Delayed germination of Douglas seeds results from the simultaneous effect of every seed components. The spermoderm at one and the same time slows down water and oxygen uptake. Embryo and endosperm are also responsible for this delayed germination owing to a certain exchange slowness between them. This process induces an embryonic dormancy the length of which depends on the seeds. Chilling treatments give rise to a fuller embryo development (after-ripening) and increase the inner exchange efficiency as also the water uptake through the spermoderm.
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