Lamina Density Research Articles

Antixenosis in Cultivars of Tomato against <i>Helicoverpa armigera</i> (Hubner)

This study on the effect of antixenosis in relation to resistance against Helicoverpa armigera in tomato was carried out at the Entomological Research Farm, LPU, Phagwara during 2020 and 2021. The result showed that the antixenosis effect of four tomato cultivars on plant morphological characters revealed that Rahul cultivar with maximum trichome density in lamina (181 and 183 cm-2), vein (89 and 93 cm), trichome length (132.43 and 136.21 µm) and leaf lamina thickness (0.21 and 0.23 mm). Correlation analysis between morphological character and larval incidence revealed significant negative correlation with trichome density (lamina and vein), trichome length, leaf area and leaf lamina thickness.

Simulation and experimental study on the influence of lamina on nanoneedle penetration into the cell nucleus.

We have developed a finite element model to simulate the penetration of nanoneedles into the cellular nucleus. It is found that the nuclear lamina, the primary supporting structure of the nuclear membrane, plays a crucial role in maintaining the integrity of the nuclear envelope and enhancing stress concentration in the nuclear membrane. Notably, nuclear lamina A exhibits a more pronounced effect compared to nuclear lamina B. Subsequently, we further conducted experiments by controlling the time of osteopontin (OPN) treatment to modify the nuclear lamina density, and the results showed that an increase in nuclear lamina density enhances the probability of nanoneedle penetration into the nuclear membrane. Through employing both simulation and experimental techniques, we have gathered compelling evidence indicating that an augmented density of nuclear lamina A can enhance the penetration of nanoneedles into the nuclear membrane.

Development characteristics and controlling factors of fractures in lacustrine shale and their geological significance for evaluating shale oil sweet spots in the third member of the Shahejie Formation in the Qikou Sag, Bohai Bay Basin

Natural fractures are critical for shale oil and gas enrichment and development. Due to the extremely high heterogeneity of shale, the factors controlling the formation of internal fractures, especially horizontal fractures, remain controversial. In this study, we integrate thin section analysis and micro-computed tomography (CT) data from several lacustrine shale samples from the third member (Es3) of the Shahejie Formation, Qikou Sag, Bohai Bay Basin, to assess the fractures in detail. The goal is to reveal the development characteristics, controlling factors, and geological significance for evaluating sweet spots in a shale oil play. The fractures in the Es3 contain high-angle structural and horizontal bed-parallel fractures that are mostly shear and extensional. Various factors influence fracture development, including lithofacies, mineral composition, organic matter content, and the number of laminae. Structural fractures occur predominantly in siltstone, whereas bed-parallel fractures are abundant in laminated shale and layered mudstone. A higher quartz content results in higher shale brittleness, causing fractures, whereas the transformation between clay minerals contributes to the development of bed-parallel fractures. Excess pore pressure due to hydrocarbon generation and expulsion during thermal advance can cause the formation of bed-parallel fractures. The density of the bed-parallel and structural fractures increases with the lamina density, and the bed-parallel fractures are more sensitive to the number of laminae. The fractures are critical storage spaces and flow conduits and are indicative of sweet spots. The laminated shale in the Es3 with a high organic matter content contains natural fractures and is an organic-rich, liquid-rich, self-sourced shale play. Conversely, the siltstone, massive mudstone, and argillaceous carbonate lithofacies contain lower amounts of organic matter and do not have bed-parallel fractures. However, good reservoirs can form in these areas when structural fractures are present and the source, and storage spaces are separated.

Lamina Variation and Its Relationship with Sedimentary Facies in Alkaline Lacustrine, Permian Fengcheng Formation, Junggar Basin, Northwest China.

The Permian Fengcheng Formation in the Mahu Sag was deposited in a volcanic-alkaline lacustrine evaporative environment and contains a unique variety of fine-grained sediments. This study examines, at a millimeter-scale, the influence of sedimentary microfacies on variability of lamina quality in fine-grained sediments in the second member of the Fengcheng Formation (P1f2). The methods used include thin-section identification, X-ray diffraction (XRD), X-ray fluorescence (XRF), scanning electron microscopy (SEM), nitrogen adsorption, and nuclear magnetic resonance (NMR). Six types of lamina were identified in two different lithofacies: fan-delta front facies (FDFF) and semideep/deep lacustrine facies (SDDLF). The laminae in FDFF are predominantly feldspar-quartz laminae (FQL), reedmergnerite laminae (RL), shortite laminae (SL), alkaline mineral laminae (AML), and chert laminae (CL). The laminae in SDDLF are predominantly FQL, RL, SL, CL, and dolomite laminae (DOL). Variations in reservoir quality, oil-bearing properties, and the fracability of laminae in different sedimentary facies are determined by the combined effects of lamina density, mineral composition, rock structure, organic matter abundance, and microfractures. Analysis of these factors indicates superior reservoir qualities in FDFF. In SDDLF, the pore structure is limited by high lamina density, chert content, and fine grain size with the NMR porosities of FQL, RL, SL, and CL being 1.32, 0.18, 0.84, and 0.39%, respectively. However, in FDFF, the combination of high organic matter content, feldspar, pyrite, and clay minerals has a superior effect on the organic matter and minerals deposited resulting in better pore structure and more storage space for shale oil. The NMR porosities of FQL, RL, SL, and CL are 2.81, 2.53, 1.80, and 1.12%, respectively. Overall, analysis of lamina variations and their relationships with sedimentary facies indicates that the reservoir in FDFF may offer more favorable targets for "sweet spot" evaluation.

Predicting the Average Compression Strength of CLT by Using the Average Density or Compressive Strength of Lamina

The compressive strength in the major direction of cross-laminated timber CLT is the key to supporting the building load when CLT is used as load-bearing walls in high-rise wood structures. This study mainly aims to present a model for predicting the average compressive strength of CLT and promoting the utilization of CLT made out of planted larch. The densities and compressive strengths of lamina specimens and CLT samples with widths of 89 and 178 mm were evaluated, and their relationship was analyzed to build a prediction model by using Monte Carlo simulation. The results reveal that the average density of the lamina and CLT were about equal, whereas the average compressive strength of the CLT was just about 72% of that of the lamina. Width exerted no significant effect on the average compressive strength of the CLT, but homogenization caused the wider CLT to have a smaller variation than that of the lamina. The average compressive strength of the lamina could be calculated by using the average density of lamina multiply by 103.10, and the average compressive strength of the CLT could be calculated according to the compression strength of lamina in major and minor direction, therefore, a new prediction model is determined to predict the average compression strength of CLT by using the average density of lamina or CLT, the average compression strength of CLT made in this study is about 74.23 times of the average density of the lamina. The results presented in this study can be used to predict the average compressive strength of CLT by using the average density of lamina and provide a fundamental basis for supporting the utilization of CLT as load-bearing walls.

The Influence of Lamina Density and Occurrence on the Permeability of Lamellar Shale after Hydration

The characteristics of laminae in lamellar shale oil reservoirs have important influences on reservoir parameters, especially permeability. In order to explore the influence of lamina density and occurrence on the permeability of lamellar shale after hydration, we studied the lamellar shale in the Chang 7 member of the Yanchang Formation of Triassic in Ordos Basin. By comparing the permeability of bedding shale and lamellar shale with different densities of laminae, it was found that the permeability anisotropy of lamellar shale was stronger. In the direction parallel to the lamina, the permeability increased approximately linearly with an increase in lamina density. The effect of hydration on rock micropore structure and permeability was studied by soaking shale in different fluids. Most of the microfracture in the lamellar shale was parallel to the lamina direction, and hydration led to a widening of the microfracture, which led to the most obvious increase in permeability parallel to the lamina. Collectively, the research results proved that lamina density, occurrence, and hydration have a significant influence on the permeability anisotropy of lamellar shale.

BIOVERA-Tree: tree diversity, community composition, forest structure and functional traits along gradients of forest-use intensity and elevation in Veracruz, Mexico.

BackgroundHere, we describe BIOVERA-Tree, a database on tree diversity, community composition, forest structure and functional traits collected in 120 forest plots, distributed along an extensive elevational gradient in Veracruz State, Mexico. BIOVERA-Tree includes information on forest structure from three levels of forest-use intensity, namely old-growth, degraded and secondary forest, replicated across eight elevations from sea-level to near the tree line at 3500 m and on size and location of 4549 tree individuals with a diameter at breast height ≥ 5 cm belonging to 216 species, 154 genera and 80 families. We also report measurements of eight functional traits, namely wood density for 143 species, maximum height for 216 species and leaf traits including: specific leaf area, lamina density, leaf thickness, chlorophyll content and leaf area for 148 species and leaf dry matter content for 145 species.New informationBIOVERA-Tree is a new database comprising data collected in a rigorous sampling design along forest-use intensity and elevational gradients, contributing to our understanding of how interactive effects of forest-use intensity and elevation affect tree diversity, community composition and functional traits in tropical forests.

Testing the competition-colonization trade-off and its correlations with functional trait variations among subtropical tree species

The competition-colonization trade-off, by which species can partition spatial niches, is a potentially important mechanism allowing the maintenance of species diversity in plant communities. We examined whether there was evidence for this trade-off among tree species in a subtropical forest and how it correlated with eight functional traits. We developed and estimated a metric for colonization ability that incorporates both fecundity and seed dispersal based on seed trap data and the sizes and distributions of adult trees. Competitive ability was estimated as survival probability under high crowding conditions based on neighborhood models. Although we found no significant relationship between colonization and competitive abilities, there was a significant negative correlation between long distance dispersal ability and competitive ability at the 5 cm size class. Colonizers had traits associated with faster growth, such as large leaves and low leaf lamina density, whereas competitors had traits associated with higher survival, such as dense wood. Our results imply that any trade-off between competition and colonization may be more determined by dispersal ability than by fecundity, suggesting that seed dispersal is an important contributor to diversity maintenance. Future work should test how competitive ability covaries with the components of colonization ability, as we did here.

KARAKTERISTIK SIFAT FISIKA DAN MEKANIKA KAYU LAMINA KOMBINASI JENIS KAYU SENGON (Paraserianthes falcataria (L.) Nilsen) DAN JENIS KAYU MERBAU (Intsia spp.)

This study aims to determine the physical and mechanical characteristics of wood lamina combination of sengon wood and merbau with Melamine Urea Formaldehyde (MUF) adhesive. The testing method used is based on the DIN standard. The test results of the average lamina density of 0.58-0.73 g / cm3 at oven drying moisture content. Modulus of Elasticity (MoE), Modulus of Rupture (MoR) and value of compression parallel to grain highest found in the combination of merbau-sengon-merbau-sengon-merbau and the lowest in the combination of sengon-merbau-sengon. While the adhesive shear strength is the highest value in the combination treatment of merbau-merbau and the lowest in the combination of sengon-sengon. Based on the modulus of elasticity (MoE) of sengon treatment as face / back and merbau as core (3 and 5 layers) can be classified into strong class IV-V, merbau as face / back and sengon as core (3 and 5 layers) can be classified into strong class II-III. Modulus of Rupture (MoR) can be classified into strong class II-III and compression parallel to grain into strong class II.

KARAKTERISTIK SIFAT FISIKA DAN MEKANIKA KAYU LAMINA KOMBINASI JENIS KAYU SENGON (Paraserianthes falcataria (L.) Nilsen) DAN JENIS KAYU MERBAU (Intsia spp.)

This study aims to determine the physical and mechanical characteristics of wood lamina combination of sengon wood and merbau with Melamine Urea Formaldehyde (MUF) adhesive. The testing method used is based on the DIN standard. The test results of the average lamina density of 0.58-0.73 g / cm3 at oven drying moisture content. Modulus of Elasticity (MoE), Modulus of Rupture (MoR) and value of compression parallel to grain highest found in the combination of merbau-sengon-merbau-sengon-merbau and the lowest in the combination of sengon-merbau-sengon. While the adhesive shear strength is the highest value in the combination treatment of merbau-merbau and the lowest in the combination of sengon-sengon. Based on the modulus of elasticity (MoE) of sengon treatment as face / back and merbau as core (3 and 5 layers) can be classified into strong class IV-V, merbau as face / back and sengon as core (3 and 5 layers) can be classified into strong class II-III. Modulus of Rupture (MoR) can be classified into strong class II-III and compression parallel to grain into strong class II.

Eighty years of food-web response to interannual variation in discharge recorded in river diatom frustules from an ocean sediment core

Little is known about the importance of food-web processes as controls of river primary production due to the paucity of both long-term studies and of depositional environments which would allow retrospective fossil analysis. To investigate how freshwater algal production in the Eel River, northern California, varied over eight decades, we quantified siliceous shells (frustules) of freshwater diatoms from a well-dated undisturbed sediment core in a nearshore marine environment. Abundances of freshwater diatom frustules exported to Eel Canyon sediment from 1988 to 2001 were positively correlated with annual biomass of Cladophora surveyed over these years in upper portions of the Eel basin. Over 28 years of contemporary field research, peak algal biomass was generally higher in summers following bankfull, bed-scouring winter floods. Field surveys and experiments suggested that bed-mobilizing floods scour away overwintering grazers, releasing algae from spring and early summer grazing. During wet years, growth conditions for algae could also be enhanced by increased nutrient loading from the watershed, or by sustained summer base flows. Total annual rainfall and frustule densities in laminae over a longer 83-year record were weakly and negatively correlated, however, suggesting that positive effects of floods on annual algal production were primarily mediated by "top-down" (consumer release) rather than "bottom-up" (growth promoting) controls.

Early-onset treadmill training reduces mechanical allodynia and modulates calcitonin gene-related peptide fiber density in lamina III/IV in a mouse model of spinal cord contusion injury.

Below-level central neuropathic pain (CNP) affects a large proportion of spinal cord injured individuals. To better define the dynamic changes of the spinal cord neural network contributing to the development of CNP after spinal cord injury (SCI), we characterized the morphological and behavioral correlates of CNP in female C57BL/6 mice after a moderate T11 contusion SCI (50 kdyn) and the influence of moderate physical activity. Compared with sham-operated animals, injured mice developed mechanical allodynia 2 weeks post injury when tested with small-diameter von Frey hair filaments (0.16 g and 0.4 g filament), but presented hyporesponsiveness to noxious mechanical stimuli (1.4 g filament). The mechano-sensory alterations lasted up to 35 days post injury, the longest time point examined. The response latency to heat stimuli already decreased significantly 10 days post injury reaching a plateau 2 weeks later. In contrast, injured mice developed remarkable hyposensitivity to cold stimuli. Animals that underwent moderate treadmill training (2 × 15 minutes; 5 d/wk) showed a significant reduction in the response rate to light mechanical stimuli as early as 6 days after training. Calcitonin gene-related peptide (CGRP) labeling in lamina III-IV of the dorsal horn revealed significant increases in CGRP-labeling density in injured animals compared with sham control animals. Importantly, treadmill training reduced CGRP-labeling density by about 50% (P < 0.01), partially reducing the injury-induced increases. Analysis of IB4-labeled nonpeptidergic sensory fibers revealed no differences between experimental groups. Abnormalities in temperature sensation were not influenced by physical activity. Thus, treadmill training partially resolves signs of below-level CNP after SCI and modulates the density of CGRP-labeled fibers.

Constraints on the lamina density of laminar bone architecture of large-bodied dinosaurs and mammals

Laminar bone tissue is commonly found in Dinosauria (including birds) and Mammalia. The tissue emerged convergently several times, and its frequent occurrence among amniotes has stimulated researchers to study some of its geometric features. One such feature is lamina thickness or lamina density (LD, expressed as number of laminae per mm). We measured LD in a sample of sauropodomorph dinosaur taxa (basal sauropodomorphs, basal sauropods and Neosauropoda) and compared it with LD of a selection of mammals. LD is relatively constrained within the groups; nonetheless mean sauropodomorph LD differs significantly from mean mammal LD. However, increasing sample size with other dinosaur groups and more perissodactyls and artiodactyls may alter this result. Among sauropods, LD does not change drastically with increasing femur length although a slight tendency to decrease may be perceived. We conclude that the laminar vascular architecture is most likely determined by a combination of structural and functional as well as vascular supply and physiological causes.

Functional trait divergence of juveniles and adults of nine Inga species with contrasting soil preference in a tropical rain forest

Summary1. Community‐wide studies have shown that functional traits of tropical trees vary with soil‐type preference, but few have examined trait diversification among closely related taxa. In this study, we asked how functional traits of adults and saplings within a speciose genus, Inga, differ in relation to their soil‐type preferences.2. We quantified soil‐type preference and functional traits of nine Inga species (Fabaceae) in the wet tropical forest at La Selva Biological Station, Costa Rica, where rich alluvial soil and less fertile residual soil of volcanic origin occur in close proximity.3. From the extensive trail network, we randomly selected 104 trail segments, each 50 m long, along which all Inga individuals &gt;50 cm tall were counted and identified to species. Five and three common species showed significant bias to residual or alluvial soil, respectively, while the remaining one showed no significant bias. For these nine species, we quantified five leaf traits for saplings and adults, as well as wood‐specific gravity (WSG) and three seed traits.4. For both adults and saplings, leaf mass per area (LMA) and lamina density were higher for residual‐soil specialists than fertile alluvial‐soil specialists. The opposite trend was found for nitrogen (N) and phosphorus (P) concentrations in leaves. Lamina thickness did not differ between soil‐type preference groups. All leaf trait values except N concentration increased from saplings to adults, with adults showing larger interspecific variations in trait values than saplings.5. Saplings and adults of the single species found on both soils had trait values intermediate between residual and alluvial‐soil specialists.6. Seed mass and seed N, as well as WSG of adults, were higher in residual‐soil specialists than alluvial‐soil specialists. Higher seed N indicates higher maternal investment of species growing on less fertile soil.7. These differences in traits among closely related species across continuous soils suggest that habitat filtering has contributed to the evolution of seed, juvenile and adult traits within a speciose tropical tree genus.

How cellulose‐based leaf toughness and lamina density contribute to long leaf lifespans of shade‐tolerant species

Cell wall fibre and lamina density may interactively affect leaf toughness and leaf lifespan. Here, we tested this with seedlings of 24 neotropical tree species differing in shade tolerance and leaf lifespan under standardized field conditions (140-867 d in gaps; longer in shade). We quantified toughness with a cutting test, explicitly seeking a mechanistic linkage to fibre. Lamina density, but not fracture toughness, exhibited a plastic response to gaps vs shade, while neither trait was affected by leaf age. Toughness corrected for lamina density, a recently recognized indicator of material strength per unit mass, was linearly correlated with cellulose content per unit dry mass. Leaf lifespan was positively correlated with cellulose and toughness in shade-tolerant species but only weakly in gap-dependent species. Leaf lifespan was uncorrelated with lamina thickness, phenolics and tannin concentrations. In path analysis including all species, leaf lifespan was directly enhanced by density and toughness, and indirectly by cellulose via its effect on toughness. Different suites of leaf traits were correlated with early seedling survival in gaps vs shade. In conclusion, cellulose and lamina density jointly enhance leaf fracture toughness, and these carbon-based physical traits, rather than phenolic-based defence, explain species differences in herbivory, leaf lifespan and shade survival.

What Makes a Leaf Tough? Patterns of Correlated Evolution between Leaf Toughness Traits and Demographic Rates among 197 Shade-Tolerant Woody Species in a Neotropical Forest

Slow-growing juveniles of shade-tolerant plant species are predicted to have tough leaves because of the high cost of leaf replacement in shade relative to potential carbon gain. We assessed the degree of correlated evolution among eight traits associated with leaf toughness and the relationships of those traits with the growth and mortality rates of 197 tree and shrub species from the understory of the 50-ha forest dynamics plot on Barro Colorado Island, Panama. Path analysis with phylogenetically independent contrasts revealed that leaves attained material toughness (resistance to fracture per unit fracture area) through increases in tissue density, percent cellulose per unit dry mass, and vein fracture toughness. Lamina density and cellulose content evolved independently and thus represent different paths to material toughness. Structural toughness (resistance to fracture per unit fracture length) depended on material toughness and lamina thickness. Mortality rates of individuals 1-10 cm in stem diameter were negatively correlated with material toughness and lamina density but were independent of structural toughness and cell wall fiber contents. Leaf toughness traits were uncorrelated with relative growth rates. Results imply that material toughness enhances resistance to natural enemies, which increases survival and offsets the biomass allocation cost of producing tough leaves in the shaded understory.

Change in central kinin B2 receptor density after exercise training in rats

Cardiovascular responses elicited by the stimulation of kinin B2 receptors in the IV cerebral ventricle, paratrigeminal nucleus or in the thoracic spinal cord are similar to those observed during an exercise bout. Considering that the kalikrein–kinin system (KKS) could act on the cardiovascular modulation during behavioral responses as physical exercise or stress, this study evaluated the central B2 receptor densities of Wistar (W) and spontaneously hypertensive rats (SHR) after chronic moderate exercise. Animals were exercise-trained for ten weeks on a treadmill. Afterwards, systolic blood pressure decreased in both trained strains. Animals were killed and the medulla and spinal cord extracted for B2 receptor autoradiography. Trained animals were compared to their sedentary controls. Sedentary groups showed specific binding sites for Hoe-140 (fmol/mg of tissue) in laminas 1 and 2 of the spinal cord, nucleus of the solitary tract (NTS), area postrema (AP), spinal trigeminal tract (sp5) and paratrigeminal nucleus (Pa5). In trained W a significant increase (p < 0.05) in specific binding was observed in the Pa5 (31.3%) and NTS (28.2%). Trained SHR showed a significant decrease in receptor density in lamina 2 (21.9%) of the thoracic spinal cord and an increase in specific binding in Pa5 (36.1%). We suggest that in the medulla, chronic exercise could hyper stimulate the KKS enhancing their efficiency through the increase of B2 receptor density, involving this receptor in central cardiovascular control during exercise or stress. In the lamina 2, B2 receptor might be involved in the exercise-induced hypotension.

Tissue‐level leaf toughness, but not lamina thickness, predicts sapling leaf lifespan and shade tolerance of tropical tree species

Leaf toughness is thought to enhance physical defense and leaf lifespan. Here, we evaluated the relative importance of tissue-level leaf traits vs lamina thickness, as well as their ontogenetic changes, for structure-level leaf toughness and regeneration ecology of 19 tropical tree species. We measured the fracture toughness of the laminas and veins of sapling leaves with shearing tests, and used principal component analysis and structural equation modeling to evaluate the multivariate relationships among traits that contribute to leaf toughness and their links to ecological performance traits. Tissue traits (density and fracture toughness of lamina and vein) were correlated positively with each other, but independent of lamina thickness. The tissue traits and lamina thickness contributed additively to the structure-level toughness (leaf mass per area and work-to-shear). Species with dense and tough leaves as saplings also had dense and tough leaves as seedlings and adults. The patterns of ontogenetic change in trait values differed between the seedling-to-sapling and sapling-to-adult transitions. The fracture toughness and tissue density of laminas and veins, but not the lamina thickness, were correlated positively with leaf lifespan and sapling survival, and negatively with herbivory rate and sapling regeneration light requirements, indicating the importance of tissue-level leaf traits.

Simultaneous determination of shear strength and shear modulus in glued-laminated timber using a full-scale shear block specimen

A newly developed full-scale shear block specimen was used to simultaneously determine the shear strength and shear modulus of glued-laminated timber. The shear modulus was calculated using the shear strain distribution measured by means of digital image correlation. To obtain the exact relationship between shear modulus and shear strength, the shear strain in the intended shear plane was measured. A relatively high correlation coefficient of 0.75 was found between the shear modulus and nominal shear strength in two types of glued-laminated timber tested: sugi (Cryptomeria japonica) and Douglas-fir (Pseudotsuga menziesii). Good correlation was also found between the density of middle lamina and shear strength. However, the relationships to density showed species dependence, which was not observed in the relationship between shear modulus and shear strength.

A mechanistic model for transverse damage initiation, evolution, and stiffness reduction in laminated composites

A constitutive model to predict stiffness reduction due to transverse matrix cracking is derived for laminae with arbitrary orientation, subject to in-plane stress, embedded in laminates with symmetric but otherwise arbitrary laminate stacking sequence. The moduli of the damaged laminate are a function of the crack densities in the damaging laminae, which are analyzed one by one. The evolution of crack density in each lamina is derived in terms of the calculated strain energy release rate and predicted as function of the applied load using a fracture mechanics approach. Unlike plasticity-inspired formulations, the proposed model does not postulate damage evolution functions and thus there is no need for additional experimental data to adjust material parameters. All that it is needed are the elastic moduli and critical energy release rates for the laminae. The reduction of lamina stiffness is an integral part of the model, allowing for stress redistribution among laminae. Comparisons with experimental data and some results from the literature are presented.