INDIVIDUALS IN wild populations of Clarkia unguiculata may have up to six chromosomes in addition to the basic diploid number of 18 (as C. elegants, Lewis, 1951). These additional chromosomes apparently represent duplications of part of the basic genome, and are perhaps repeatedly produced by certain structural heterogyzotes (Lewis, 1951, 1954). These naturally occurring supernumerary chromosomes have no known effect on the phenotype (Lewis, 1951). Nevertheless, species formation in the genus Clarkia has often been accompanied by the addition of whole chromosomes to the basic genome (Lewis, 1953), but in these instances, the addition of a chromosome is associated with morphological differences, differences in habitat preference, and barriers to gene exchange.