Flint Hills Research Articles

Kansas Windscape - 2016 Status

Kansas has played a significant role for the growth of wind energy in the United States. Kansas ranks sixth among states for wind-energy capacity, and is third in the nation for per-capita production of electricity from the wind. Wind farms are sited in topographically high positions on uplands, drainage divides, escarpments, and other rural sites in which there are few or no obstacles to interrupt near-surface wind flow. The best regions of the state include the High Plains, especially in southwestern Kansas, the Blue Hills and Flint Hills, and on the Missouri-Arkansas drainage divide in the Osage Cuestas. Rapid development of wind energy has taken place in these regions during the past few years, except for the Flint Hills, which is an exclusion zone for new wind farms. Bird and bat mortality remains an issue, but with appropriate wildlife mitigation much more wind energy could be developed in Kansas. Aesthetic issues include visibility and noise. Developing safe and renewable energy resources that have...

Patch‐burn grazing increases habitat heterogeneity and biodiversity of small mammals in managed rangelands

AbstractHabitat heterogeneity is a key driver of biodiversity in many ecosystems. Wildlife inhabiting the native prairies of North America evolved in a heterogeneous mosaic of habitat conditions created by fire and grazing by native ungulates. Current rangeland management practices in the tallgrass prairie ecosystem evenly distribute fire and grazing across management units and promote homogeneous habitat conditions. Patch‐burn grazing is a rangeland management strategy that seeks to restore heterogeneity to rangelands via fire‐grazing interactions. Our 3.5‐year study tested the effects of patch‐burn grazing on habitat heterogeneity and small mammal community dynamics in the Flint Hills ecoregion of eastern Kansas. To study the ecological effects of patch‐burn grazing, we sampled habitat conditions and the small mammal community. We assessed habitat conditions once each growing season in a negative control that was annually burned and grazed, a positive control that was burned every four years and ungrazed, and within each of three units of a patch‐burn grazing experiment (PBG) managed with rotational fire. Habitat conditions were significantly different among treatments, and a principal components analysis showed that the patch‐burn grazing treatment had higher canopy cover of forbs and habitat heterogeneity than our two control units. To sample the small mammal community, we conducted monthly live trapping of small mammals on two randomly located trap grids in each of our two controls and three units of our PBG treatment. Small mammal diversity was significantly higher in the patch‐burn grazing treatment and in the positive control, vs. the negative control. Moreover, a canonical correspondence analysis showed that a fire‐grazing interaction was the major driver structuring small mammal communities. Patch‐burn grazing is an effective strategy for restoring heterogeneity to vegetative structure and composition, and can increase biodiversity of small mammals in managed rangelands in the tallgrass prairie ecosystem.

High dissimilarity within a multiyear annual record of pollen assemblages from a North American tallgrass prairie.

Grassland vegetation varies in composition across North America and has been historically influenced by multiple biotic and abiotic drivers, including fire, herbivory, and topography. Yet, the amount of temporal and spatial variability exhibited among grassland pollen assemblages, and the influence of these biotic and abiotic drivers on pollen assemblage composition and diversity has been relatively understudied. Here, we examine 4 years of modern pollen assemblages collected from a series of 28 traps at the Konza Prairie Long‐Term Ecological Research Area in the Flint Hills of Kansas, with the aim of evaluating the influence of these drivers, as well as quantifying the amount of spatial and temporal variability in the pollen signatures of the tallgrass prairie biome. We include all terrestrial pollen taxa in our analyses while calculating four summative metrics of pollen diversity and composition – beta‐diversity, Shannon index, nonarboreal pollen percentage, and Ambrosia:Artemisia – and find different roles of fire, herbivory, and topography variables in relation to these pollen metrics. In addition, we find significant annual differences in the means of three of these metrics, particularly the year 2013 which experienced high precipitation relative to the other 3 years of data. To quantify spatial and temporal dissimilarity among the samples over the 4‐year study, we calculate pairwise squared‐chord distances (SCD). The SCD values indicate higher compositional dissimilarity across the traps (0.38 mean) among all years than within a single trap from year to year (0.31 mean), suggesting that grassland vegetation can have different pollen signatures across finely sampled space and time, and emphasizing the need for additional long‐term annual monitoring of grassland pollen.

Contribution of regional-scale fire events to ozone and PM2.5 air quality estimated by photochemical modeling approaches

Two specific fires from 2011 are tracked for local to regional scale contribution to ozone (O3) and fine particulate matter (PM2.5) using a freely available regulatory modeling system that includes the BlueSky wildland fire emissions tool, Spare Matrix Operator Kernel Emissions (SMOKE) model, Weather and Research Forecasting (WRF) meteorological model, and Community Multiscale Air Quality (CMAQ) photochemical grid model. The modeling system was applied to track the contribution from a wildfire (Wallow) and prescribed fire (Flint Hills) using both source sensitivity and source apportionment approaches. The model estimated fire contribution to primary and secondary pollutants are comparable using source sensitivity (brute-force zero out) and source apportionment (Integrated Source Apportionment Method) approaches. Model estimated O3 enhancement relative to CO is similar to values reported in literature indicating the modeling system captures the range of O3 inhibition possible near fires and O3 production both near the fire and downwind. O3 and peroxyacetyl nitrate (PAN) are formed in the fire plume and transported downwind along with highly reactive VOC species such as formaldehyde and acetaldehyde that are both emitted by the fire and rapidly produced in the fire plume by VOC oxidation reactions. PAN and aldehydes contribute to continued downwind O3 production. The transport and thermal decomposition of PAN to nitrogen oxides (NOX) enables O3 production in areas limited by NOX availability and the photolysis of aldehydes to produce free radicals (HOX) causes increased O3 production in NOX rich areas. The modeling system tends to overestimate hourly surface O3 at routine rural monitors in close proximity to the fires when the model predicts elevated fire impacts on O3 and Hazard Mapping System (HMS) data indicates possible fire impact. A sensitivity simulation in which solar radiation and photolysis rates were more aggressively attenuated by aerosol in the plume reduced model O3 but does not eliminate this bias. A comparison of model predicted daily average speciated PM2.5 at surface rural routine network sites when the model predicts fire impacts from either of these fires shows a tendency toward overestimation of PM2.5 organic aerosol in close proximity to these fires. The standard version of the CMAQ treats primarily emitted organic aerosol as non-volatile. An alternative approach for treating organic aerosol as semi-volatile resulted in lower PM2.5 organic aerosol from these fires but does not eliminate the bias. Future work should focus on modeling specific fire events that are well characterized in terms of size, emissions, and have extensive measurements taken near the fire and downwind to better constrain model representation of important physical and chemical processes (e.g. aerosol photolysis attenuation and organic aerosol treatment) related to wild and prescribed fires.

Assessing the Potential for Transitions from Tallgrass Prairie to Woodlands: Are We Operating Beyond Critical Fire Thresholds?

A growing body of evidence suggests humans are pushing ecosystems near or beyond key ecological thresholds, resulting in transitions to new, sometimes undesirable phases or states that are costly to reverse. We used remotely sensed fire data to assess if the Flint Hills—a landscape of tallgrass prairie in the Central Great Plains, United States—is operating beyond fire frequency thresholds. Long-term fire experiments and observational evidence suggests that applying prescribed fire at return intervals > 3 yr can lead to transitions from grassland to shrubland. Fire return intervals > 10 yr and complete fire suppression, in particular, can result in transitions to woodlands over 30−50 yr. Once shrublands and woodlands are established, restoration back to grassland is difficult with prescribed fires. We applied these fire frequency cutoffs to remotely sensed fire data from 2000 to 2010 in the Flint Hills, identifying the extent of tallgrass prairie susceptible to shrub and tree expansion. We found that 56% (15 620 km2) of grasslands in this region are burned less than every 3 yr and are therefore susceptible to conversion to shrub or tree dominance. The potential effects of this large-scale shift are greater risk for evergreen (Juniperus virginiana) woodland fires, reduced grazing potential, and increased abundance of woodland adapted species at the expense of the native grassland biota. Of the 12 127-km2 area likely to remain grassland, 43% is burned approximately annually, contributing to vegetative homogenization and potential air-quality issues. While this synthesis forecasts a precarious future for tallgrass prairie conservation and their ecosystem services, increases in shrub or tree dominances are usually reversible until fire frequency has been reduced for more than 20 yr. This delay leaves a small window of opportunity to return fire to the landscape and avoid large-scale transformation of tallgrass prairie.

Effects of Intensive Late-Season Sheep Grazing Following Early-Season Steer Grazing on Population Dynamics of Sericea Lespedeza in the Kansas Flint Hills

Sericea lespedeza (Lespedeza cuneata; SL) is a high-tannin, invasive forb in the Tallgrass Prairie ecosystem. In Kansas, sericea lespedeza infests 980 square miles of pasture, primarily in the Flint Hills region. Sericea lespedeza infestations reduce native grass production by up to 92% through a combination of aggressive growth, prolific reproduction, canopy dominance, and chemical inhibition (allelopathy). Herbicides retard the spread of sericea lespedeza, but application is laborious and expensive; moreover, herbicides are lethal to ecologically-important, non-target plant species. Increased grazing pressure on sericea lespedeza by domestic herbivores may slow its spread and facilitate some measure of biological control. Unfortunately, mature plants contain high levels of condensed tannins, which are a strong deterrent to grazing by beef cattle. Small ruminants have greater tolerance for condensed tannins than beef cattle. Sheep, in particular, appear less susceptible to certain plant toxins than beef cattle and may be useful to selectively pressure noxious weeds like sericea lespedeza. The predominant grazing management practice in the Flint Hills region of Kansas involves annual spring burning followed by intensive grazing with yearling beef cattle from April to August. During seasonal grazing, 40 to 60% of annual graminoid production is removed and pastures remain idle for the remainder of the year. Under this prevailing management practice, invasion by sericea lespedeza into the Tallgrass Prairie biome has steadily increased. Sericea lespedeza flowers and produces seed in late summer from August to September. The absence of grazing pressure during this interval strongly promotes seed production, seed distribution, and continued invasion of the Flint Hills ecoregion by this noxious weed. Therefore, the objective of our study was to evaluate effects of late-season sheep grazing following locally-conventional steer grazing on vigor and reproductive capabilities of sericea lespedeza.

Effects of Growing-Season Prescribed Burning on Vigor of the Noxious Weed Sericea Lespedeza (Lespedeza cuneata) in the Kansas Flint Hills

Sericea lespedeza (SL) was introduced into the United States from Asia in the late 19th century. Early land managers recognized that SL was adaptable; tolerant of shallow, acidic or low-fertility soils; and resistant to insects and disease. This combination of traits made SL a widely-used plant for reseeding strip-mined lands, highway right-ofways, dams, and waterways in the US for nearly a century. Regrettably, SL is highly fecund. Individual plants are capable of producing up to 850 lb of seed per acre annually. Vigorous seed production allows SL to rapidly infiltrate native grasslands that are adjacent to reseeding projects; seed can be transported great distances via the alimentary canal and hair of wild and domestic herbivores. In Kansas alone, SL has infested approximately 980 square miles of pasture, primarily in the Flint Hills region. The resulting damage to native habitats for wildlife and pasture quality for domestic herbivores has been devastating. The predominant grazing management practice in the Kansas Flint Hills involves annual spring burning in March or April, followed by intensive grazing with yearling beef cattle for a relatively short period from April to August. During seasonal grazing, 40 to 60% of annual graminoid production is removed and grazing lands then remain idle for the remainder of the year. Under this prevailing management practice, invasion by SL into the Tallgrass Prairie biome has steadily increased. Oklahoma State University researchers speculated that dormant-season, spring fires may stimulate SL growth by scarifying seeds lying on the surface of the soil. In contrast, plants with robust canopies respond more strongly to growing-season prescribed burns than to dormant-season prescribed burns. Previous research reported that application of growing season fire at 3-yr intervals decreased the rate of SL invasion. Therefore, the objective of our study was to evaluate the effects of growing-season prescribed burning of native tallgrass range on vigor of sericea lespedeza.

Emporia State University's Natural Areas: A General Overview of the Areas and Research Pertaining to Hydrogeology and Climate Change

Emporia State University (ESU) manages eight sites (5 – 81 ha) that serve as outdoor laboratories for our students, faculty, and visiting scientists. The sites provide regional representation of major ecological communities, including tallgrass prairie, forest, streams, ponds, and wetlands. The sites have been used for decades, with the oldest site, the Ross Natural History Reservation, having been acquired in 1958. Common uses of our Natural Areas include field courses in ecology, botany, zoology, geospatial analysis, soil science, geology, and hydrology. Recent research has included investigations of hydrogeology of the Neosho and Cottonwood River basins and the effects of shrub encroachment on carbon dynamics in tallgrass prairie. The former has included long-term monitoring of rivers and groundwater at ESU's on-campus Hydrogeology Teaching and Research Station. Hydrologic conditions at a newly acquired, riverine wetland site in the Flint Hills are also currently under investigation.

A Statistical Approach for Predicting Grassland Degradation in Disturbance-Driven Landscapes

Maintaining a land base that supports safe and realistic training operations is a significant challenge for military land managers which can be informed by frequent monitoring of land condition in relation to management practices. This study explores the relationship between fire and trends in tallgrass prairie vegetation at military and non-military sites in the Kansas Flint Hills. The response variable was the long-term linear trend (2001-2010) of surface greenness measured by MODIS NDVI using BFAST time series trend analysis. Explanatory variables included fire regime (frequency and seasonality) and spatial strata based on existing management unit boundaries. Several non-spatial generalized linear models (GLM) were computed to explain trends by fire regime and/or stratification. Spatialized versions of the GLMs were also constructed. For non-spatial models at the military site, fire regime explained little (4%) of the observed surface greenness trend compared to strata alone (7% - 26%). The non-spatial and spatial models for the non-military site performed better for each explanatory variable and combination tested with fire regime. Existing stratifications contained much of the spatial structure in model residuals. Fire had only a marginal effect on surface greenness trends at the military site despite the use of burning as a grassland management tool. Interestingly, fire explained more of the trend at the non-military site and models including strata improved explanatory power. Analysis of spatial model predictors based on management unit stratification suggested ways to reduce the number of strata while achieving similar performance and may benefit managers of other public areas lacking sound data regarding land usage.

A Critical Examination of Timing of Burning in the Kansas Flint Hills

Frequent burning is a crucial ecological and economic component of the Kansas Flint Hills. Although burning is important for the preservation of tallgrass prairie and improving livestock production, it has become a controversial societal issue because of its potential impact on air quality standards. Over the past 80years, recommendations on burning in Kansas have ranged from total fire exclusion to burning only in late April; and for the past 40years, the concept that burning should only occur in late spring has become ingrained in the cultural practices of rangeland management. Yet the scientific basis for these recommendations has received little rigorous scrutiny. Herein, we critically review the research on dormant-season burning in the Flint Hills that formed the foundation for modern burn practices in Kansas. Close examination of the historical data does not support the tenet that burning must be limited to a narrow window in late spring. Many conclusions of the research that led to recommending burning only in late spring were ambiguous, not subjected to statistical analysis, or were influenced by an antiburn bias. Current research suggests that timing of a burn is not as critical as ranchers have been led to believe and burning does not have to be restricted to a narrow window in late April. There is an absence of scientific evidence that burning earlier in the spring adversely affects forage production, plant species composition, soil moisture, or cattle weight gain. Although there is a need for research on the consequences of burning grazed pastures at different times of the year, expanding the window for burning earlier in the dormant season should help alleviate air quality issues downwind of the burned areas and potentially be beneficial to ranchers.

Bison and cattle grazing management, bare ground coverage, and links to suspended sediment concentrations in grassland streams

AbstractThis study quantified the impact of bison and cattle grazing management practices on bare ground coverage at the watershed, riparian, and forested riparian scales within the Flint Hills ecoregion in Kansas. We tested for correlations between bare ground coverage and fluvial suspended sediment concentrations during base‐flow and storm‐flow events. We used remotely sensed imagery combined with field surveys to classify ground cover and quantify the presence of bare ground. Base‐flow water samples were collected bi‐monthly during rain‐free periods and 24 h following precipitation events. Storm‐flow water samples were collected on the rising limb of the hydrograph, using single‐stage automatic samplers. Ungrazed treatments contained the lowest coverage of bare ground at the watershed, riparian, and forested riparian scales. Bison treatments contained the highest coverage of bare ground at the watershed scale, while high‐density cattle treatments contained the highest coverage of bare ground at the riparian and forested riparian scales. In bison and cattle‐grazed treatments, a majority of bare ground was located near fence lines, watershed boundaries, and third‐ and fourth‐order stream segments. Inorganic sediment concentrations at base flow were best predicted by riparian bare ground coverage, while storm‐flow sediment concentrations were best predicted by watershed scale bare ground coverage.

Dynamic Disturbance Processes Create Dynamic Lek Site Selection in a Prairie Grouse.

It is well understood that landscape processes can affect habitat selection patterns, movements, and species persistence. These selection patterns may be altered or even eliminated as a result of changes in disturbance regimes and a concomitant management focus on uniform, moderate disturbance across landscapes. To assess how restored landscape heterogeneity influences habitat selection patterns, we examined 21 years (1991, 1993–2012) of Greater Prairie-Chicken (Tympanuchus cupido) lek location data in tallgrass prairie with restored fire and grazing processes. Our study took place at The Nature Conservancy’s Tallgrass Prairie Preserve located at the southern extent of Flint Hills in northeastern Oklahoma. We specifically addressed stability of lek locations in the context of the fire-grazing interaction, and the environmental factors influencing lek locations. We found that lek locations were dynamic in a landscape with interacting fire and grazing. While previous conservation efforts have treated leks as stable with high site fidelity in static landscapes, a majority of lek locations in our study (i.e., 65%) moved by nearly one kilometer on an annual basis in this dynamic setting. Lek sites were in elevated areas with low tree cover and low road density. Additionally, lek site selection was influenced by an interaction of fire and patch edge, indicating that in recently burned patches, leks were located near patch edges. These results suggest that dynamic and interactive processes such as fire and grazing that restore heterogeneity to grasslands do influence habitat selection patterns in prairie grouse, a phenomenon that is likely to apply throughout the Greater Prairie-Chicken’s distribution when dynamic processes are restored. As conservation moves toward restoring dynamic historic disturbance patterns, it will be important that siting and planning of anthropogenic structures (e.g., wind energy, oil and gas) and management plans not view lek locations as static points, but rather as sites that shift around the landscape in response to shifting vegetation structure. Acknowledging shifting lek locations in these landscapes will help ensure conservation efforts are successful by targeting the appropriate areas for protection and management.

College and Career Ready: What’s the Library Got to Do With It?

College and Career Ready, Career Clusters, and Career Pathways are buzz words often heard but not always understood. These terms will be discussed as information about the collaborative activities between Emporia High School (EHS) and Flint Hills Technical College (FHTC) is shared. One responsibility of Emporia High School guidance counselors is to assist students in exploring and choosing career pathway courses. To support the counselors, the librarians at both institutions have created resources and activities that engage students in career exploration. The FHTC librarian created a LibGuide that includes online handbooks, reports, and crosswalks from FHTC programs to the six career fields and sixteen career clusters. Resources and activities created by the EHS librarian for the career fair, which helped students explore their learning styles and career interests, will be discussed. Although the activities are specific to these institutions they can be generalized for other settings.

Alternative Rangeland Management Strategies and the Nesting Ecology of Greater Prairie-Chickens

Population declines of grassland birds over the past 30 yr have followed the widespread implementation of intensive rangeland management practices that create homogenous grassland habitats. Patch-burn grazing (PBG) was tested as an alternative management technique that is ecologically similar to historically heterogeneous fire and grazing regimes and holds promise as a rangeland management tool that may benefit grassland wildlife. We conducted a 3-year study to compare nest-site selection and nest survival of greater prairie-chickens, an umbrella species for tallgrass prairie conservation, on private lands managed with PBG or intensive fire and grazing in the Flint Hills of Kansas. The goal of our field study was to evaluate the relationships among rangeland management practices, habitat conditions, and nesting ecology of greater prairie-chickens. Nest-site selection and nest survival of prairie-chickens were both directly related to vertical nesting cover, which was determined by the fire return interval of a pasture. Nesting habitat was affected little by stocking rate in PBG management regimes because preferred nest sites were unburned patches that were not grazed by cattle. Overall, the quantity and quality of nesting sites was improved under PBG management when compared with more intensive rangeland management regimes. Our results join a growing body of evidence that rangeland management strategies that mimic historical heterogeneous fire and grazing regimes benefit native species of prairie wildlife.

Resolution of Respect: Paul Risser, 1939–2014

Resolution of Respect: Paul Risser, 1939–2014

Multiple watershed alterations influence fish community structure in Great Plains prairie streams

AbstractStream fish distributions are commonly linked to environmental disturbances affecting terrestrial landscapes. In Great Plains prairie streams, the independent and interactive effects of watershed impoundments and land cover changes remain poorly understood despite their prevalence and assumed contribution to declining stream fish diversity. We used structural equation models and fish community samples from third‐order streams in the Kansas River and Arkansas River basins of Kansas, USA to test the simultaneous effects of geographic location, terrestrial landscape alteration, watershed impoundments and local habitat on species richness for stream‐associated and impoundment‐associated habitat guilds. Watershed impoundment density increased from west to east in both basins, while per cent altered terrestrial landscape (urbanisation + row‐crop agriculture) averaged ~50% in the west, declined throughout the Flint Hills ecoregion and increased (Kansas River basin ~80%) or decreased (Arkansas River basin ~30%) to the east. Geographic location had the strongest effect on richness for both guilds across basins, supporting known zoogeography patterns. In addition to location, impoundment species richness was positively correlated with local habitat in both basins; whereas stream‐species richness was negatively correlated with landscape alterations (Kansas River basin) or landscape alterations and watershed impoundments (Arkansas River basin). These findings suggest that convergences in the relative proportions of impoundment and stream species (i.e., community structure) in the eastern extent of both basins are related to positive effects of increased habitat opportunities for impoundment species and negative effects caused by landscape alterations (Kansas River basin) or landscape alterations plus watershed impoundments (Arkansas River basin) for stream species.

Ecological consequences of shifting the timing of burning tallgrass prairie.

In the Kansas Flint Hills, grassland burning is conducted during a relatively narrow window because management recommendations for the past 40 years have been to burn only in late spring. Widespread prescribed burning within this restricted time frame frequently creates smoke management issues downwind. A potential remedy for the concentrated smoke production in late spring is to expand burning to times earlier in the year. Yet, previous research suggested that burning in winter or early spring reduces plant productivity and cattle weight gain while increasing the proportion of undesirable plant species. In order to better understand the ecological consequences of burning at different times of the year, plant production and species abundance were measured for 20 years on ungrazed watersheds burned annually in autumn, winter, or spring. We found that there were no significant differences in total grass production among the burns on either upland or lowland topographic positions, although spring burned watersheds had higher grass culm production and lower forb biomass than autumn and winter burned watersheds. Burning in autumn or winter broadened the window of grass productivity response to precipitation, which reduces susceptibility to mid-season drought. Burning in autumn or winter also increased the phenological range of species by promoting cool-season graminoids without a concomitant decrease in warm-season grasses, potentially widening the seasonal window of high-quality forage. Incorporating autumn and winter burns into the overall portfolio of tallgrass prairie management should increase the flexibility in managing grasslands, promote biodiversity, and minimize air quality issues caused by en masse late-spring burning with little negative consequences for cattle production.

Woodland Voles Captured Among Sparse Shrubs in Native Tallgrass Prairie on Konza Prairie Biological Station, Kansas

From autumn 1981 through spring 2009, woodland voles (Microtus pinetorum) were captured rarely and only in woodland habitats (four in ca. 15,000 trap nights) and never in grassland habitats (ca. 300,000 trap nights) on the Konza Prairie Biological Station, Kansas. Konza Prairie is located in the Flint Hills region of eastern Kansas in Geary and Riley counties (see Knapp and Seastedt 1998 or http://keep.konza.ksu.edu/visit/map. htm for maps and location). We summarized characteristics of these voles (all four voles captured in moist gallery forest; Kaufman and Kaufman 2009) as well as their locations and habitat features because small mammals only were to be sampled in grassland sites away from gallery forest habitats as we completed our long-term studies of small mammals during autumn 2009 through spring 2013 (nearly 30,000 more trap nights during this time period). Unexpectedly, two woodland voles were captured in a prairie site during March 2013, which was the last sampling period of our 32 year study. Herein, we describe the characteristics of the two voles captured along a prairie trapline plus features of the habitat surrounding capture locations.

Consumption and performance by beef heifers provided dried distillers grains in a self-fed supplement containing either 10 or 16% salt while grazing Flint Hills native grass (2014)

Introduction Optimizing cattle performance and maintaining pasture health are important considerations when striving to maximize profitability and sustainability on a Flint Hills pasture yearling grazing operation. The two growing seasons prior to initiation of this study were droughty and stressful to pastures. This situation provided an opportunity to evaluate the value of dried distillers grains with solubles (DDGS) as a self-fed supplement to ensure that nutritional resources were adequate for a 78-day grazing period. Grazing density was increased from 200 to either 225 or 250 lb of beef while simultaneously providing salt-limited DDGS supplements containing 10 and 16% salt, respectively.

Winter Bird Habitat Use in a Heterogeneous Tallgrass Prairie

In the Flint Hills of Kansas and Oklahoma, U.S.A., grazing land is managed predominantly to promote homogeneous grassland structure. This potentially limits the diversity of habitats for grassland obligate songbirds with narrow habitat preferences during the breeding season, prompting ecologists and conservationists to call for managing rangelands for increased heterogeneity. The Flint Hills also hosts multiple species of conservation concern during winter, but avian habitat requirements are less well known during this period and seldom considered in management recommendations. We investigated the influence of vegetation structure on occurrence of songbirds overwintering at The Nature Conservancy's Tallgrass Prairie Preserve in Osage County, Oklahoma, where fire and grazing are used to increase structural and compositional heterogeneity. During the winters of 2008–2009 and 2009–2010, we used an area-search method to survey 149, 1 ha plots distributed among patches that represented a wide gradient in vegetation structure and 1–3 growing seasons since fire. We modeled occurrence of Le Conte's Sparrow (Ammodramus leconteii), Sedge Wren (Cistothorus platensis), Savannah Sparrow (Passerculus sandwichensis), and Smith's Longspur (Calcarius pictus) with a combination of fine-scale vegetation (e.g., litter depth in sampling plots), landscape-scale factors (e.g., distance to utility poles), and time since fire. Vegetation height was the most consistent predictor for three species, with Le Conte's Sparrow and Sedge Wren associated with tall structure and Smith's Longspur associated with shorter structure. Occurrence of Smith's Longspur was too low to adequately model effects of burn age, but fine-scale vegetation structure used by Le Conte's Sparrow and Sedge Wren was available in plots irrespective of time since fire. A variety of management techniques for creating habitat heterogeneity have the potential to promote grassland songbird diversity during winter, but further study is needed.