Eocene Research Articles

First Australian amber fossil of Podonominae (Diptera: Chironomidae) from the late Middle Eocene

A non-biting midge of the cosmopolitan family Chironomidae (Podonominae) is recorded in amber from the Anglesea Coal Measures, southeastern Australia. The new morphotype is likely a new species but is not formally described here because of insufficient diagnostic character states. To date, only five amber deposits in the Northern Hemisphere have yielded fossils of Podonominae, making our discovery the first amber-hosted representative of the group from the Southern Hemisphere. The single adult male individual has been assigned to the subfamily Podonominae, and shows similarities to the Austrochlus cluster group including small size, the lack of veins R2 + 3, and the distally located cross-vein m-cu. Synchrotron X-ray microtomographic imaging reveals some internal structures of the wing musculature and brain capsule, highlighting the taphonomic significance of this unique specimen. Maria Blake* [ maria.paulsen@monash.edu ], School of Earth, Atmosphere and Environment, 9 Rainforest Walk, Monash University, Clayton, Victoria, 3800, Australia Enrique Peñalver [ e.penalver@igme.es ], Instituto Geológico y Minero de España, CSIC, Valencia, Spain Jeffrey D. Stilwell [ jeffrey.stilwell@monash.edu ], School of Earth, Atmosphere and Environment, 9 Rainforest Walk, Monash University, Clayton, Victoria, 3800, Australia, and Australian Museum, Australian Museum, 1 William Street, Sydney, NSW 2010, Australia Joseph J. Bevitt [ jbv@ansto.gov.au ], Australian Centre for Neutron Scattering, Australian Nuclear Science and Technology Organisation, Lucas Heights, New South Wales, Australia Daniel J. Bickel [ Dan.Bickel@australian.museum ] Australian Museum, 1 William Street, Sydney, New South Wales, 2000, Australia.

Fish otoliths from the basal Oligocene Viborg Formation in Denmark

The Viborg Formation is the lowest part of the early Oligocene in Denmark and represents the upper part of nannoplankton zone NP21. Here we describe a small assemblage of otoliths from the Viborg Formation obtained from five localities that yielded 88 specimens of nine species. The composition of the otolith-based f ish species is transitional between the terminal Eocene (Priabonian) faunas from the Latdorf Formation of northern Germany and its equivalents and the middle and upper Rupelian. The Viborg otolith association witnessed the first arrivals of the subsequently dominant Gadidae (Trisopterus elegans) and the migration of certain species of the merlucciid genus Palaeogadus from the Eastern Paratethys through the Polish gateway. Furthermore, the otoliths from the Viborg Formation f ill an important gap in the previous knowledge of late Eocene and early Oligocene otolith associations in the North Sea Basin and thus facilitate some refinement of their biostratigraphic usage.

New fossil woods from the middle Eocene climate optimum of north-central Turkey

The middle Eocene climate optimum, a crucial greenhouse event in Earth’s life history, occurred approximately 40 Ma. Fossil wood identifications of middle Eocene age reveal important evidence for the terrestrial tropical/subtropical conditions during this period. The purpose of the present study is to identify new fossil woods from the middle Eocene of north-central Turkey and to evaluate the paleoclimate and paleovegetational history of the north-central Turkey based on new records. Seven fossil woods were collected from the middle Eocene Göynücek Volcanics of the Çekerek Formation (Sarıkaya village, close to Çekerek, Yozgat). Two new species Apocynoxylon umut-tuncii Akkemik and Mantzouka, n. sp., and Ficoxylon anatolica Akkemik and Mantzouka, n. sp., are described, together with two more coniferous species Pinuxylon cf. P. tarnocziense (Tuzśon, 1901) Greguss, 1954 and Juniperoxylon cf. J. acarcaeaAkkemik, 2021a. Although dealing with a rather low number of fossil wood findings, indistinct boundaries of the tree rings, low xeromorphy ratios and high conductivity values in these two new fossil angiosperm species and less visible and indistinct boundaries in the two fossil coniferous species may reflect the warm, humid tropical conditions of the middle Eocene of north-central Turkey. The results are consistent with our former paleoecological evaluation based on the fossil woods including Actinodaphnoxylon zileensis Akkemik and Mantzouka in Akkemik et al., 2021 from the same geological unit and Palmoxylon sabaloidesGreguss, 1969 from the same area.

Lutetian–Bartonian (Middle Eocene) micropalaeontology, biostratigraphy and palaeoecology of the Mengen Coal Basin, Northwest Türkiye (Turkey)

This study focuses on the upper Lutetian–lower Bartonian Tokmaklar Formation in the Mengen Basin of Northwest Türkiye (Turkey), examining various aspects such as biostratigraphy, palaeoecology, and isotope data (δ18O and δ13C), providing insights into the geological history and environmental conditions during this interval. Moreover, the study incorporates coal petrography on four samples to explore a potential correlation between the coalification process and the preservation of palynomorphs. The formation is informally divided into two distinct categories: a coal-bearing sequence positioned at the base of the formation and a coal-free sequence above it. However due to dense vegetation and soil cover in the area, the availability of suitable outcrops for measured sections and obtaining samples is severely limited. Therefore, we relied on drilling data from Turkish Coal Enterprises (TCE) for the purpose of coal exploration, as well as two partial sections (Ender and Çorak) from underground mining. Microapalaeontological evidences involving benthic foraminiferans, ostracods, and palynomorphs, along with isotope data (δ18O and δ13C), suggest that the Tokmaklar Formation was deposited as a result of a local transgressive process during the late Lutetian–early Bartonian. Furthermore, palynological data, particularly the existence of mangroves such as Nypa, lepidocaryoid palm and Acrostichum aureum, coupled with quantitative palaeoclimate analysis, the negative δ18O values, and the lower δ13C values indicate a warm, probably tropical climate under high rainfall.

Unravelling controls on multi‐source‐to‐sink systems: A stratigraphic forward model of the early–middle Cenozoic of the SW Barents Sea

AbstractSource‐to‐sink dynamics are subjected to complex interactions between erosion, sediment transfer and deposition, particularly in an evolving tectonic and climatic setting. Here we use stratigraphic forward modelling (SFM) to predict the basin‐fill architecture of a multi‐source‐to‐sink system based on a state‐of‐the‐art numerical approach. The modelling processes consider key source‐to‐sink parameters such as water discharge, sediment load and grain size to simulate various sedimentary processes and transport mechanisms reflecting the dynamic interplay between erosion in the catchment area, subsidence, deposition and filling of the basin. The Cenozoic succession along the SW Barents Shelf margin provides a key area to examine controls on source‐to‐sink systems along a transform margin that developed during the opening of the North Atlantic when Greenland and Eurasian plates were separated (ca. 55 Ma onwards). Moreover, the gradual cooling which culminated in major glaciations in the northern hemisphere during the Quaternary (ca. 2.7 Ma), has affected the spatio‐temporal evolution of the sediment routing along the western Barents Shelf margin. This study aims to characterize the relative importance of different source areas within the source‐to‐sink framework through SFM. In the early Eocene, the SW Barents Shelf experienced a relatively equal sediment delivery from three principal source areas: (i) Greenland to the north, (ii) the Stappen High to the east, representing a local source terrain, and (iii) a major southern source (Fennoscandia). In the middle Eocene, our best‐fit modelling scenario suggests that the northern and the local eastern sources dominated over the southern source, collectively supplying large amounts of sand into the basin as evidenced by the submarine fans in Sørvestsnaget Basin. In the Oligocene (ca. 33 Ma) and Miocene (ca. 23 Ma), significant amounts of sediments were sourced from the east due to shelf‐wide uplift. Finally, this study highlights the dynamic nature and controls of sediment transfer in multi‐source‐to‐sink systems and demonstrates the potential of SFM to unravel tectonic and climatic signals in the stratigraphic record.

Mineral chemistry and garnet U-Pb dating in the Bizmişen iron skarn deposit, Erzincan, East-Central Türkiye

In Bizmişen area, Middle Eocene (46.3–42.0 Ma) plutonic rocks (quartz diorite) were intruded into Triassic-Cretaceous limestones and Upper Cretaceous ophiolites and caused to skarn zones containing prograde (garnet, clinopyroxene) and retrograde (amphibole, epidote, calcite, magnetite, phlogopite, serpentinite and chlorite) minerals. Due to contact relationships of quartz diorites with both serpentinized ultramafic rocks and limestone blocks, the mineralogical associations represent calcic (grossular/andradite, amphibole, chlorite and epidote) as well as magnesian (diopside, serpentine, phlogopite, talc) skarns. During the skarnization process, garnet, diopside, epidote, scapolite and amphibole minerals developed in the endoskarn zone within quartz-diorite, and diopside, tremolite, serpentine, phlogopite, talc, Mg-chlorite and calcite minerals developed in the exoskarn zones in ophiolites (serpantinite and listwaenite) and limestones. Skarn and magmatic clinopyroxenes in serpentinized ultramafic hosted exoskarn zone have a compositional range of Wo45-53En35-53Fs0–15 and Wo24-51En38-76Fs0–11, and classified as diopside and augite, respectively. The end members of garnets are Adr46–54Grs43–50Prp1–2Sps1.1–5 in amphibole-epidote-calcite-garnet skarn in endoskarn zone, whereas Adr40–93Grs4–57Prp0.2–2.3Sps0.9–2 in epidote-calcite-garnet skarn in exoskarn zone, and classified as andradite-rich grandite garnets. FeO, MnO and partially MgO contents gradually increase from the edges to the center in zoned garnet grains. Amphibole compositions represent to calcic (Ca >1 atoms per formula unit) group (pargasite) and indicate the high oxygen fugacity conditions which caused the increase in the rate of magnetite crystallization. Al2O3 and FeO contents of epidotes range 23.68–27.63 and 9.10–14.45 wt%, respectively, correspond to composition of epidote and clinozoisite. The iron oxide contents (FeO, wt%) of magnetites change from 91.72 to 98.36 correspond to structural Fe3+1.88–1.97 and Fe2+0.91–1.0 values. The contents of Ti, divalent and trivalent metal cations correspond to ulvospinel-magnetite compositional line at the magnetite end-member. The mineral chemistry of skarn minerals shows similarities to those of most iron skarn deposits. Garnet U-Pb ages (46.95–45.63 Ma) are consistent with the cooling ages of the Bizmişen pluton. The prograde stage of skarn should be occurred syn-intrusion, whereas retrograde stage should be developed later than youngest garnet age (45.6 Ma) and before the oldest late stage argillic alteration (37.5 Ma).

Shallow-water whale-fall communities: Evidence from the middle–late Eocene basilosaurid whale bones, Wadi El-Hitan, Fayum, Egypt

The present study examines the basilosaurid whale bone specimens collected from the Sandouk El-Borneta section, Wadi El-Hitan (Valley of Whales), Fayum, Egypt. These specimens are embedded in highly fossiliferous calcareous sandstones of the middle Eocene Gehannam and the upper Eocene Birket Qarun formations. These whale bones display some post-mortem alterations, representing good signs of at least three distinct stages in what is called the whale fall. The co-occurrence of shark, ray, and crocodile fossil specimens with the examined whale bones may indicate the first mobile-scavenger stage. Borings of the siboglinid polychaete genus Osedax represent the most common worm type that deeply bioeroded the whale bones, contributing to their rapid degradation, and representing the enrichment-opportunist stage. These fossil traces of the bone-eating worm Osedax represent the first record in the Eocene Epoch regionally and inter-regionally, filling the gap between the Late Cretaceous and the Oligocene occurrences. Subsequently, the studied whale bones served as hard substrates for some calcareous tube-dwelling encrusting polychaetes, balanoid barnacles, sheet‐like cheilostome bryozoans, and scleractinian corals. Furthermore, a bioerosion structure produced by polychaete annelids was also observed. The presence of these sclerobionts assemblage confirms the well-developed final reef stage with prolonged exposure and colonisation of these whale bones prior to final burial. Based on the recorded post-mortem alterations, together with other sedimentological and palaeontological data, the studied whale bones were deposited in a shallow open marine bay to sheltered gulf environments, which were characterised by low depositional energy, low to moderate rate of sedimentation, and high surface water productivity.

Ypresian–Lutetian platform evolution in the Indus Basin, Southwestern Pakistan: An interplay between local and regional tectonic changes

The present investigation aims to analyze the depositional environment, sequence stratigraphy, and geochemistry of the Early–Middle Eocene succession in the Laki Range of the Southern Indus Basin, Pakistan. The sequence is subsequently correlated with other sections within the Southern Indus Basin, Central Indus Basin, Upper Indus Basin, and Hazara Basin in Pakistan. For this study, the Lakhra and Laki formations were sampled from four sections in the Laki Range. Based on detailed outcrop observations, eight lithofacies were identified including four clastic lithofacies (CLF-1 to 4) and four limestone lithofacies (LLF-1 to 4). Similarly, based on detailed petrographic examination, seven microfacies were identified in limestone units. Considering the biotic paleoecology, facies texture, and chemical composition, depositional environments were assigned to the microfacies and lithofacies. All the microfacies, and the shale and sandstone lithofacies (CLF-1 and CLF-4) represent shallow inner to deeper outer-shelf settings whereas the other two lithofacies (CLF-2 and CLF-3) represent deltaic and tidal flat settings, respectively. All the microfacies and lithofacies information was used to establish sequence stratigraphy for the studied strata. The top of the Lakhra Formation and the whole of the Laki Formation in the Bara Nala Section (BNS) represent two complete, and two partial, third-order sequences, further divided into fourth order and small-scale cycles. The sequence's development is primarily influenced by regional and local tectonics. The regional correlation of the Indus Basin and Hazara Basin indicates that the regional basin's bathymetry and resultant depositional sequence were significantly influenced by the Himalayan Orogeny. The regional depositional pattern indicates that subsequent to India–Asia collision, the closure of eastern Tethys took place in a temporal succession from northwestern to the southwestern parts of the Indus Basin. In the Central Indus Basin, marine environments persisted until the Priabonian Stage (SBZ20) compared to those of the Upper (SBZ11–12) and Southern (SBZ13–14) Indus basins. This study offers valuable insights into both the local and regional depositional frameworks and the influence of local and regional tectonics on a carbonate platform evolution.

Heterogeneous occurrence of evergreen broad-leaved forests in East Asia: Evidence from plant fossils

Evergreen broad-leaved forests (EBLFs) are widely distributed in East Asia and play a vital role in ecosystem stability. The occurrence of these forests in East Asia has been a subject of debate across various disciplines. In this study, we explored the occurrence of East Asian EBLFs from a palaeobotanical perspective. By collecting plant fossils from four regions in East Asia, we have established the evolutionary history of EBLFs. Through floral similarity analysis and paleoclimatic reconstruction, we have revealed a diverse spatio-temporal pattern for the occurrence of EBLFs in East Asia. The earliest occurrence of EBLFs in southern China can be traced back to the middle Eocene, followed by southwestern China during the late Eocene–early Oligocene. Subsequently, EBLFs emerged in Japan during the early Oligocene and eventually appeared in central-eastern China around the Miocene. Paleoclimate simulation results suggest that the precipitation of wettest quarter (PWetQ, mm) exceeding 600 mm is crucial for the occurrence of EBLFs. Furthermore, the heterogeneous occurrence of EBLFs in East Asia is closely associated with the evolution of the Asian Monsoon. This study provides new insights into the occurrence of EBLFs in East Asia.

The first mydid fly (Diptera: Mydidae) from the Paleocene maar of Menat (France)

AbstractPaleomydas menatensis gen. et sp. nov., the third known fossil mydid fly, is described from the Paleocene of Menat (France). The fossil fly is remarkable in its very broad hind femur, a rather infrequent character in this family, which is mainly present in the Neotropical genera Mapinguari and Ceriomydas, and to a less pronounced degree, in the Nearctic genus Phyllomydas. But the lack of information on the antenna and genitalia structures forbids us to specify its relationships within the Mydidae more precisely. Thus we propose to attribute it to a new genus. The systematic affiliation of the late Eocene Mydas miocenicus from Florissant is discussed. Because of the rarity of Diptera in the insect assemblage of the Menat outcrop, the present discovery could suggest that this nowadays rather rare family was more frequent during the Paleocene than today.

A new Trachyaspis-like pan-cheloniid turtle from the Bartonian of Kazakhstan

The article describes a fossil pan-cheloniid Protrachyaspis shorymensis gen. et sp. nov. from the Karakeshi, Kert, Kuyulus, and Monata localities of the Shorym Formation (Bartonian, middle Eocene), as well as unknown localities of the Mangyshlak Peninsula, Kazakhstan. In addition, the shell bones of small pan-cheloniids from the Kuyulus and Tuzbair localities of the Shorym Formation with some traits of P. shorymensis are described, which probably represent remains of juvenile specimens of this species. The new taxon is characterized by a number of features rarely found in pan-cheloniids, including serrated dentaries, distally displaced lateral process of the humerus, and deeply sculptured external carapace surface. These features indicate a likely herbivorous diet and pelagic lifestyle of the new pan-cheloniid. According to the results of the cladistic analysis, P. shorymensis is sister to the Neogene species Trachyaspis lardyi Meyer, 1843, from which it differs in the absence of ridges on the carapace in adult individuals, the configuration of the scutes on the parietal bone, and the structure of the plastron. The stratigraphic gap between the appearance of P. shorymensis and its sister T. lardyi suggests a long ghost lineage of members of this clade throughout the Bartonian – Aquitanian (ca. 20.7 million years) preceding the appearance of T. lardyi. In most trees, the P. shorymensis + T. lardyi clade is located within the Chelonini clade, which, taking into account the middle Eocene age of P. shorymensis, indicates the early divergence of crown cheloniids, previously established based on molecular data. The new taxon is similar to the fragmentary remains of pan-cheloniids with a sculptured external surface of the shell, previously described from four localities of the upper part of the Buchak and lower part of Kiev formations in the south of European Russia and Ukraine, which makes it possible to determine these materials as cf. Protrachyaspis sp., and probably extends the appearance of the P. shorymensis + T. lardyi clade back to the middle Lutetian.

PETROLEUM GEOLOGY OF THE CENOZOIC SUCCESSION IN THE ZAGROS OF SW IRAN: A SEQUENCE STRATIGRAPHIC APPROACH

The Cenozoic stratigraphy of the Zagros records the ongoing collision between the Arabian and Eurasian Plates and the closure of NeoTethys. A Paleogene NW‐SE trending foreland basin was inherited from a Late Cretaceous precursor. Widespread progradation into the foredeep was a feature of both margins which, allied to ongoing tectonism, had by the late Eocene led to the narrowing and subsequent division of the foredeep into the Lurestan – Khuzestan and Lengeh Troughs, separated by the northward continuation of the rejuvenated Qatar‐Fars Arch. This sub‐division strongly influenced subsequent deposition and the petroleum geology of the area. In addition, the diachronous nature of the Arabian – Eurasian collision led to strong diachroneity in lithostratigraphic units along the length of the Zagros. Hence its petroleum geology is best understood within a regional sequence stratigraphic framework. This study identifies three tectono‐megasequences (TMS 10, TMS 11a, TMS 11b) and multiple depositional sequences.The Cenozoic contains a world class hydrocarbon province with prolific oil reservoirs in the Oligo‐Miocene Asmari Formation sealed by the evaporite‐dominated Gachsaran Formation, mostly contained within giant NW‐SE trending “whaleback” anticlines concentrated in the Dezful Embayment. Reservoirs in the SW are dominantly siliciclastic or comprise mixed siliciclastics and carbonates, whereas those to the east and NE are dominated by fractured carbonates. There remains untested potential in stratigraphic traps, especially in deeperwater sandstone reservoirs deposited along the SW margin of the foredeep.Late Miocene to Pliocene charge to the Asmari reservoirs was mostly from Aptian – Albian Kazhdumi Formation source rocks. In some fields, an additional component was from organic‐rich late Eocene to earliest Oligocene Pabdeh Formation source rocks confined to the narrowing Lurestan – Khuzestan Trough. Where mature, the latter source rock is also a potential unconventional reservoir target, although the prospective area is limited due to recent uplift and erosion. Deeper Jurassic source rocks contributed to the Cheshmeh Khush field in Dezful North. Silurian source rocks charged gas‐bearing structures in the Bandar Abbas region.

Evaluation of Reservoir Properties of Sylhet Limestone of Jaintia Group, North-Eastern Sylhet, Bangladesh

The Sylhet Limestone in the Bengal Basin, formed in the Eocene Epoch and known for its fossil content, is significant in understanding the reservoir characteristics. Despite exposure in Jaflong and Takerghat of the Bengal Basin, little research has been conducted on reservoir characteristics. This study aimed to comprehensively examine the Sylhet Limestone Formation, encompassing its crystalline upper and fossiliferous lower sections. It utilized field investigations and laboratory analyses to address the gaps of sporadic or insufficient earlier studies. A thorough examination of thin sections from the Sylhet Limestone, exposed in the Dauki River area, provides insights into the textural and mineralogical attributes and the presence of skeletal fossils within the limestone. Based on the analysis of thin sections, the limestones are categorized as Rudestones and Packstones. The porosity observed in the exposed rocks ranges from 5% to 12%, with most pores associated with interconnected fractures and joints. However, thin-section studies also indicate evidence of diagenetic recrystallization and calcite cementation. Hence, closely spaced, interconnected joints and fractures filled with diagenetic calcite might deteriorate the reservoir quality. Notably, this limestone exhibits fossilized specimens such as Nummulite, Discocyclina, Alveolina, Assilina, and Ostracoda, among others. The combination of the fossil assemblage, limestone texture, and composition strongly suggests that this limestone formation was deposited in a shallow marine environment with minimal sediment input under a warm and humid climate. The petrographic analysis of the limestones concludes that the upper portion of the formation is fine-grained while the lower part is coarse-grained. The Dhaka University Journal of Earth and Environmental Sciences, Vol. 12(2), 2023, P 119-137

Mesozoic crustal growth and recycling along the Southern margin of Eurasia: Magmatic rocks from the Sanandaj-Sirjan Zone of Iran

The Sanandaj-Sirjan Zone of western Iran is characterized by significant Mesozoic magmatic activity (from the early Jurassic to the late Cretaceous), with Cenozoic rocks being less common. Granitoids are prevalent in this zone, displaying A-, I-, and S-type geochemical characteristics. Despite several studies on these granitoids, there are still some questions regarding their sources– especially the S- and A-type granitoids – and the crustal growth and recycling in this zone, which require further studies using isotopes. This study focuses on the Aligoodarz, Kolah-Ghazi, and Golpayegan granitoids from the central SaSZ to better understand crustal growth and recycling in the Sanandaj-Sirjan Zone. The Aligoodarz and Kolah-Ghazi granitoids are identified as S-type granites, showing strong peraluminous signatures. Geochemical evidence suggests a mixture of melts from sub-crustal metasedimentary and meta-igneous sources, with factors such as source diversity and disequilibrium melting contributing to their formation. in contrast, the Golpayegan granitoids exhibit high total alkali (Na2O + K2O) contents and high field strength elements, indicating alkaline magmatic affinity. Their geochemical signatures suggest derivation from an enriched portion of an Indian MORB-like protolith. Golpayegan syenites share similarities with within-plate A1 granites, suggesting they are related to anorogenic silica-oversaturated granitoids. Golpayegan A-type granites probably originated from high-degree fractionation of alkali basic magmas. Golpayegan gabbros may be derived from a MORB or OIB-like mantle source, with characteristics indicating the presence of garnet in the mantle source and potentially deep melting from a mantle plume. Newly compiled zircon UPb data from the Sanandaj-Sirjan Zone show a complex magmatic chronology spanning from approximately 180 million years ago to the Eocene epoch, revealing peaks of magmatic activity at various intervals. Distinct phases of A-, I-, and S-type magmatism are identified, and synthesis of zircon UPb ages and HfO isotope data reveals the simultaneous occurrence of S-type and I-type magmas during specific periods. Furthermore, our study explores the tectonic settings underlying these magmatic processes and proposes scenarios involving subduction, continental rifting, and asthenospheric upwelling to elucidate the geological dynamics of the SaSZ.

Digital restoration of the snout of Khirtharia inflata (Raoellidae, Artiodactyla) from the middle Eocene of northwest Himalaya

Digital restoration of the snout of Khirtharia inflata (Raoellidae, Artiodactyla) from the middle Eocene of northwest Himalaya

The bivalve-bearing carbonate platform on the east Tethys during the middle Eocene and its response to the Tethys transgression

The Eocene was a typical greenhouse climate period in the history of the Earth, with a high global sea level and extensive carbonate deposits developed in Tethys. During the Eocene, a carbonate platform was deposited in the western Tarim Basin, which belongs to the easternmost Tethys. However, the details of this carbonate platform and its complications for paleoecology and paleoclimate are still unclear. This research focuses on the bivalve-bearing carbonates of the Kalataer Formation in the western Tarim Basin, and detailed analyses of microfacies, biostratigraphy, paleoecology, and sea-level change were carried out. The bivalves of the Kalataer Formation are dominated by the Ostrea (Turkostrea) strictiplicata, Ostrea (Turkostrea) cizancourti, Ostrea (Turkostrea) strictiplicata, Ostrea (Turkostrea) strictiplicata, and Sokolowia buhsii, and the age is middle Lutetian of the Eocene. The biota of the Kalataer Formation mainly lives in open shallow sea environments, with medium to low energy and rich nutrients and oxygen, representing a typical shallow marine ecosystem of the carbonate platform. Microfacies and paleoecology indicate a large marine transgression event occurred in the western Tarim Basin during the middle Eocene.

A new primate community from the earliest Oligocene of the Atlantic margin of Northwest Africa: Systematic, paleobiogeographic, and paleoenvironmental implications

We report a new Paleogene primate community discovered in the uppermost part of the Samlat Formation outcropping on the continental shore of the Rio de Oro, east of the Dakhla peninsula (in the south of Morocco, near the northern border of Mauritania). Fossils consist of isolated teeth, which were extracted by wet screening of estuarine sediments (DAK C2) dating from the earliest Oligocene (ca. 33.5 Ma). These dental remains testify to the presence of at least eight primate species, documenting distinct families, four of which are among the Anthropoidea (Oligopithecidae [Catopithecus aff. browni], Propliopithecidae [?Propliopithecus sp.], Parapithecidae [Abuqatrania cf. basiodontos], and Afrotarsiidae [Afrotarsius sp.]) and four in the Strepsirrhini (a Djebelemuridae [cf. ‘Anchomomys’ milleri], a Galagidae [Wadilemur cf. elegans], a possible lorisiform [Orogalago saintexuperyi gen. et sp. nov.], and a strepsirrhine of indeterminate affinities [Orolemur mermozi gen. et sp. nov.]). This record of various primates at Dakhla represents the first Oligocene primate community from Northwest Africa, especially from the Atlantic margin of that landmass. Considering primates plus rodents (especially hystricognaths), the taxonomic proximity at the generic (even specific) level between DAK C2 (Dakhla) and the famous Egyptian fossil-bearing localities of the Jebel Qatrani Formation (Fayum Depression), either dating from the latest Eocene (L-41) or from the early Oligocene, suggests the existence of an east–west ‘trans-North African’ environmental continuum during the latest Eocene–earliest Oligocene time interval. The particularly diverse mammal fauna from DAK C2, recorded within the time window of global climate deterioration characterizing the Eocene/Oligocene transition, suggests that this tropical region of northwest Africa was seemingly less affected, if at all, by the cooling and associated paleoenvironmental and biotic changes documented at that time or at least that the effects were delayed. The expected densely forested paleoenvironment bordering the western margin of North Africa at the beginning of the early Oligocene probably offered better tropical refugia than higher latitudes or more inland areas during the cooling episode.

Genomes of the Orestias pupfish from the Andean Altiplano shed light on their evolutionary history and phylogenetic relationships within Cyprinodontiformes

BackgroundTo unravel the evolutionary history of a complex group, a comprehensive reconstruction of its phylogenetic relationships is crucial. This requires meticulous taxon sampling and careful consideration of multiple characters to ensure a complete and accurate reconstruction. The phylogenetic position of the Orestias genus has been estimated partly on unavailable or incomplete information. As a consequence, it was assigned to the family Cyprindontidae, relating this Andean fish to other geographically distant genera distributed in the Mediterranean, Middle East and North and Central America. In this study, using complete genome sequencing, we aim to clarify the phylogenetic position of Orestias within the Cyprinodontiformes order.ResultsWe sequenced the genome of three Orestias species from the Andean Altiplano. Our analysis revealed that the small genome size in this genus (~ 0.7 Gb) was caused by a contraction in transposable element (TE) content, particularly in DNA elements and short interspersed nuclear elements (SINEs). Using predicted gene sequences, we generated a phylogenetic tree of Cyprinodontiformes using 902 orthologs extracted from all 32 available genomes as well as three outgroup species. We complemented this analysis with a phylogenetic reconstruction and time calibration considering 12 molecular markers (eight nuclear and four mitochondrial genes) and a stratified taxon sampling to consider 198 species of nearly all families and genera of this order. Overall, our results show that phylogenetic closeness is directly related to geographical distance. Importantly, we found that Orestias is not part of the Cyprinodontidae family, and that it is more closely related to the South American fish fauna, being the Fluviphylacidae the closest sister group.ConclusionsThe evolutionary history of the Orestias genus is linked to the South American ichthyofauna and it should no longer be considered a member of the Cyprinodontidae family. Instead, we submit that Orestias belongs to the Orestiidae family, as suggested by Freyhof et al. (2017), and that it is the sister group of the Fluviphylacidae family, distributed in the Amazonian and Orinoco basins. These two groups likely diverged during the Late Eocene concomitant with hydrogeological changes in the South American landscape.

Phylogenomics and biogeography of the small carpenter bees (Apidae: Xylocopinae: Ceratina)

Small carpenter bees in the genus Ceratina are behaviourally diverse, species-rich, and cosmopolitan, with over 370 species and a range including all continents except Antarctica. Here, we present the first comprehensive phylogeny of the genus based on ultraconserved element (UCE) phylogenomic data, covering a total of 185 ingroup specimens representing 22 of the 25 current subgenera. Our results support most recognized subgenera as natural groups, but we also highlight several groups in need of taxonomic revision – particularly the nominate subgenus Ceratina sensu stricto – and several clades that likely need to be described as new subgenera. In addition to phylogeny, we explore the evolutionary history of Ceratina through divergence time estimation and biogeographic reconstruction. Our findings suggest that Ceratinini split from its sister tribe Allodapini about 72 million years ago. The common ancestor of Ceratina emerged in the Afrotropical realm approximately 42 million years ago, near the Middle Eocene Climatic Optimum. Multiple subsequent dispersal events led to the present cosmopolitan distribution of Ceratina, with the majority of transitions occurring between the Afrotropics, Indomalaya, and the Palearctic. Additional movements also led to the arrival of Ceratina in Madagascar, Australasia, and a single colonization of the Americas. Dispersal events were asymmetrical overall, with temperate regions primarily acting as destinations for migrations from tropical source regions.

Low-Temperature Thermochronology Records the Convergence between the Anatolide–Tauride Block and the Arabian Platform along the Southeast Anatolian Orogenic Belt

SE Anatolia is witnessing the final stage of the Wilson Cycle, where a continental collision between the Tauride–Anatolide block and Arabian platform occurred, and a 1.5 km Eastern Tauride mountain chain formed. We present new low-temperature thermochronology (LTT) ages, including eight apatite fission track (AFT) and seven apatite and zircon U-Th-Sm/He (AHe, ZHe) ages, for the metamorphic rocks from the Nappe Zone of the Southeast Anatolian Orogenic Belt. The ZHe ages vary from 51.2 ± 0.7 Ma to 30.4 ± 0.6 Ma, the AFT ages range from 33.1 ± 1.6 Ma to 18.1 ± 0.9 Ma, and the AHe ages range from 23.6 ± 2.5 Ma to 6 ± 1.9 Ma. The LTT data show a continuous slow uplift of the region. However, the thermal modeling results suggest an Eocene and middle–late Miocene fast uplift of the region. Similar to our results, the LTT studies along the SAOB show that the vertical movements initiated during the Eocene period have continued in a steady-state regime to recent times. The Eocene epoch is identified by arc–back-arc setting in the region, whereas the Miocene epoch is marked by the continental collision. Within this tectonic framework, vertical movements on the overriding plate are controlled by both extensional and compressional tectonics. The LTT data obtained along the SAOB show fingerprints of thrust propagation from north to south.