Summary. A laboratory and theoretical study of the stability of conduits of buoyant fluid in a viscous shear flow has been conducted. The object of the study is to explain the formation of discrete islands in island chains such as the Hawaiian Emperor seamount chain, and to investigate a new method by which the variation of shear with depth in the mantle may be determined. The conduits were made by injecting oil into a more viscous oil of greater density. Initially a growing chamber of lower viscosity oil formed near the injector, but when the chamber got sufficiently large it rose as a buoyant spheroid. Behind this trailed a vertical cylindrical conduit through which fluid could continue to rise to the surface as long as the source continued. If the more viscous fluid was sheared laterally the conduit was gradually rotated to a more horizontal position. The diameter of the conduit increased with time due to a decreasing component of gravitational force along the axis of the conduit. When the conduit was tilted to more than 60 with the vertical, it began to go unstable by developing bumps which ultimately initiated a new chamber which rose to a new spot. In addition, if the Reynolds number of the conduit was greater than approximately ten, an axisymmetric wavy instability appeared in the walls of the conduit and the conduit had to be tilted less before a new chamber was initiated. If shear under the Pacific plate has to tilt buoyant mantle plumes to as much as 60 to form the relatively regular island chains associated with hot spots, most of the shear would be found in a zone with a vertical extent of less than 200 km.