with adults in a range of social and ecological environments. These findings compel us to consider how childhood may have also evolved to facilitate flexible participation in productive activities in early life.

Fluidities of personhood in the idioms of the Maloti-Drakensberg, past and present, and their use in incorporating contextual ethnographies in southern African rock art research

ABSTRACT In the Maloti-Drakensberg mountains of southern Africa, beliefs about snakes and their representations in rock art images are emblematic of hybrid histories of regional societies. The snake symbol initially represented an attempt at ‘reaching out’ as forager societies incorporated a prominent figure in the mythologies of incoming societies into their own – a figure which became a symbolic reference to cross-cultural symbiosis and admixture. Reflecting the long history of such contact, the ritual uses and ontological positions of snakes in contemporary knowledge systems of the Maloti-Drakensberg are coherent with those of earlier societies. This offers fertile ground for novel forms of interpretation. Using contextual historic and modern ethnographic material, this paper presents a relational account of regional idioms. It dwells on the language of taming and domestication that permeate these ethnographies, and the concern they show for the mitigation of ‘wild’, sometimes ‘monstrous’, consequences of spiritual power in the social world. Symbolic resolutions of these consequences are discernible in rock art images, particularly those of snakes, demonstrating the ritual brokerage of relations between human and non-human communities, with both forms of agency depicted in various states of ‘domestication’, bridging forager and farmer understandings of human–animal relations.

Case study of a rare occipital osteoma related to cranial trauma.

Burial 280 is a 31- to 40-year-old male Native American individual who lived in a 10th to 16th century coastal foraging society in California. His remains display a large ectocranial elevation on the occipital, measuring 23.3 mm anteroposteriorly, 25.3 mm mediolaterally, and is elevated to 20.1 mm. It is located along the superior nuchal line on the left side of the occipital and represents an extremely rare occipital localization of a particular type of trauma-related osteoma. In our prior study of Burial 280, trauma etiology was not considered; herein, we present evidence for such a cause. Such trauma-related neoplasms are sometimes called ballooned (or giant, depending on size) osteomas. The presence of this tumor appears to have resulted in asymmetrical cranial muscle use and consequent skeletal changes. This case study illustrates that cranial trauma may sometimes induce benign tumor elevations, rather than a depression or pronounced fracture.

Reproductive seasonality in the Baka Pygmies, environmental factors and climatic changes.

Reproductive seasonality is a phenomenon common to human and animal populations and driven by, among others, climatic variables. Given the currently changing climate and its impacts on both the environment and human lives, the question arises of its potential effects on reproductive seasonality. Few studies have specifically explored the seasonality of reproduction among hunter-gatherers and anyone investigated how current climate change might affect this phenomenon. In this study we addressed reproductive seasonality in the Baka Pygmy living in African rain forests. Since reproductive seasonality can be linked to weather patterns, we explore this possibility. However, climatic variables driving weather patterns have changed over the years, so we assessed whether this has influenced the Baka reproductive pattern. Based on 34 years of written birth records and oral questionnaires from 13 years of systematic fieldwork, we observed a bimodal birth pattern with two birth peaks at 6-month intervals. Our results demonstrate that precipitation at conception or at birth potentially has effects, respectively negative and positive on the monthly number of births; and temperature has a role in controlling other variables that do affect the reproductive pattern. Changing weather patterns appear to be affecting the reproductive seasonality in the Baka, suggesting that attention needs to be given to the influence of global climate change on forager societies.

The Cost of Gathering Among the Baka Forager-Horticulturalists From Southeastern Cameroon

What present-day foragers do for their living and what they eat have long been privileged areas for exploring human behavior, global health, and human evolution. While many studies have focused on hunting and meat acquisition, less attention has been given to gathering and plant foods. Despite evidence of variation in both nutritional quality and energetic costs of gathering different plants, the overall effort spent on gathering in relation to other subsistence tasks is still under explored. In the current context of economic, climate, and social changes, many forager societies also rely on other subsistence strategies, including agriculture and wage labor. In this study, we aim to explore the place of gathering in the livelihood of a mixed economy society, the Baka forager-horticulturalists of southeastern Cameroon, by comparing the involvement and the costs of activities related to food acquisition. From a pool of 153 adult participants (97 women and 56 men), we collected 246 daily records using a GPS (Global Positioning System) tracker combined with heart rate monitor and time allocation recalls. We compared the duration, distance traveled, and the intensity of work, measured by calculating the metabolic equivalent of task (MET), of subsistence activities related to food acquisition. Results from this work show that gathering activities, performed by both women and men, are energetically costly, with higher MET values than hunting and fishing activities. Furthermore, the MET values vary depending on the targeted plant foods. We discuss these insights in the overall framework of subsistence patterns, merging them with the socio-cultural and environmental factors that might explain Baka livelihood and subsistence strategy.

Co-occurrence of Ostensive Communication and Generalizable Knowledge in Forager Storytelling : Cross-Cultural Evidence of Teaching in Forager Societies.

Teaching is hypothesized to be a species-typical behavior in humans that contributed to the emergence of cumulative culture. Several within-culture studies indicate that foragers depend heavily on social learning to acquire practical skills and knowledge, but it is unknown whether teaching is universal across forager populations. Teaching can be defined ethologically as the modification of behavior by an expert in the presence of a novice, such that the expert incurs a cost and the novice acquires skills/knowledge more efficiently or that it would not acquire otherwise. One behavioral modification hypothesized to be an adaptation for teaching is ostensive communication-exaggerations of prosody and gesture that signal intent to transmit generalizable knowledge and indicate the intended receiver. On this view, the use of ostensive communication in conjunction with the transmission of generalizable knowledge constitutes evidence of teaching. Oral storytelling appears to meet these criteria: Indigenous peoples regard their traditions as important sources of ecological and social knowledge, and oral storytelling is widely reported to employ paralinguistic communication. To test this hypothesis, descriptions of performed narrative in forager societies were coded for the use of 14 ostensive-communicative behaviors and the presence of generalizable knowledge. Although biased toward North America, the study sample comprised 53 forager cultures spanning five continents, 34 language families, and diverse biomes. All cultures evinced the predicted behaviors. Results suggest that foragers use storytelling as a mode of instruction, thus providing cross-cultural evidence of teaching in forager populations.

Dynamics of Wealth Inequality in Simple Artificial Societies

A simple generative model of a foraging society generates significant wealth inequalities from identical agents on an equal opportunity landscape. These inequalities arise in both equilibrium and non-equilibrium regimes with some societies essentially never reaching equilibrium. Reproduction costs mitigate inequality beyond their affect on intrinsic growth rate. The highest levels of inequality are found during non-equilibrium regimes. Inequality in dynamic regimes is driven by factors different than those driving steady state inequality.

Comparative Study of Territoriality across Forager Societies

Land tenure systems vary considerably across forager societies. The economic defensibility model explains this variation by considering the costs and benefits of defending resources that vary in density and predictability. The purpose of our study was to describe and explain the variation in tenure systems across 30 hunter-gatherer societies from across the world using the economic defensibility model. We used qualitative data from the Human Relations Area Files (HRAF) database on defense and sharing of resources among groups, and quantitative data from Binford’s Frames of References (2001) as indicators of resource density, resource predictability, and competition for resources. However, we were unable to explain the variation in territoriality using these independent variables. We argue that the diversity and complexity of foragers’ subsistence strategies is one of the main reasons for our failure to find correlations between territoriality and resource density and predictability. We propose that it may better to conceptualize tenure systems in foraging societies as assemblages of multiple property regimes.

Inter- and intra-cultural variation in learning-through-participation among Hadza and BaYaka forager children and adolescents from Tanzania and the Republic of Congo

We examined cross-cultural variation in children’s learning-through-participation in economic work in two forager societies; the Hadza of Tanzania and the BaYaka of the Republic of Congo. We used observational data from 46 Hadza (41% female) and 65 BaYaka (48% female) children and adolescents between the ages of 3 and 18; interview data from 73 Hadza (49% female) and 52 BaYaka (56% female) adults; and ethnographic observations from both populations. Results showed that by providing tools, assigning chores, and foraging with children, Hadza and BaYaka adults provided opportunities for autonomous learning through facilitating participation. Furthermore, although both Hadza and BaYaka children foraged alongside adults when they could be of help, Hadza children were more likely than BaYaka children to forage independently, and BaYaka children were more likely than Hadza children to participate in domestic tasks. We argue that these strategies provided children with opportunities to learn while contributing economically.Nous avons examiné les variations interculturelles dans l’apprentissage des enfants dans le contexte de leur participation à des travaux économiques dans deux sociétés de chasseurs-cueilleurs; les Hadza de Tanzanie et les BaYaka du Congo. Nous avons utilisé les données d’observation de 46 enfants et adolescents Hadza (41% femmes) et 65 BaYaka (48% femmes) âgés de 3 à 18 ans, celles d’interviews de 73 adultes Hadza (49% femmes) et de 52 adultes BaYaka (56% femmes), ainsi que des observations ethnographiques sur ces deux groupes. Les résultats montrent qu’en leur fournissant des outils, en leur assignant des tâches et en chassant et en cueillant avec les enfants, les adultes Hadza et BaYaka leur offrent des possibilités d’apprentissage autonome tout en facilitant leur participation. En outre, alors que les enfants Hadza et BaYaka chassent et cueillent aux côtés des adultes quand ils peuvent, les enfants Hadza sont plus susceptibles que les enfants BaYaka de recourir à la chasse et à la cueillette. Ces derniers sont aussi plus susceptibles que les enfants Hadza de participer aux tâches domestiques. Nous en concluons que ces stratégies fournissent aux enfants des opportunités d’apprentissage tout en contribuant, parallèlement, aux ressources économiques de leur groupe.

Coalitional Play Fighting and the Evolution of Coalitional Intergroup Aggression

Dyadic play fighting occurs in many species, but only humans are known to engage in coalitional play fighting. Dyadic play fighting is hypothesized to build motor skills involved in actual dyadic fighting; thus, coalitional play fighting may build skills involved in actual coalitional fighting, operationalized as forager lethal raiding. If human psychology includes a motivational component that encourages engagement in this type of play, evidence of this play in forager societies is necessary to determine that it is not an artifact of agricultural or industrial conditions. We examine whether coalitional play fighting appears in the hunter-gatherer record and includes motor skills used in lethal raiding. Using the ethnographic record, we generated a list of motor patterns regularly used in forager warfare. Then, using Murdock's Ethnographic Atlas, we identified 100 culture clusters containing forager societies and searched the ethnographic records of these societies for descriptions of coalitional play fighting, operationalized as contact games played in teams. Resulting games were coded for the presence of eight motor patterns regularly used in forager lethal raiding. Although play does not tend to be systematically documented in the hunter-gatherer literature, sufficiently detailed descriptions of coalitional play were found for 46 of the 100 culture clusters: all 46 exhibited coalitional play using at least one of the predicted motor patterns; 39 exhibited coalitional play using four or more of the eight predicted motor patterns. These results provide evidence that coalitional play fighting (a) occurs across a diverse range of hunter-gatherer cultures and habitats, (b) regularly recruits motor patterns used in lethal raiding, and (c) is not an artifact of agricultural or industrial life. This is a first step in a new line of research on whether human male psychology includes motivations to engage in play that develops the deployment of coordinated coalitional action involving key motor patterns used in lethal raiding.

Late Pleistocene Lithic Procurement and Geochemical Characterization of the Cerro Kaskio Obsidian Source in South‐western Bolivia

Primary questions regarding the foraging behaviour of the first hunter–gatherers who colonized the New World are how they found, procured and utilized high‐quality raw materials for manufacturing stone tools. In this paper, we present evidence from the late Pleistocene site of Cueva Bautista in the highlands of south‐western Bolivia, which demonstrates that a substantial portion of the recovered stone tool assemblage originated in Cerro Kaskio, a recently discovered obsidian source located 15 km south‐west of the site. In addition to describing the geological and geochronological setting, we provide the first geochemical characterization of the Cerro Kaskio source by means of instrumental neutron activation analysis and energy‐dispersive X‐ray fluorescence. Supported on the technological analysis and archaeometric sourcing of the obsidian lithics found in Cueva Bautista, we discuss the nature of the procurement strategies practised by the earliest mobile hunter–gatherers who explored and colonized the Andean highlands. We conclude that opportunistic lithic resource procurement was probably an important component of the first foraging societies that explored the highland Andes.