The direction of arrival (DOA) estimation technique is to obtain the direction information of the source when it reaches the array by processing and analyzing the received signal. In recent years, the DOA estimation of an array signal has been a research hotspot. For application scenarios with a small number of snapshots and a low signal-to-noise ratio, the compressive sensing theory has been commonly used to estimate the DOA of an array signal to achieve better estimation performance. However, the DOA estimation methods based on compressive sensing theory require information on source sparsity. Moreover, the influence of a complex underwater acoustic environment limits the accuracy of estimation algorithms. To address this limitation, this study proposes a high-precision DOA estimation model for underwater acoustic signals based on sparsity adaptation. The proposed model includes mainly two parts. In the first part, a source sparsity adaptive model based on a causal convolutional neural network is proposed. The model is used to address the constraint that the source sparsity should be known a priori when compressed sensing is used for DOA estimation. In the second part, a differential combination matching pursuit (DCMP) algorithm is adopted. First, a differentiated path filtering strategy is employed to reduce algorithm complexity and avoid the problem of invalid filtering. In addition, the combined optimization strategy is used to improve the prediction accuracy of the algorithm, providing an efficient error correction idea for the compressed sensing application to DOA estimation. The results of simulations conducted under seven different signal-to-noise ratios and using three different array types show that the proposed source sparsity adaptive model can reach an average prediction accuracy of 89.6%. In addition, compared with the other reconstruction algorithm accuracy, on the basis of ensuring low time complexity, the proposed DCMP algorithm can achieve an accuracy improvement of 9.99%–19.94% under seven different signal-to-noise ratio values. Moreover, the mean absolute error of the proposed DCMP algorithm is lower by approximately 0.05°–14° than those of the OMP and MMP algorithms.