Variations In Elevation Research Articles

Effect of elevated temperature variations on the corrosion resistance of S31603 and SS2562 austenitic stainless steels in chloride-sulphate environments

Effect of environmental temperature variation at 308K, 318K, 333K and 353K on the corrosion resistance of S31603 and SS2562 austenitic stainless steels in 2.5M H2SO4/3.5% NaCl solution was studied by potentiodynamic polarization, potentiostatic measurement and optical microscopy. S31603 exhibited higher general corrosion resistance compared to SS2562 with values of 0.529 mm/y and 8.701 mm/y (308K and 353K) compared to 0.911 mm/y and 17.492 mm/y for SS2562. The corrosion potential of SS2562 shifted to cathodic values compared to S31603 which varies between cathodic and anodic values. At 308K, SS2562 exhibited metastable pitting activity at potential of 0.02 V. Passivation occurs at 0.09 V before breakdown of the protective oxide. Beyond 308K, passivation was absent on SS2562. S31603 exhibited unstable passivation behaviour at 308K and elevated temperatures. The protective oxide of S31603 formed at certain potentials before collapsing at all temperatures studied. Images of both steels showed localized corrosion deterioration at the grain boundaries. Larger micro-pits were visible on SS2562 compared to S31603 which had numerous shallow pits on its surface. At 353K, the extent morphological deterioration has advanced further on SS2562 compared to S31603 with the grain boundaries becoming more visible. The microstructural deterioration on S31603 at 308K and 353K is comparatively minimal.

Two-Dimensional Spatial Coherence for Ultrasonic DMAS Beamforming in Multi-Angle Plane-Wave Imaging

Ultrasonic multi-angle plane-wave (PW) coherent compounding relies on delay-and-sum (DAS) beamforming of two-dimensional (2D) echo matrix in both the dimensions PW transmit angle and receiving channel to construct each image pixel. Due to the characteristics of DAS beamforming, PW coherent compounding may suffer from high image clutter when the number of transmit angles is kept low for ultrafast image acquisition. Delay-multiply-and-sum (DMAS) beamforming exploits the spatial coherence of the receiving aperture to suppress clutter interference. Previous attempts to introduce DMAS beamforming into multi-angle PW imaging has been reported but only in either dimension of the 2D echo matrix. In this study, a novel DMAS operation is proposed to extract the 2D spatial coherence of echo matrix for further improvement of image quality. The proposed 2D-DMAS method relies on a flexibly tunable p value to manipulate the signal coherence in the beamforming output. For p = 2.0 as an example, simulation results indicate that 2D-DMAS outperforms other one-dimensional DMAS methods by at least 9.3 dB in terms of ghost-artifact suppression. Experimental results also show that 2D-DMAS provides the highest improvement in lateral resolution by 32% and in image contrast by 15.6 dB relative to conventional 2D-DAS beamforming. Nonetheless, since 2D-DMAS emphasizes signal coherence more than its one-dimensional DMAS counterparts, it suffers from the most elevated speckle variation and the granular pattern in the tissue background.

Forest structure determines spatial changes in avian communities along an elevational gradient in tropical Africa

AbstractAimTo test if tree species richness and forest structure drive spatial variation in avian communities along a tropical elevation gradient and to present information about the role of detailed forest parameters.LocationA 2,000‐m long elevational gradient of tropical forest on Mt. Cameroon, west‐central Africa.TaxonBirds and trees.MethodsWe performed bird censuses and vegetation mapping at the same plots across six forested sites at elevations of 350, 650, 1,100, 1,500, 1,850, and 2,200 m a.s.l., with 16 plots per elevation. We tested the effects of elevation, forest structure and tree diversity on the species richness, functional diversity and β‐diversity of birds (Bray–Curtis dissimilarity). We used conditional inference trees based on random forests (RF) to investigate these relationships across all elevation sites as well as within elevations.ResultsBoth tree and bird species richness declined monotonically with elevation. Vegetation structure correlated with elevation, and all vegetation attributes significantly differed among elevations. The RF explained 70% of the variance in avian species richness, with the most important predictors being elevation, proportion of dead trees, tree species richness and herb layer coverage. We found that elevation (and shrub layer) was a particularly important predictor of avian functional diversity. We identified no important predictor of bird species richness after standardization within elevations, and the proportion of dead trees was the sole important predictor of functional diversity. Within‐elevation β‐diversity in avian community composition was determined by the dissimilarity of the tree community and differences in leaf area index, solar radiation and spatial distance. The functional dissimilarity was best explained by leaf area index.Main conclusionsApart from elevation itself, spatial distance even within elevations correlated with compositional and functional variation among avian assemblages. Forest structural traits can have a significant influence on distribution of birds. Thus, gaps in the spatial distribution of species such as along elevations might be caused by fine‐scale recognition of suitable habitats.

Elevation, aspect, and local environment jointly determine diatom and macroinvertebrate diversity in the Cangshan Mountain, Southwest China

Abstract Freshwater ecosystems in high-mountain regions are subject to emerging threats such as global warming and expanding human activities. Stream diatoms and macroinvertebrates form an essential component of freshwater ecosystems in high-mountain regions. Although these organisms are sensitive to environmental changes, knowledge regarding their elevational diversity patterns remains limited. Opposite aspects (e.g., north vs south; west vs east) usually receive different amounts of solar radiation and precipitation, leading to distinct in-stream characteristics such as discharge, flow regime, and water temperature. Despite the suggested strong influence of aspect on biodiversity patterns in mountains, its effect on stream diatoms and macroinvertebrates has been largely overlooked. The aims of our study were to 1) investigate whether macroinvertebrate and diatom taxon richness follows the same pattern along an elevational gradient; 2) test the effect of aspect on the elevational diversity (i.e., taxon richness and assemblage dissimilarity) patterns of macroinvertebrate and diatom assemblages; and 3) examine the relative importance of elevation, aspect, and the local environment (e.g., in-stream physicochemical variables) in shaping macroinvertebrate and diatom assemblages. We investigated macroinvertebrate and diatom assemblages in six nearly parallel streams (three streams on the east aspect and three on the west) in the Hengduan Mountains region. We found that the taxon richness of both macroinvertebrates and diatoms showed a monotonic increase with elevation (1623–2905 m a.s.l.) when aspect was not accounted for. When aspect was taken into consideration, macroinvertebrate taxon richness still showed a monotonically increasing elevational pattern on both the east and west aspects, but with significantly different model slopes, while a monotonical pattern for diatoms only remained on the west aspect. In addition, taxon richness of macroinvertebrates may also follow a potential unimodal pattern. The distance-decay relationships followed the same patterns as taxon richness, suggesting that the influence of aspect on diatom and macroinvertebrate assemblages was not negligible. Our results suggested that the diversity of both diatoms and macroinvertebrates was jointly determined by elevation, aspect and local environment. Compared to macroinvertebrates, the diatom assemblages were more strongly influenced by the local environment. To gain a better understanding of the underlying mechanisms driving elevational patterns of stream biodiversity in high-mountain regions, comparative studies that involve multiple organisms, streams, and mountains across a large elevational range are needed.

Erosion Hazard Model Mapping for Orlu Zone, Imo State, Nigeria Using GIS

Soil erosion is a significant environmental issue of common concern in the world today. For the past five decades, the study area (Orlu Zone) in Imo State, Nigeria has been experiencing environmental degradation and loss of properties due to soil erosion. Impact of soil erosion on man and his environment is natural and as old as the time of Earths creation. It has been supported that a number of factors combined in contributing to soil erosion and its negative environmental consequences ranging from social, economic and property losses of which serious runoff and soil loss has become one of the main factors restraining local economic developments. The aim of this study is therefore, to produce an erosion hazard model map that identifies areas of high and low risks for erosion. Study identified seven susceptibilities factors (soil characteristics, variation in elevation, slope, land use, geology, population density and drainage density) of soil erosion, converted them to grid format and reclassified each factor into five potential after assigning erosion risk values based on acceptable hazard potential of each factor. Thereafter, weighed on Arc GIS version 10.2 software weighted overlay toolbox to produce erosion risk map showing Orlu Zone lands classified into various hazard potentials. Result showed that highest vulnerable land areas to soil erosion occurred in the Northern part of the study area while the lowest vulnerable areas occurred on the Southern counterpart.

A multi-objective optimization model for decision support in water reclamation system planning

Abstract Water reclamation and reuse not only reduce the negative impacts of wastewater to the environment, but also provide an alternative to withdrawal from natural water resources, forming a closed-loop water supply chain. The design of such a supply chain requires an appropriate sustainability assessment, which simultaneously accounts for economic, environmental, and social dimensions. In this study, a multi-objective optimization model was developed to minimize the costs and carbon footprint (as an environmental indicator), and maximize the value of resource recovery (representing a social benefit) of the water reclamation systems by locating the treatment facility, allocating the treatment capacity, selecting the treatment technology, and allocating customers (final reclaimed water users). The optimizer CPLEX 12.7 was applied and the Triangle Splitting Method was used for the optimization algorithm. The expansion of a water reclamation system in Hillsborough County, Florida was evaluated to illustrate the use of the model. The impacts of population density and elevation variation in the water service area on the model outputs were also investigated. Although the centralization of treatment facilities takes advantage of the economies of scale, the results revealed that simultaneous consideration of economic and environmental indicators favored decentralization of treatment facilities in the study area. This was mainly due to the significant decrease in water transfer requirements, especially in less populous areas. Moreover, the results revealed that contribution of population density to the optimal degree of decentralization of treatment facilities was significant.

Vertical zonality of the water cycle and the impact of land-use change on runoff in the Qingshui River Basin of Wutai Mountain, China

ABSTRACTHydrological processes in hilly watersheds are significantly affected by variations in elevation; however, the hydrological functions of different vertical vegetation belts, have rarely been reported. The distributed hydrological model WEP-L (Water and Energy transfer Process in Large river basins) was applied to analyse vertical variations in the hydrological processes of Qingshui River basin (QRB), Wutai Mountain (altitude: 3058 m a.s.l.), China. The results show that the highest ratio of evapotranspiration to precipitation occurs at 1800 m a.s.l. Below 1800 m, evapotranspiration is mainly controlled by precipitation, and in regions above1800 m it is controlled by energy. The runoff coefficients for different vertical vegetation belts may be ranked as follows: farmland > grassland > subalpine meadow > evergreen coniferous shrub forest > deciduous broad-leaved forest. Grassland is the largest runoff production area, contributing approximately 39.10% to the annual water yield of the QRB. The runoff from forested land decreased to a greater extent than the grassland runoff. Increasing forest cover may increase evapotranspiration and reduce runoff. These results are important, not only for further understanding of the hydrological mechanisms in this basin, but also for implementing the sustainable management of water resources and ecosystems in other mountainous regions.

The response of mixed-species bird flocks to anthropogenic disturbance and elevational variation in southwest China

Abstract Avian mixed-species flocks (MSFs) are an important example of species interactions threatened by the biodiversity crisis. They are found throughout the world in forested habitats but are generally reduced in size or frequency by human disturbance. In southern China, a unique MSF system is led by several species of closely- related fulvettas (Alcippe morrisonia, A. hueti, and A. davidi). Our objective was to understand how this system is distributed across elevational gradients, especially moving west into the Hengduan Mountains, and how it responds to human disturbance. We sampled leadership and composition of 375 MSFs over 2 yr in and surrounding 5 nature reserves in Guangxi Zhuang Autonomous Region and Yunnan Province, with transects placed in different land use types—protected forest, buffer areas, or agriculture—and ranging from 400 to 3,200 m. We also sampled birds outside of MSFs. We found MSFs led by fulvettas (A. fratercula and A. davidi) in forests across the region up to 2,900 m. Elevation was not a significant influence on MSF size or prevalence in models that also included land use. We found that MSFs were encountered at only one-third of the frequency in agriculture as in forest, and had strongly different composition and leadership. Although MSFs in buffer areas were more similar to those in forest, birds in buffer areas had lower flocking propensity, different flock leaders, and less complex social networks. In particular, buffer transects that were seeded pine had low numbers of fulvetta-led MSFs and forest specialists in MSFs. In the future, it is important to understand which vegetational characteristics allow MSFs, particularly fulvetta-led MSFs, to persist in buffer habitats.

Fifteen important questions in the spatial ecology of diatoms

Abstract Recently, studies about the spatial distribution of microbial species and communities have increased substantially. Freshwater diatoms are used as one of the most popular model taxa in such macroecological studies, but the most exciting patterns in the spatial ecology of diatoms have not been reviewed. Here, we identify 15 important questions about freshwater diatom ecology, review the major findings, and suggest novel research avenues. We found that diatoms do not support classical latitudinal diversity gradient, species–area relationship or Rapoport's rule. We found some, albeit inconsistent support for an elevational diversity gradient, Bergmann's rule and niche conservation. Diatoms do support a positive interspecific abundance–occupancy relationship like nearly all other taxa. With regards to metacommunity dynamics, diatom communities are typically structured by species sorting, but neutral dynamics or mass effects are also evident at large or small spatial scale, respectively. Taken together, this review suggests that diatoms follow some classical biogeographical patterns similarly to larger taxa, but patterns are typically weaker than for macro‐organisms. This may result because diatoms are typically relatively strongly controlled by local factors and often exhibit efficient stochastic dispersal processes both at small and large scales. More research is needed to build a more comprehensive view of diatom species–area relationships and latitudinal gradients in diversity, for example. In the future, well‐replicated macroecological field experiments, diatom studies with intercontinental or global coverage, trait‐based approaches, and DNA‐based species identification would be fruitful avenues to shed more light into the spatial ecology of diatoms.

Do Deltas Along the Crustal Dichotomy Boundary of Mars in the Gale Crater Region Record a Northern Ocean?

AbstractDeltaic deposits mapped along the Martian crustal dichotomy boundary scarp have been suggested to delineate an ancient ocean in the northern lowlands of Mars. Using recently acquired orbital data, we have expanded the dichotomy delta inventory and performed an updated analysis of delta front elevations, a proxy for paleo‐water levels. Our analysis focused near Gale crater, home of the Curiosity rover. We found that delta front elevations vary by approximately 2,400 m, but these elevation variations do not correspond to modeled deformation from true polar wander or Tharsis. Locally, delta front elevations vary by ≤60 m, and using present‐day topography, they correspond to distinct enclosed basins. We infer that these deltas formed in paleo‐lakes up to approximately 13,000 km2 and approximately 0.4 km deep, perhaps coeval with paleo‐lakes in Gale. Our results suggest that a northern ocean is not needed to explain the deltaic deposits in the Gale crater region.

A 2D vertical model for simulating surface and subsurface flows using finite element–finite volume methods

AbstractA numerical model is presented for simulation of hydrodynamics of a 2D vertical free surface domain consisting of an arbitrary partitioned porous and non-porous area. To this end, modified Navier–Stokes equations are considered which could be applied in surface water and in subsurface flows, simultaneously. A wide range of Reynolds number has been considered, from which non-Darcy effects have also been taken into account. A fractional step method has been used in the time discretization procedure, where the convection and diffusion terms are separated from the pressure term while solving the momentum equations. To include the variation of surface elevation in computation, the domain has been divided into two parts, namely, ‘interior subdomain’, which never gets dry during the simulation period, covered by fixed unstructured triangular grids and ‘top layer’ with only a one layer structured grid, the position of which varies with the water surface. The validation of the proposed model has been achieved by comparison of its results with both theoretical and experimental data reported in the literature.

Allopatric populations of the invasive larch casebearer differ in cold tolerance and phenology

1. Traits of non‐native insect herbivores may vary spatially due to local genetic differences, rapid post‐introduction evolution, and/or novel host plant associations.2. Populations of larch casebearer, Coleophora laricella Hübner, originally from Europe have likely been isolated for > 60 years in North America on eastern larch, Larix laricina (Du Roi) K. Koch, and western larch, Larix occidentalis Nutt.3. This study investigated cold tolerance and phenology of larvae collected from eastern larch in Minnesota, and western larch in Oregon, Idaho, and Montana, U.S.A.4. Mean supercooling points of larvae from Minnesota were up to 10 °C lower than supercooling points of larvae from Oregon, Idaho, and Montana.5. At ambient environmental conditions in spring, overwintering larvae from Minnesota required a mean (± SE) of 172 ± 19 degree‐days above 5 °C to break winter quiescence and actively wander, significantly more than required by larvae from Oregon (66 ± 4), Idaho (64 ± 1), and Montana (60 ± 2).6. Across all assays and despite substantial latitudinal and elevational variation among western larch sites, no significant differences in any traits were detected among larvae collected from western larch.7. Spatial variation in cold tolerance and phenological traits of larch casebearer may be attributable to insect genetic differences and/or host plant effects, but exact mechanisms remain unknown. Differences in thermal biology between regions may result in disparate effects of climate change on insect populations and should be accounted for when forecasting insect dynamics across large spatial scales.

Evaluating earthquake-related ground failure mapping by combined traditional and modern methods

Abstract. A wide range of ground failure such as earthquake faulting (surface rupture), landslides, and liquefaction occur after a large earthquake. In this study, in order to rapidly determine the distribution of failure over a wide area after an earthquake, we combined traditional methods such as aerial photo interpretation and modern methods such as unmanned aerial vehicle (UAV) or interferometric synthetic aperture radar (InSAR) techniques. Moreover, elevation variations obtained using DEMs and the structure from motion and multi-view stereo (SfM-MVS) technique were employed to understand local ground deformation factors, such as reclaimed valley deformation. Using ortho-mosaic images, surface fissures caused by the 2016 Kumamoto Earthquake in Japan were rapidly interpreted and mapped, which enabled early interpretation of the ground failure situation. Furthermore, surface displacement properties extracted from SAR interferograms allowed for more advanced earthquake fault detection; surface displacement associated with liquefaction was also identified from SAR interferograms. In addition, InSAR was used to detect reclaimed valley deformation. Comparing this with the reclaimed valley distribution map created by the SfM-MVS technique improved our understanding of this phenomenon. However, many of these techniques require large amounts of manpower and time and can be influenced by differences in analyst skill level. In future, the development of mechanically automated ground failure identification will improve earthquake disaster responses.

Increasing phylogenetic stochasticity at high elevations on summits across a remote North American wilderness.

At the intersection of ecology and evolutionary biology, community phylogenetics can provide insights into overarching biodiversity patterns, particularly in remote and understudied ecosystems. To understand community assembly of the high alpine flora in the Sawtooth National Forest, USA, we analyzed phylogenetic structure within and between nine summit communities. We used high-throughput sequencing to supplement existing data and infer a nearly completely sampled community phylogeny of the alpine vascular flora. We calculated mean nearest taxon distance (MNTD) and mean pairwise distance (MPD) to quantify phylogenetic divergence within summits, and assessed whether maximum elevation explains phylogenetic structure. To evaluate similarities between summits, we quantified phylogenetic turnover, taking into consideration microhabitats (talus vs. meadows). We found different patterns of community phylogenetic structure within the six most species-rich orders, but across all vascular plants phylogenetic structure was largely not different from random. There was a significant negative correlation between elevation and tree-wide phylogenetic diversity (MPD) within summits: overdispersion degraded as elevation increased. Between summits, we found high phylogenetic turnover driven by greater niche heterogeneity on summits with alpine meadows. Our results provide further evidence that stochastic processes may also play an important role in the assembly of vascular plant communities in high alpine habitats at regional scales. However, order-specific patterns suggest that adaptations are still important for assembly of specific sectors of the plant tree of life. Further studies quantifying functional diversity will be important in disentangling the interplay of eco-evolutionary processes that likely shape broad community phylogenetic patterns in extreme environments.

Regional terrain complexity evaluation based on GIS and K-means clustering model: a case study of Ningdu County, China

In order to accurately quantify the terrain complexity, a simple and accurate terrain complexity assessment (TCA) model is proposed. Taking Ningdu county in Jiangxi Province of China as an example, firstly, six terrain factors (named slope, topographic relief degree, surface cutting depth, surface roughness, elevation variation coefficient and topographic factors) of Ningdu county are extracted based on the Digital Elevation Model (DEM) with 30 m resolution and ARCGIS 10.2 software. Secondly, terrain complexity indexes of Ningdu County are obtained using k-means clustering. Results show that a current and effective spatial distribution characteristic of topographic complexity in Ningdu county is produced, and the very low and low terrain complexity indexes account for 33.28%, 28.35% respectively. The terrain complexity can be evaluated effectively by k-means clustering model. The terrain complexity can be provided for environmental protection and land use planning.

Allopatric divergence drives the genetic structuring of an endangered alpine endemic lizard with a sky‐island distribution

AbstractAnthropogenic climate change is causing a world‐wide reduction of alpine habitat, leaving many high‐elevation species restricted to sky‐islands and vulnerable to extinction. Understanding the genetic parameters of these populations provides key insight into species diversity, dispersal capacity and vulnerability to disturbance. We examined the impact of past climatic variation on a threatened alpine endemic lizard, the Guthega skink, Liopholis guthega. We analysed SNP and mtDNA data to determine the population structure and phylogeny within this species, providing an understanding of the species’ relatedness, dispersal and viability. We identified significant genetic structure, with the split between populations in Koscuiszko National Park, New South Wales (NSW) and the Bogong High Plains, Victoria (VIC) consistent with Plio‐Pleistocene divergence. However, we also detected evidence of possible historical introgressive hybridization between some NSW populations and the VIC populations. Marked within‐site population structure and significant population differentiation among sites within each state were found, indicating a limited dispersal capacity. Higher levels of genetic diversity within NSW support the correlation between elevation and diversity and implicate Kosciuszko National Park as a historic refugia. Low contemporary habitat availability, little to no capacity for elevational progression and low genetic diversity, particularly in VIC, leaves L. guthega highly vulnerable to threatening processes associated with climate change. Conservation management should consider genetic rescue as a potential method to enhance genetic diversity across this species’ range.

Seasonal near-surface air temperature dependence on elevation and geographical coordinates for Pakistan

This study evaluates the monthly, seasonal, and annual characteristics of near-surface air temperature as a function of both elevation and geographical coordinates over Pakistan, using a climatic data set from 53 observation stations ranging between 5 and 2317 m asl and stepwise multiple regression. Forcing processes for temperature gradient (TG) variation were determined by analyzing its relationship with moisture flux (qv) and gradients of rainfall and saturation vapor pressure (es). The bi-model pattern of TGE (temperature gradient with topographic elevation), which earlier studies (TAAC 113(3):671-682, TAAC 132:1129-1141) from the southern slopes of the Himalayas have observed, is also consistent in this study region. However, the forcing strengths, mechanisms, and processes for the monthly variations in TGs magnitude and their diurnal range are somewhat different. The annual cycle of TGE in this study is inversely (positively) associated with the rainfall (es) gradient. Monsoon and orographic controls for TGE are stronger in summer, especially in the day, while the influences of inversions and mountainous topoclimate are higher during the non-monsoon period, in particular in winter and at night. Mountain barrier effects are higher in summer, which considerably hamper the relationship between temperature and geographical coordinates and rainfall and elevations. Steeper values of mean TGEs in the non-monsoon period are the result of strong dry convectional cooling at higher elevations due to high thermal forcing effects at lower elevations. Sensible heating reduction due to westerly-driven precipitation and cloud cover in daytime in the northern mountainous region (or higher latitudes) further strengthens the TGE value in this period. This effect is weaker in the surrounding regions; therefore, the TGE magnitude in this study is expected to be higher, especially in the pre-monsoon season. Gradients are more shallow in winter (excluding latitude), but values are greater than those in summer, due to distinct variations in moisture and cloud amount, i.e., lower in winter and higher in summer, particularly at lower elevations. Observed distinct differences in TG diurnal range between elevation (high) and latitude (low) in this study are due to a wide variation in temperature, topography, and elevations within the same latitudinal belts. In this study, the lowest gradient values for elevation are observed in August and the highest in May, 1 month later than observed in adjacent mountain regions due to the late arrival of monsoon moisture in summer and intense thermal forcing effects following the wet early pre-monsoon months. Distinct variability of gradient magnitudes between the northern mountainous region and the southern flat terrain is due to differences in topoclimates, moisture amounts, elevations, and geographical coordinates.

Dynamics of Selected Soil Quality Indicators in Response to Land Use/Cover and Elevation Variations in Wanka Watershed, Northwestern Ethiopian Highlands

Abstract Soil quality (SQ) dynamics assessment vis-à-vis land use/land cover (LULC) and elevation variations in Ethiopia is desirable as elevation impact on land use is highly pronounced. This study examined SQ indicators dynamics across LULC and elevation variations. For this, surface soil samples (0−20 cm) were collected from the recognized LULC categories of different elevations in Wanka watershed, northwestern Ethiopian highlands. Both disturbed and undisturbed soil samples that were taken from three adjacent LULC (natural forest, grazing and cultivated lands) and elevation (2238–2300, 2400–2600, and 2700–2800) classes analysed for the selected physico-chemical SQ indicators. Two-way ANOVA, Tukey’s multiple comparison test and SQ deterioration index were computed. The impact of LULC and elevation was found significant on key SQ indicators. In cultivated and grazing lands, soil organic matter (SOM) and soil nutrients like total nitrogen declined significantly (p < 0.01). Conversely, bulk density increased significantly (p < 0.01). The divalent basic cations (Ca2+ and Mg2+), cation exchange capacity and pH significantly (p < 0.01) decreased in upper elevation. Synergetic effect of LULC and elevation variations was found significant (p < 0.01) on SOM, total nitrogen, available phosphorus, water content at field capacity and soil particle distribution (silt and clay). Thus, elevation specific land management strategies that improve these SQ indicators need to be emphasized.

Elevational and microclimatic drivers of thermal tolerance in Andean Pristimantis frogs

AbstractAimWe analysed elevational and microclimatic drivers of thermal tolerance diversity in a tropical mountain frog clade to test three macrophysiological predictions: less spatial variation in upper than lower thermal limits (Bretts’ heat‐invariant hypothesis); narrower thermal tolerance ranges in habitats with less variation in temperature (Janzen's climatic variability hypothesis); and higher level of heat impacts at lower elevations.LocationForest and open habitats through a 4,230‐m elevational gradient across the tropical Andes of Ecuador.MethodWe examined variability in critical thermal limits (CTmax and CTmin) and thermal breadth (TB; CTmax–CTmin) in 21 species of Pristimantis frogs. Additionally, we monitored maximum and minimum temperatures at the local scale (tmax, tmin) and estimated vulnerability to acute thermal stress from heat (CTmax–tmax) and cold (tmin–CTmin), by partitioning thermal diversity into elevational and microclimatic variation.ResultsOur results were consistent with Brett's hypothesis: elevation promotes more variation in CTmin and tmin than in CTmax and tmax. Frogs inhabiting thermally variable open habitats have higher CTmax and tmax and greater TBs than species restricted to forest habitats, which show less climatic overlap across the elevational gradient (Janzen's hypothesis). Vulnerability to heat stress was higher in open than forest habitats and did not vary with elevation.Main conclusionsWe suggest a mechanistic explanation of thermal tolerance diversity in elevational gradients by including microclimatic thermal variation. We propose that the unfeasibility to buffer minimum temperatures locally may explain the rapid increase in cold tolerance (lower CTmin) with elevation. In contrast, the relative invariability in heat tolerance (CTmax) with elevation may revolve around the organisms’ habitat selection of open‐ and canopy‐buffered habitats. Secondly, on the basis of microclimatic estimates, lowland and upland species may be equally vulnerable to temperature increase, which is contrary to the pattern inferred from regional interpolated climate estimators.

Increased variation in numbers of presacral vertebrae in suspensory mammals.

Restricted variation in numbers of presacral vertebrae in mammals is a classic example of evolutionary stasis. Cervical number is nearly invariable in most mammals, and numbers of thoracolumbar vertebrae are also highly conserved. A recent hypothesis posits that stasis in mammalian presacral count is due to stabilizing selection against the production of incomplete homeotic transformations at the lumbo-sacral border in fast-running mammals, while slower, ambulatory mammals more readily tolerate intermediate lumbar/sacral vertebrae. We test hypotheses of variation in presacral numbers of vertebrae based on running speed, positional behaviour and vertebral contribution to locomotion. We find support for the hypothesis that selection against changes in presacral vertebral number led to stasis in mammals that rely on dorsomobility of the spine during running and leaping, but our results are independent of running speed per se. Instead, we find that mammals adapted to dorsostability of the spine, such as those that engage in suspensory behaviour, demonstrate elevated variation in numbers of presacral vertebrae compared to dorsomobile mammals. We suggest that the evolution of dorsostability and reduced reliance on flexion and extension of the spine allowed for increased variation in numbers of presacral vertebrae, leading to departures from an otherwise stable evolutionary pattern.