Understanding the air velocity profile over random and three-dimensional wind-waves is crucial for evaluating momentum and energy transfer between air and water. Unfortunately, determining accurate air velocity profiles in field conditions is nearly impossible. It is well accepted, though, that the airflow above water waves has a logarithmic profile usually expressed in terms of the effective roughness parameter. However, this parameter cannot be evaluated directly from the wave measurements at the water surface. For young wind waves, it was recently shown that the airflow over the waves maintains wall similarity. In this case, the airflow vertical velocity distribution can be described by the Nikuradse fully rough logarithmic profile, using the root mean square value of the water surface elevation as the equivalent sand grain roughness height. The existence of mean current in water in either co- or counter-wind direction may significantly modify the wave field, the vertical wind-velocity profile and thus the momentum and energy transfer from air to water. The effect of the water current on the spatially developing boundary layer over young wind waves and the wall similarity is examined. Combined laboratory measurements at several fetches of finely resolved mean air velocity profile above the water surface and of the characteristics of the wind-wave field are performed at multiple wind-forcing and mean water current conditions. The shear stress at the air–water interface estimated using two independent approaches is weakly dependent on current, while the resultant wave field differs significantly.