Eastern Baltic Region Research Articles

Performance variation of lodgepole pine provenances in Latvia

Lodgepole pine ( var. Engelm.) has been extensively introduced in Scandinavia on less productive sites. Under a changing climate, it also has a high potential in the eastern Baltic region; still, its performance there has scarcely been reported. This study investigated the performance of 36 Canadian provenances in 14 trials in western Latvia. Tree dimensions showed notable provenance and provenance-by-environment variation, implying that local selection by provenance can be applied for improved yield. Southern provenances showed the best height growth, while southwestern (more oceanic) provenances excelled in diameter growth. Most of the quality traits were affected by provenance or provenance-by-environment interaction, yet the variation was lower than for the growth traits.Pinus contortalatifolia

Зооморфные и антропоморфные пряжки и контакты Восточной Прибалтики с Византией в эпоху переселения народов

Зооморфные и антропоморфные пряжки и контакты Восточной Прибалтики с Византией в эпоху переселения народов

Evaluation and comparison of the genetic structure of Bunias orientalis populations in their native range and two non-native ranges

We studied the invasive warty cabbage Bunias orientalis (Brassicaceae) in three geographically distinct areas. Using inter-simple sequence repeat fingerprinting, we analyzed warty cabbages, including non-native populations, from the eastern Baltic and western Siberian regions and native populations from southwestern Russia. The eastern Baltic region and western Siberia represent the two opposite directions of B. orientalis spread in climatically different zones. The genetic structures of the native and non-native B. orientalis populations were assessed through analysis of molecular variance (AMOVA) and the Bayesian clustering method and by determining the main measures of genetic diversity. AMOVA revealed considerable population differentiation in both the native and invasive ranges. Our results did not indicate a decrease in genetic diversity in the non-native populations of B. orientalis. Similar measures of genetic diversity and genetic structure were determined in the invasive populations in two geographically and ecologically distinct, non-native regions located in Europe and Asia. In both of these regions, higher genetic diversity was detected in the non-native populations than in the native region populations, which may be due to multiple introductions. However, Bayesian clustering analysis revealed slightly different sources of invasive populations in the two non-native regions. Genetic diversity patterns revealed the lack of isolation by distance between populations and confirmed the influence of anthropogenic factors on the spread of B. orientalis. The significance of native populations as germplasm resources for breeding is discussed.

Atmospheric teleconnections between the Arctic and the eastern Baltic Sea regions

Abstract. The teleconnections between meteorological parameters of the Arctic and the eastern Baltic Sea regions were analysed based on the NCEP-CFSR and ERA-Interim reanalysis data for 1979–2015. The eastern Baltic Sea region was characterised by meteorological values at a testing point (TP) in southern Estonia (58° N, 26° E). Temperature at the 1000 hPa level at the TP have a strong negative correlation with the Greenland sector (the region between 55–80° N and 20–80° W) during all seasons except summer. Significant teleconnections are present in temperature profiles from 1000 to 500 hPa. The strongest teleconnections between the same parameter at the eastern Baltic Sea region and the Arctic are found in winter, but they are clearly affected by the Arctic Oscillation (AO) index. After removal of the AO index variability, correlations in winter were below ±0.5, while in other seasons there remained regions with strong (|R| > 0.5, p < 0.002) correlations. Strong correlations (|R| > 0.5) are also present between different climate variables (sea-level pressure, specific humidity, wind speed) at the TP and different regions of the Arctic. These teleconnections cannot be explained solely with the variability of circulation indices. The positive temperature anomaly of mild winter at the Greenland sector shifts towards east during the next seasons, reaching the Baltic Sea region in summer. This evolution is present at 60 and 65° N but is missing at higher latitudes. The most permanent lagged correlations in 1000 hPa temperature reveal that the temperature in summer at the TP is strongly predestined by temperature in the Greenland sector in the previous spring and winter.

Drought identification in the eastern Baltic region using NDVI

Abstract. Droughts are phenomena that affect large areas. Remote sensing data covering large territories can be used to assess the impact and extent of droughts. Drought effect on vegetation was determined using the normalized difference vegetation index (NDVI) and Vegetation Condition Index (VCI) in the eastern Baltic Sea region located between 53–60° N and 20–30° E. The effect of precipitation deficit on vegetation in arable land and broadleaved and coniferous forest was analysed using the Standardized Precipitation Index (SPI) calculated for 1- to 9-month timescales. Vegetation has strong seasonality in the analysed area. The beginning and the end of the vegetation season depends on the distance from the Baltic Sea, which affects temperature and precipitation patterns. The vegetation season in the southeastern part of the region is 5–6 weeks longer than in the northwestern part. The early spring air temperature, snowmelt water storage in the soil and precipitation have the largest influence on the NDVI values in the first half of the active growing season. Precipitation deficit in the first part of the vegetation season only has a significant impact on the vegetation on arable land. The vegetation in the forests is less sensitive to the moisture deficit. Correlation between VCI and the same month SPI1 is usually negative in the study area. It means that wetter conditions lead to lower VCI values, while the correlation is usually positive between the VCI and the SPI of the previous month. With a longer SPI scale the correlation gradually shifts towards the positive coefficients. The positive correlation between 3- and 6-month SPI and VCI was observed on the arable land and in both types of forests in the second half of vegetation season. The precipitation deficit is only one of the vegetation condition drivers and NDVI cannot be used universally to identify droughts, but it may be applied to better assess the effect of droughts on vegetation in the eastern Baltic Sea region.

A Review of the Earliest Evidence of Agriculture in Lithuania and the Earliest Direct AMS Date on Cereal

Current knowledge of the beginnings of crop cultivation in Lithuania is based mainly on Cerealia-type pollen data supplemented by other indirect evidence such as agricultural tools. We argue that these records, predating carbonized remains of cultivated plants, are not substantial enough indicators of the early stages of agriculture in Lithuania. Here, we demonstrate that the macroremains of cultural plants that were previously reported from two Neolithic settlements in Lithuania were either mistakenly identified as domestic crops or incorrectly ascribed to the Neolithic period due to movement through the stratigraphic sequence and the absence of direct dating of cereal grains. Furthermore, we present a charred Hordeum vulgare grain from the Bronze Age settlement of Kvietiniai in western Lithuania. It was AMS-dated to 1392–1123 cal bc, and at present represents the earliest definite evidence for a crop in the eastern Baltic region. We conclude that, presently, there are no grounds to suggest that crop cultivation took place in Lithuania during the Neolithic.

Rimutė Rimantienė: Founder of the School of Stone Age Explorations in Lithuania and the Eastern Baltic Region

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Comparison of input data with different spatial resolution in landscape pattern analysis – A case study from northern Latvia

A suitable spatial scale needs to be selected in geographical and landscape ecological research, and this requires great consideration as different scales have profound effect on derived landscape spatial patterns. Numerous studies have investigated the effects of different scales on landscape metrics using simulated patterns, but few have been conducted to compare different data sources with variable scale for regional- and landscape-scale assessments. Possibly this has occurred because researchers have been prone to use the best available source, a well-known standard, and easiest to use. This study was conducted to assess the impact of input data resolution on values of landscape pattern metrics in four landscapes at scales 1:10 000, 1:50 000 and 1:100 000. The aim was to determine the applicability of three data sources for thematic models in landscape pattern analyses in the Eastern Baltic region. We found that the utility of CORINE Land Cover data for comprehensive structural assessment in mosaic-type landscapes was very limited, as the level of cartographic generalization excluded many small and linear landscape structure elements with potentially high importance for landscape functioning, such as habitat continuity. We also found that actual area harvested using clearcuts was considerably higher than shown in CORINE data, due to clearcuts size being much smaller than the minimum mapping unit. In the light of this, we suggest using data with spatial resolution corresponding to a cartographic scale of at least 1:50 000, in cases when spatial patches have size up to 25 ha.

Exotic aphid species Brachycaudus divaricatae in Central Europe: Distribution, host specificity and molecular diversity

AbstractAphid species

Palaeoseismic deformations in the Eastern Baltic region (Kaliningrad District of Russia)

Palaeoseismic deformations in the Eastern Baltic region (Kaliningrad District of Russia)

Pseudoscorpions (Arachnida: Pseudoscorpiones) in Estonia: new records and an annotated checklist

The current knowledge of pseudoscorpions in Estonia is summarized, based on specimens preserved in various Estonian collections and new material collected from 1993 to 2015. Altogether 474 studied specimens belonged to 14 species, including nine species recorded from Estonia for the first time. Supplemented by Apocheiridium rossicum, listed by literature data only, the number of known Estonian pseudoscorpions is now set at 15. The findings of Mesochelifer ressli are the northernmost known for the species. Some ecological and phenological observations were made on the most common species, along with notes on the habitat preferences of other species and the use of different trapping methods. The composition of the known Estonian fauna is compared with the published records from the neighbouring countries, revealing a highest similarity to the Finnish fauna.The possible occurrence of other species in Estonia and the latitudinal species diversity gradient in the Eastern Baltic region are shortly discussed.

Genetic diversity of Great Cormorants Phalacrocorax carbo in the eastern Baltic region

A rapid expansion of the continental subspecies of the Great Cormorant (Phalacrocorax carbo sinensis) has been recorded in Europe since the 1980s. Evaluation of genetic variability of the Great Cormorant using molecular markers is necessary for investigation of the mechanisms of formation of the newly established breeding populations in the eastern Baltic region (in the Kaliningrad region of Russia and in Lithuania). The samples for molecular investigation were collected in the largest breeding colonies of Great Cormorants located on the coast of the Curonian Lagoon of the Baltic Sea and in their smaller, later formed breeding colonies located in the continental part of Lithuania. After sequencing and alignment of partial fragments of mtDNA control region, 21 different haplotypes, including 8 new haplotypes never identified before, and 13 haplotypes described earlier were found. They were distributed with different frequency in different sampling sites. The haplotype network constructed using 342 bp D-loop sequences identified during the current study and all available sequences of Great Cormorants deposited in GenBank by previous investigators revealed haplotypes attributed to subspecies P. c. sinensis being distinguished from haplotypes of subspecies P. c. carbo in the separate part of haplotype network. The newly described haplotypes did not form a phylogenetically uniform group indicating possible colonization of the Kaliningrad region and the continental part of Lithuania by individuals descending from the largest breeding colonies spread at the coast zone of the Baltic Sea. A high level of genetic population diversity in different breeding colonies recorded in the Kaliningrad region and in eastern Lithuania confirms the formation of a highly variable and well-adapted population of the Great Cormorant participating in the process of colonization of new breeding areas in the Baltic region.

The beginnings of rye (Secale cereale) cultivation in the East Baltics

In this paper we review the earliest Secale (rye) records, both pollen and macroremains, from the eastern Baltic region (northeast Poland, Lithuania, Latvia, Estonia and Finland) in order to understand the processes evident in the beginnings of rye cultivation. By taking note of the abundance of recovered Secale grains and pollen in relation to other cereals found in published archaeobotanical data, we try to understand when rye began to be used regionally as a staple food. The clarification of the timing of Secale in the East Baltic, that had social and economic implications, is of particular importance. In this publication we also present a radiocarbon date directly derived from dating a Secale grain from a Roman period hillfort in Lithuania. The date obtained constitutes the earliest record of Secale in the eastern Baltic region, showing that rye cultivation traditions and concomitant innovations in agricultural technology started there much earlier than previously suggested.

Deconstructing the concept of Subneolithic farming in the southeastern Baltic

The paper presents a critical review of the zooarchaeological, macrobotanical, palynological and archaeological data from Lithuania and their previous interpretations, which formerly served as the basis for the concept of development of pre-Neolithic or Subneolithic low intensity farming and/or livestock breeding in the eastern Baltic region. Moreover, it presents the first direct AMS dates from the crop remains and domestic animal bones discovered in Lithuanian Subneolithic and Neolithic settlements. An investigation proved that most of, or possibly all, the early farming “evidence” came from the wrong identification of the plant and animal species and incorrect dating of crop remains and domestic animal bones. The errors of dating were caused by the fresh water reservoir effect being ignored when dating the bulk lacustrine sediment samples, by the failure to evaluate the impact of the palimpsest and bioturbation phenomena on the formation of an archaeological layer, and by insufficient attention to stratigraphy and spatial documentation of the finds during very extensive archaeological excavations in the second half of the 20th century. To date, no credible evidence is available in Lithuania that domestic animals had been kept and crops grown before the Neolithic Globular Amphora and Corded Ware cultures in 3200/2700 cal bc. However, this does not mean such evidence may not appear in the future, provided direct AMS dating of animal and crop residues from Subneolithic contexts continues, and systematic macrobotanical studies finally start not only in the lake settlement and fishing sites, but also in higher altitude areas.

King Wenceslas’ relations with the Teutonic Order in light of the dispute over the Archbishopric of Riga (1392–1397)

ABSTRACTKing Wenceslas of the Romans (1378–1400) and of Bohemia (1378–1419) supported the Teutonic Knights, but his involvement in the Eastern Baltic region during the 1390s turned him into an enemy of the Knights. This paper argues that the change in Wenceslas’ policy was solely in the interests of his courtier Duke Swantibor of Pomerania-Stettin, who sought to establish his son as Archbishop of Riga. Wenceslas’ support for Swantibor owed to the traditions of courtly favor and instability in Bohemia, not out of any personal grudge against the Knights, as was claimed in earlier historiography.

GIS‐based multiproxy coastline reconstruction of the eastern Gulf of Riga, Baltic Sea, during the Stone Age

We present early to mid‐Holocene palaeocoastline reconstructions for the Tolkuse‐Rannametsa area on the eastern coast of the Gulf of Riga. Radiocarbon‐dated sediment stratigraphies and diatom analyses, supported by ground‐penetrating radar data and GIS‐based modelling, were used in the palaeogeographical reconstructions to strip back younger sediments and landforms from the LiDAR‐derived digital elevation model. The reconstructions show an open coast environment during the Ancylus Lake stage and the formation of a peatland after the lake drainage at about 9000–8600 cal. a BP. The waters of the Litorina Sea flooded the study area between 8200 and 7700 cal. a BP, forming a 25‐km‐long and up to 5.5‐m‐deep lagoon with two connections to the sea. This brackish water lagoon existed in the area for about 4000 years and terminated around 3800–3500 cal. a BP as a result of isostatic uplift and growth of the barrier spit. The reconstructions illustrate the relationships between the lagoonal system and three Mesolithic and Neolithic settlement sites found in the study area. A comparative analysis which besides palaeogeographical and ecological data included a data set with more than 100 Stone Age sites from the eastern Baltic region, was used to propose the most suitable locations for coastal settlements in the Tolkuse‐Rannametsa area. However, some of these areas are presently covered by a peat layer that is several metres thick, which may explain the relative scarcity of the archaeological finds on the shores of the palaeo‐lagoon. The proposed multiproxy approach has the capability to delimit the areas that would have been favourable for prehistoric habitation, but might not be observable in the present‐day topography. In return, this increases the likelihood of locating any well‐preserved preserved settlement sites and artefacts that are buried under the protective sediment layer.

The new Late Bronze Age hoard find from Kobbelbude (former Eastern Prussia, district Fischhausen) and the first results of its archaeometallurgical investigations

This article deals with a new discovered Late Bronze Age hoard find from former Kobbelbude (district Fischhausen, Eastern Prussia, now district Kaliningrad, Russian Federation). The hoard find is of special importance because it is the first time that metallurgical investigations were carried out on its artefacts. Archaeometallurgical analysis is, apart from few exceptions, still a desideratum of research in the eastern Baltic region. Also, for the first time, lead isotope ratios were determined. The results of the isotopic composition of the lead show a first hint towards a possible Eastern Alpine origin of the used ores. Former archaeometallurgical research of Bronze Age metal finds from the eastern Baltic region suggested a more abstract assumption about the origin of the ores from Middle Europe. Moreover, with the analysis of the chemical composition of the artefacts from Kobbelbude, a material classification to the type “dilute” fahlore copper with nickel, which we know from Late Bronze Age Middle Europe, was drawn. The results presented in this paper are a first step in the field of archaeometallurgical research in the eastern Baltic region and open a new perspective on Bronze Age metallurgy in relation to cultural history.

Palaeoenvironmental evidence for the impact of the crusades on the local and regional environment of medieval (13th–16th century) northern Latvia, eastern Baltic

This paper evaluates the impact of the crusades on the landscape and environment of northern Latvia between the 13th–16th centuries (medieval Livonia). The crusades replaced tribal societies in the eastern Baltic with a religious state (Ordenstaat) run by the military orders and their allies, accompanied by significant social, cultural and economic developments. These changes have previously received little consideration in palaeoenvironmental studies of past land use in the eastern Baltic region, but are fundamental to understanding the development and expansion of a European Christian identity. Sediment cores from Lake Trikāta, located adjacent to a medieval castle and settlement, were studied using pollen, macrofossils, loss-on-ignition and magnetic susceptibility. Our results show that despite continuous agricultural land use from 500 BC, the local landscape was still densely wooded until the start of the crusades in AD 1198 when a diversified pattern of pasture, meadow and arable land use was established. Colonisation followed the crusades, although in Livonia this occurred on a much smaller scale than in the rest of the Ordenstaat; Trikāta is atypical showing significant impact following the crusades with many other palaeoenvironmental studies only revealing more limited impact from the 14th century and later. Subsequent wars and changes in political control in the post-medieval period had little apparent effect on agricultural land use.

Limiting climatic factors and habitats ofErica tetralixat the eastern edge of its distribution range

We determined the limiting climatic factors, as well as the preferred habitats, of Erica tetralix L. at the eastern limit of its distribution range in the eastern Baltic region. It was found that E. tetralix in this region is a typical bog woodland plant preferring the wettest Sphagnum-rich sites. Northern Atlantic wet heath fragments, base-rich fens and species-rich Nardus grasslands were other habitats for the species. The species composition in E. tetralix habitats resembled that found in appropriate habitats within its main distribution range, although the habitats in the eastern Baltic region lack many Atlantic floristic elements characteristic of wet heath. A generalised linear model describing the climatic niche of E. tetralix in Latvia suggests that mild winters, which exhibit a combination of a shallow maximum depth of soil freezing and a large number of days when the air temperature exceeds 0°C, as well as abundant annual and winter precipitation, are the main factors allowing the presence of E. tetralix in Latvia. The climate parameters at the eastern limit of the species’ distribution range are consistent with those recorded as being suitable for E. tetralix elsewhere. Our results reveal a strong relationship between oceanity and the distribution of E. tetralix. The climatic niche model suggests more climatically suitable areas in the Coastal Lowland geobotanical region where the species could potentially be found, most likely in bog woodlands. The future prospects to restore open areas of northern Atlantic wet heath with E. tetralix in Latvia are poor due to the decline in traditional land-use; these areas have turned into bog woodlands or been converted into agricultural land.

Aphis pomi and Aphis spiraecola (Hemiptera: Sternorrhyncha: Aphididae) in Europe - new information on their distribution, molecular and morphological peculiarities

Aphid species Aphis pomi (de Geer, 1773) is oligophagous on pomoideous host plants, whilst Aphis spiraecola Patch, 1914 is a polyphagous species alternating between Spiraea spp., its primary host, and a wide variety of secondary hosts, also including pomoideous species. Despite the biological distinction, these species are difficult to separate using their morphological characters. Partial sequences of mitochondrial COI and nuclear EF-1α genes were analyzed for samples from Central and Eastern Europe, Germany, Bulgaria, Italy, Turkey, China together with available data from GenBank. Interspecific pairwise sample divergences of the COI fragment ranged from 3.1 to 4.3%. One COI haplotype of A. pomi was predominant (n = 24), with a pan European distribution. The most abundant COI haplotype of A. spiraecola (n = 16) occurred in Lithuania, Latvia, Poland, Italy, Turkey and China. Interspecific pairwise sample divergences of the EF-1α fragment ranged from 0.6 to 1.2%. Analyzed partial sequences of EF-1α were identical in A. pomi. The most abundant EF-1α haplotype of A. spiraecola (n = 14) occurred in Lithuania, Poland, Italy, Turkey and China. The length of ultimate rostral segment appeared to be the most reliable morphological character for discrimination between apple and spirea aphid species. It allowed a 100% correct identification of A. pomi (n = 143) and 91.5% of A. spiraecola (n = 94) specimens in the European samples used for the molecular analysis. The existence of A. spiraecola in the Eastern Baltic region of Europe is documented for the first time.