Denali National Park Research Articles

Influence of atmospheric state on variability of long-term residual ambient sound level measurements in a subalpine valley.

Two natural influences on the acoustic environments of mountainous parks and communities are flowing water and shifting weather. A central purpose of the acoustic measurement design used by the United States National Park Service is to provide spectral estimates of residual ambient sound level metrics at a seasonal time scale. Acoustic monitoring sampling methodologies are often designed using a sequence of similar measurements. When source and residual ambient spectra overlap, an estimate of variability in the latter is beneficial to successful monitoring design. The observed and modelled effects of atmospheric state on sound level are analyzed to reveal variability due to these effects at a long-term monitoring site in Denali National Park, Alaska. The analysis of variability incorporates a covariate that is otherwise challenging to estimate in remote settings: vertical temperature gradients in the atmospheric boundary layer. Results reveal inversions (positive gradients) in the atmosphere ≥30% between 19:00 and 09:00. Inversion strengths above 0.06 °C/m are associated with 10-15 dB increases in sound level over hourly time scales. Because inversions tend to occur during otherwise quiescent times of day, they ultimately reduce seasonal variability at the site and corresponding uncertainty in noise metrics for transportation noise arriving from varied directions.

Suspected True Hermaphroditism in a Free-ranging Gray Wolf (Canis lupus) in Interior Alaska, USA.

Hermaphroditism or intersexuality occurs endogenously in most mammal species. We document the behavior and physical and anatomic appearance of a suspected true hermaphroditic free-ranging wild gray wolf (Canis lupus) in Denali National Park and Preserve, Alaska, USA.

Methods for managing soundscape quality: Evidence from Denali National Park and Preserve

The purpose of the United States national park system is to preserve resources unimpaired for the enjoyment of future generations. At Denali National Park (DENA), high levels of air tour traffic and air transportation present challenges to preserving wilderness character. Accordingly, the park attached high priority to measuring visitor impressions of soundscape quality. Visitors in four distinct settings rated the acceptability of five randomly chosen recordings of aircraft noise. We used a cumulative link mixed model to fit visitor’s response to acoustic and nonacoustic factors. The negative effects of increased noise were moderated for visitors who were interested in taking an air tour. To be conservative, we examined visitors uninterested in an air tour and found the probability of rating aircraft noise as unacceptable at 54 dB LAeq, 30 s or higher was 26%. For context, noise above 55dB is incompatible with outdoor, rural activities. Visitor response predictions were joined to a spatial model of aircraft noise propagation to create a map depicting the acceptability of aircraft noise in DENA’s frontcountry area. The map can be used to forecast the range of soundscape conditions park visitors are exposed to, inform hiking recommendations for visitors, and evaluate park management zones.

Transformative potential of nature-based values that influence the relationships between reported and intended pro-environmental behavior

Protected area landscapes embody multiple values of nature that can create meaning in everyday life. Though the values ascribed to these environments theoretically inspire changes in human behavior, surprisingly few studies have empirically evaluated how ‘specific values’ affect actions that benefit the environment. We used Public Participation in Geographic Information Systems (PPGIS) methods to evaluate the relationships among four nature-based values and the patterns of both reported and intended behavior among visitors to Denali National Park and Preserve, Alaska, USA (n = 667). We found that wilderness, recreation, ecological integrity, and scientific qualities of places were particularly important for characterizing the Denali landscape and accounted for more variation in intended than reported pro-environmental behaviors. We provide new insights on how nature-based values underpin the decisions of visitors and lead to transformative changes after experiencing a high profile, charismatic protected area. Understanding the reasons why people forge connections with natural areas and modeling how these associations relate to different types of behavior advances knowledge of how to effectively build environmental stewardship and guide public land management decisions. Management implicationsFindings from this study provide public land management agencies with insight on what people value in protected areas, the locations they appreciate, and how their experiences may influence their behaviors after returning home. We contend that “nature-based values” that encompass recreation, ecological integrity, wilderness, and scientific qualities of nature are particularly important to visitors and can provide a basis for communication about high and low priority places, as well as spatially explicit guidance for management agencies. Also, we show that as nature-based values increase, so do the reported and intended actions of people who visit Denali National Park and Preserve. As such, these multiple values of nature should be harnessed to inspire and energize more environmentally friendly practices in the future.

Early Holocene ice on the Begguya plateau (Mt. Hunter, Alaska) revealed by ice core 14C age constraints

Abstract. Investigating North Pacific climate variability during warm intervals prior to the Common Era can improve our understanding of the behavior of ocean–atmosphere teleconnections between low latitudes and the Arctic under future warming scenarios. However, most of the existing ice core records from the Alaskan and Yukon region only allow access to climate information covering the last few centuries. Here we present a surface-to-bedrock age scale for a 210 m long ice core recovered in 2013 from the summit plateau of Begguya (Mt. Hunter; Denali National Park, Central Alaska). Combining dating by annual layer counting with absolute dates from micro-radiocarbon dating, a continuous chronology for the entire ice core archive was established using an ice flow model. Calibrated 14C ages from the deepest section (209.1 m, 7.7 to 9.0 ka cal BP) indicate that basal ice on Begguya is at least of early Holocene origin. A series of samples from a shallower depth interval (199.8 to 206.6 m) were dated with near-uniform 14C ages (3 to 5 ka cal BP). Our results suggest this may be related to an increase in annual net snow accumulation rates over this period following the Northern Hemisphere Holocene Climate Optimum (around 8 to 5 kyr BP). With absolute dates constraining the timescale for the last >8 kyr BP, this paleo-archive will allow future investigations of Holocene climate and the regional evolution of spatial and temporal changes in atmospheric circulation and hydroclimate in the North Pacific.

Vertebrate ichnology and palaeoenvironmental associations of Alaska’s largest known dinosaur tracksite in the Cretaceous Cantwell Formation (Maastrichtian) of Denali National Park and Preserve

ABSTRACT The Upper Cretaceous Cantwell Formation in Denali National Park and Preserve (DENA) preserves an abundant and diverse ichnofossil record of high latitude dinosaurs. Field-based investigations of the formation remain limited due to its wide areal extent and remoteness, leaving questions concerning its temporal and faunistic relationships to other units unaddressed. Here we describe the largest tracksite known in DENA and all of Alaska – a 7500 square metre outcrop of steeply dipping beds known as the Coliseum. The site exposes 66.3 metres of vertical section consisting of laterally extensive fine- to medium-grained sandstone, mudstone, calcareous shale, and bentonite. Vertebrate trace fossils include true tracks, transmitted tracks, natural casts, and trackways documented via handheld and UAV-assisted photogrammetry, enabling three-dimensional mapping of the site. Facies analysis reveals a depositional environment of crevasse splay and overbank deposits within a fluvial floodplain. U-Pb dating of bentonite collected from the site returned an age of 69.3 ± 0.9 Ma, improving the limited temporal constraints of the formation. We document several ichnotaxa, some new to the formation, representing ornithopods, ceratopsids, and large and small-bodied avian and non-avian theropods. The Coliseum provides a rare window into a high latitude, Late Cretaceous forested ecosystem deposited in a Greenhouse World.

High-Latitude Depositional Systems, Provenance, and Basinal Setting of the Late Cretaceous Cantwell Basin, Denali National Park and Preserve, Alaska: A Stratigraphic Framework for Paleontological and Paleoclimatic Studies

The Cantwell Formation of the central Alaska Range provides a robust archive of high-latitude, Late Cretaceous depositional systems and paleo-floral/faunal assemblages. Our stratigraphic analysis defines two mappable members. The lower member (1500–2000 m thick) represents vegetated alluvial fan and braided fluvial systems that transition up-section to fluvial–estuarine systems that drained into an inland continental seaway. The upper member (~2000 m thick) represents estuarine–marginal marine and lacustrine systems. Previous paleontological studies demonstrate that the Cantwell basin was populated by various dinosaurs, fishes, bivalves, birds, and marginal marine micro-organisms. Integration of new and published geologic mapping allows for reconstruction of depositional systems at the basin scale and provides additional paleogeographic context. The northern basin margin was defined by a previously unrecognized south-verging thrust belt, whereas the southern margin of the basin was defined by a north-verging thrust belt inboard of an active magmatic arc. Sediment sources interpreted from U-Pb detrital zircon geochronology included the coeval magmatic arc and older Cretaceous plutons, and Proterozoic–Mesozoic strata exhumed along the basin margins. Results of our study provide a depositional, stratigraphic, and structural framework that may serve as a guide for future paleontological and paleoclimatic investigations of Late Cretaceous Arctic environments of the Cantwell basin.

How much noise is too much? Methods for identifying thresholds for soundscape quality and ecosystem services

The United States National Park Service mandate is to conserve park resources and provide superlative visitor experience. In the context of acoustic resources, Denali National Park and Preserve provides an advantageous opportunity to understand the effect of aircraft noise on visitor experience because it possesses high levels of air tour traffic in a park renowned for its remote, wilderness character. Park visitors in four different settings were asked to rate the acceptability of recordings of aircraft noise, presented in randomized order relative to noise level. A cumulative link mixed model fitted visitor assessments to acoustic and nonacoustic factors. In addition to noise level, interest in an air tour was an important predictor of sound clip acceptability. For visitors uninterested in an air tour, the probability of rating aircraft noise as unacceptable at 54 dB LAeq,30 s or higher was 26%. For reference, this aligns with federal guidance that identified 55 dB as a threshold for interference with outdoor activities at rural residences and schools. Predictions of visitor response were joined to a spatial model of aircraft noise propagation to map visitor acceptability of aircraft noise in Denali’s entrance area (frontcountry). This map can be used to assess the condition of park management zones, to inform hiking recommendations for visitors, and to predict the range of soundscape conditions experienced by park visitors.

Tree and shrub expansion at treeline drive contrasting responses in a subarctic passerine community.

Inferences about the mechanisms of distributional change are often made from simple assessments of variation in the geographical positions of populations. However, direct assessments of species' responses to local habitat change may be necessary for proper understanding of the drivers of distributional dynamics. Amplified climate warming is inducing cascading impacts in boreal-tundra regions including the expansion of conifers and deciduous shrubs (shrubs). In Denali National Park (Denali), Alaska, passerine birds are exhibiting rapid upslope shifts in distribution but the relative roles of conifer and shrub (woody vegetation) expansion in driving these shifts are unknown. Without directly assessing passerine-vegetation dynamics, the assumption has been that the observed upslope shifts are indicative of shrub-adapted passerines tracking the upslope expansion of shrubs. Here, we jointly investigate the processes of conifer and shrub expansion and their relationship to changes in passerine abundance in Denali. We used a remotely sensed vegetation cover timeseries (1985-2020) to assess the topographic and edaphic correlates of conifer and shrub expansion. We then assessed the impacts of changes in shrub and conifer cover on the relative abundance of 12 passerine species (1995-2020). Shrub and conifer colonization rates were highest at intermediate elevations near treeline. However, forest- and shrub-adapted passerines differed in terms of the location in which their response was concentrated relative to treeline. The population growth rates of forest-adapted passerines exhibited stronger effects of woody vegetation expansion at sites that were initially above treeline (IAT). In contrast, the population growth rates of shrub-adapted passerines exhibited the negative effects of conifer expansion together with the positive effects of shrub expansion at initially below treeline sites. However, they showed a weak response to woody vegetation expansion at sites that were IAT. Below treeline conifer infilling appears to be pushing the elevational distributions of shrub-adapted passerines upslope rather than these species following the pull of modest shrub expansion above treeline, as previously assumed. Overall, our findings illustrate the need for explicitly accommodating heterogeneity in habitat change at small spatial scales to properly view the distributional response, particularly when habitat change is concentrated at ecotones.

Examining Visitor Collected Species Data from Denali National Park and Preserve

In citizen science programs, the concept of “participant” is often used as a catch-all without considering how different participants (e.g., young, old, wealthy, poor, differently-abled, local inhabitants, and visitors) affect scientific outcomes of citizen science programs (e.g., the data collection, data analysis, publications, etc.). This research advances the understanding of tourist participants’ ability to produce data comparable to participants who live near the study area. To examine data collected by tourist participants, we performed a case study on wildlife observation data collected through the Map of Life-Denali program in Denali National Park and Preserve in Alaska, USA. The species observation data collected by tourists and Alaska Residents were compared visually using heatmaps and statistically with the Ripley’s K function variation, the L function. Results from the analysis show that the tourist and resident data have similar point patterns for the three species we compared, <em>Ursus arctos</em> (Grizzly Bear), <em>Rangifer tarandus</em> (Caribou), and <em>Alces alces</em> (Moose). Our results indicate that tourists can be effective citizen science participants. And show the potential for leveraging this large pool of untapped participants in popular tourist destinations such as U.S. National Parks.

Integrating social values with GPS tracks through Denali National Park and Preserve

This study advanced knowledge of the geospatial relationships between social values elicited during a participatory mapping exercise and on-ground travel patterns understood through Global Positioning System (GPS) tracking of backcountry visitors to a protected area in Alaska. As one of the first studies to combine social values and real-time use of a protected area landscape, we showcase how these combined forms of knowledge can be better understood when compared against biophysical conditions. Contrary to previous research, we observed that perceived social value hotspots, defined by an abundance of point data, did not fully align with use patterns, suggesting that visitors value areas that are not experienced first-hand. Specifically, backcountry travel routes in Denali were less dispersed than areas perceived to be important. Use was mostly concentrated in backcountry units close to the middle sections of the park road while highly valued units coincided with major landmarks, such as the peak of Denali. Travel cost induced by terrain conditions (summarized by elevation, slope and landcover), accessibility (measured by proximity to the park road), and long-view visual resources all contributed to how observed travel behavior deviated from perceived social values. These findings help inform policy and management decisions about outdoor recreation, visitor safety, and visual resource stewardship.

Water source dynamics influence macroinvertebrate communities across groundwater-fed streams in a glacierized catchment

Groundwater contributions to streamflow significantly influence the structure and function of riverine ecosystems, particularly in glacierized catchments where there are marked differences in water sources and subsurface flow paths. Here, we investigated spatial and temporal variation in relationships between water sources, flow paths, physical and chemical processes, organic matter, microbial biofilms, and macroinvertebrates across groundwater-fed streams in the glacierized Toklat River catchment of Denali National Park, Alaska. Streams fed predominantly by seepage from the valley sides were perennial, whereas streams sustained by glacial meltwater seepage were ephemeral. Differences in environmental conditions between flow regimes appeared to influence spatial and temporal patterns of organic matter, linking to macroinvertebrate community dynamics. Macroinvertebrates in perennial streams were supported by fine particulate organic matter from subsurface flow paths during summer, transitioning to a combination of fine particulate matter and leaf litter in autumn. In comparison, macroinvertebrates inhabiting ephemeral streams, which only flowed during autumn, were supported by leaf litter. Some macroinvertebrate taxa were unaffected by turnover in organic matter, indicating potential plasticity in organic matter resource use. Findings highlight the importance of considering spatial and temporal variation in groundwater-fed streams, considering that projected hydrological changes under a changing climate may have significant implications for these systems.

Arthropod communities along an elevation gradient in Denali National Park and Preserve, Alaska: Rapidly shrinking tundra hosts a unique assemblage of specialists

ABSTRACT Arthropods at high latitudes and elevations are likely to be vulnerable to effects from climate change such as increased temperatures and shifting vegetation boundaries. Though range shifts northwards and upslope have been reported for many arthropod taxa in temperate latitudes, baseline data needed to track such changes are scarce at northern latitudes. We investigated the influence of climate and vegetation cover on the abundance, diversity, and species composition of pollinators and epigeic arthropods along an elevation gradient in Denali National Park and Preserve, Alaska. We compared arthropods across three habitat types: low-elevation forest, mid-elevation shrubs, and high-elevation tundra along five replicate transects over three years. We collected 35,473 arthropods representing 510 species. Arthropod communities differed distinctly across the three habitat types, with tundra having the highest number of strong indicators and unique species. Elevation, air temperature, and vegetation structure were strong drivers for the ordination of sites. As treeline and shrubline shift upslope with climate change, we predict that distributions of some arthropods will shift to track these habitat boundaries and that tundra-associated arthropods will be most vulnerable as their habitat shrinks. Long-term monitoring of arthropods along elevation gradients at northern latitudes is needed to detect such declines.

Long-term changes in macroinvertebrate communities across high-latitude streams.

Long-term records of benthic macroinvertebrates in high-latitude streams are essential for understanding climatic changes, including extreme events (e.g. floods). Data extending over multiple decades are typically scarce. Here, we investigated macroinvertebrate community structural change (including alpha and beta diversity and gain and loss of species) over 22 years (1994-2016) in 10 stream systems across Denali National Park (Alaska, USA) in relation to climatological and meteorological drivers (e.g. air temperature, snowpack depth, precipitation). We hypothesised that increases in air temperature and reduced snowpack depth, due to climatic change, would reduce beta and gamma diversity but increase alpha diversity. Findings showed temporal trends in alpha diversity were variable across streams, with oscillating patterns in many snowmelt- and rainfall runoff-fed streams linked to climatic variation (temperature and precipitation), but increased over time in several streams supported by a mixture of water sources, including more stable groundwater-fed streams. Beta-diversity over the time series was highly variable, yet marked transitions were observed in response to extreme snowpack accumulation (1999-2000), where species loss drove turnover. Gamma diversity did not significantly increase or decrease over time. Investigating trends in individual taxa, several taxa were lost and gained during a relative constrained time period (2000-2006), likely in response to climatic variability and significant shifts in instream environmental conditions. Findings demonstrate the importance of long-term biological studies in stream ecosystems and highlight the vulnerability of high-latitude streams to climate change.

Human‐caused mortality triggers pack instability in gray wolves

Transboundary movement of wildlife results in some of the most complicated and unresolved wildlife management issues across the globe. Depending on the location and managing agency, gray wolf (Canis lupus) management in the US ranges from preservation to limited hunting to population reduction. Most wildlife studies focus on population size and growth rate to inform management, but relatively few examine species biological processes at scales aside from that of the population. This is especially important for group‐living species such as the gray wolf, for which the breeding unit is the social group. We analyzed data for gray wolf packs living primarily within several US National Park Service units (years of data): Denali National Park and Preserve (33 years), Grand Teton National Park (23 years), Voyageurs National Park (12 years), Yellowstone National Park (27 years), and Yukon‐Charley Rivers National Preserve (23 years). We identified two gray wolf biological processes that differed from population size – namely, pack persistence and reproduction – and determined that while human‐caused mortality had negative effects on both, pack size had a moderating effect on the impacts of mortality.

Climate Indicators of Landslide Risks on Alaska National Park Road Corridors

Landslides along road corridors in Alaska national parks pose threats to public safety, visitor access, subsistence activities, and result in costly remediation of damaged infrastructure. Landslide risk in these areas, which contain near-surface permafrost, is associated with mean annual air temperatures (MAATs) above freezing and heavy precipitation events. Historical (1981–2020) values of MAAT and summer precipitation (JJA PCPT) from the fifth generation European Centre for Medium-Range Weather Forecasts (Reading, UK) atmospheric reanalysis (ERA5) were compared to mid-century (2021–2060) and late-century (2061–2100) downscaled climate model projections across Gates of the Arctic National Park and Preserve (GAAR), Denali National Park and Preserve (DENA), and Wrangell-St. Elias National Park and Preserve (WRST). ERA5 showed that all locations historically had MAAT values below freezing, but all three parks were warming significantly (0.3–0.6 °C per decade). Observed trends of MAAT from 18 stations showed warming trends with 11 of the 18 being significant at the 95% confidence level using the Mann–Kendall non-parametric test. Road corridor values are given for the: (1) proposed Ambler Road through GAAR, (2) Denali Park Road in DENA, and (3) McCarthy Road in WRST. Elevated risk from MAAT was projected in the mid-century period for the Denali Park Road and McCarthy Road and across all three park road corridors in the late-century period; elevated risk from JJA PCPT was projected in all periods for all road corridors.

Provenance of Anthropogenic Pb and Atmospheric Dust to Northwestern North America.

Industrial activities release aerosols containing toxic metals into the atmosphere, where they are transported far from their sources, impacting ecosystems and human health. Concomitantly, long-range-transported mineral dust aerosols play a role in Earth’s radiative balance and supply micronutrients to iron-limited ecosystems. To evaluate the sources of dust and pollutant aerosols to Alaska following the 2001 phase-out of leaded gasoline in China, we measured Pb-Sr-Nd isotopic compositions of particles collected in 2016 from snow pits across an elevational transect (2180–5240 m-a.s.l) in Denali National Park, USA. We also determined Pb flux and enrichment from 1991–2011 in the Denali ice core (3870 m-a.s.l). Chinese coal-burning and non-ferrous metal smelting account for up to 64% of Pb deposition at our sites, a value consistent across the western Arctic. Pb isotope ratios in the aerosols did not change between 2001 and 2016, despite the ban on lead additives. Emissions estimates demonstrate that industrial activities have more than compensated for the phase-out of leaded gasoline, with China emitting ∼37,000 metric tons year–1 of Pb during 2013–2015, approximately 78% of the Pb from East Asia. The Pb flux to Alaska now equals that measured in southern Greenland during peak pollution from North America.

The first documentation of the Nearctic-Paleotropical migratory route of the Arctic Warbler.

The Arctic Warbler (Phylloscopus borealis) is a cryptically plumed songbird with an uncommon Nearctic–Paleotropical migratory strategy. Using light‐level geolocators, we provide the first documentation of the migratory routes and wintering locations of two territorial adult male Arctic Warblers from Denali National Park and Preserve, Alaska. After accounting for position estimation uncertainties and biases, we found that both individuals departed their breeding grounds in early September, stopped over in southeastern Russia and China during autumn migration, then wintered in the Philippines and the island of Palau. Our documentation of Arctic Warbler wintering on Palau suggests that additional study is needed to document their wintering range. Our study provides hitherto unknown information on stopover and wintering locations for Arctic Warblers and indicates that this species may migrate further overwater than previously thought.

Surface water area in a changing climate: Differential responses of Alaska’s subarctic lakes

Lake surface area in arctic and sub-arctic Alaska is changing in response to permafrost deterioration, changes in precipitation, and shifts in landscape hydrology. In interior Alaska, the National Park Service’s Central Alaska Network Shallow Lakes program studies lakes and ponds in a wide range of geomorphological settings ranging from alpine lakes to low lying lakes on fluvial plains. The purpose of this study was to determine if and how lake area was changing across this diverse environment. Using the USGS Dynamic Surface Water Extent product, we tested landscape-scale trends in surface water area from 2000–2019 in 32 distinct ecological areas, or ecological subsections, within the three parks. Surface water area declined in 9 subsections, largely in glaciated landscapes with coarse substrates and areas underlain by ice-rich permafrost. Surface water increase was seen in one subsection in the floodplain of the Copper River in Wrangell-St. Elias National Park. No net change was observed in many subsections, but individual lake analysis showed that within several ecological subsections some lakes were increasing in area while others decreased in area, masking changes in lake surface area within the subsection. Over the course of the study period, surface water area in all parks experienced similar fluctuations, likely due to oscillations in regional climate. Periods of high surface water area coincided with relatively warm, wet periods. Climate change models project increases in both temperature and precipitation in Alaska; our results suggest periods of regional wetting may mask longer-term declines in surface water area in some geomorphological settings. Overall, lake surface area declined over the study period; declines were greatest in the Glaciated Lowlands in Denali National Park and Preserve.

Operational Processing of Big Satellite Data for Monitoring Glacier Dynamics: Case Study of Muldrow Glacier

Frequent acquisition of Synthetic Aperture Radar (SAR) data by the European Sentinel-1 satellites provides an opportunity for monitoring the dynamics of worldwide glaciers. We present a fully-automated processing system for producing multi-dimensional time series of glacier flow. We then use this fully-automated processing system to investigate the dynamics of Muldrow Glacier, located in the Denali National Park and Preserve (Alaska, AK, USA) during the October 2014—November 2021 period. We compute north, east, and vertical Surface-Parallel-Flow (SPF) and non-Surface-Parallel-Flow (nSPF) components of flow velocity and displacement with an average temporal resolution of 9 days and grid spacing of 100 m. During this period, we observe a glacier surge, a manifold increase in glacier flow velocity, that started as early as 2017 and continues until the present; however, the near completion of this surge is apparent. This glacier previously surged in 1906–1912 (the exact date is unknown) and in 1956–1957. We present our results in different ways to emphasize various aspects of the observed surge and demonstrate the full capability of our processing system. As the availability of SAR data improves, we expect that the fully-automated processing systems, similar to the one presented here, will play an increasingly dominant role and soon entirely replace manual processing.