Defoliation Date Research Articles

The Impact of Accumulating Herbage Masses in Autumn on Perennial Ryegrass Sward Characteristics

Autumn grazing management aims to accumulate herbage for defoliation prior to a decrease in growth rates for the extension of the grazing season. The current study investigated the impact of building different target herbage masses (THMs) in autumn and imposing one of three different defoliation dates (DDs) between mid-October and late November on light transmitted to the base of the sward, free leaf lamina (FLL), leaf stage and internode elongation. Four THMs (low, medium, high and very high) and three DDs (DD1—15 October, DD2—7 November and DD3—21 November) were assigned to a 4 × 3 split plot design over two years. Light transmitted to the base of the sward was greatest in the low THM and decreased in all other THMs. Internode elongation increased in tillers in the medium to the very high THMs. Defoliation of the medium, high and, in particular, the very high THMs earlier in autumn reduced the effect of decreased light transmission on internode elongation. This study highlights that, as light transmitted to the base of the sward decreases, internode elongation increases, and this could negatively impact sward structure.

A superior genetic source for late leafing in walnut 'Ahir Nut'

One of the main objectives of walnut crossbreeding programmes is to produce cultivars with late leafing dates. Sixteen years ago, a walnut crossbreeding programme was initiated by Prof. Mehmet Sütyemez, and now a new walnut genotype, namely ‘Ahir Nut’, has been generated with a leafing date that starts later than all commercially-famous walnut cultivars in the world. The present study describes this walnut genotype. For a detailed analysis, ‘Ahir Nut’ was compared with two French-origin walnut cultivars, i.e., ‘Franquette’ and ‘Ronde de Montignac’, which have had the latest leafing dates until now. In an experimental orchard in Turkey, the leafing date in ‘Ahir Nut’ was 14 May, whereas the leafing date in ‘Franquette’ and ‘Ronde de Montignac’ were 23 April and 25 April, respectively. This approximately 3-week difference between ‘Franquette’ and ‘Ronde de Montignac’ to ‘Ahir Nut’ reveals how valuable this genotype is for walnut breeding studies. The harvest dates for ‘Ahir Nut’, ‘Franquette’ and ‘Ronde de Montignac’ were 4 October, 3 October, and 7 October, respectively. The defoliation date for ‘Ahir Nut’ was 4 December. The nut weight and kernel percentage of ‘Ahir Nut’ were calculated as 11.61 g and 49.84%, respectively. Remarkable phenological traits were found in ‘Ahir Nut’, making it a promising option for future walnut cultivation. In the present study, the late leafing date of ‘Ahir Nut’ prompted further research into the molecular and walnut crossbreeding potentials. This new genotype has been conserved as an important genetic resource for future walnut breeding programmes around the world.

Morphology and forage quality in buffel, rhodes, and blue grama grasses in Valle del Mezquital

This study aims to evaluate the production and quality of buffel (Pennisetum ciliare), rhodes (Chloris gayana), and blue grama (Bouteloua gracilis) forage with three defoliation dates —at 50, 80, and 110 days after regrowth (dar)— in Valle del Mezquital. Crude protein (CP), neutral detergent fiber (NDF), acid detergent fiber (ADF), total digestible nutrients (TDN), relative feed value (RFV), and metabolizable energy (ME) were determined. A completely randomized design with three repetitions, a Tukey test for mean separation (α 0.05), and SAS Proc GLM for data analysis were used. The highest forage production occurs in the rhodes forage at 110 dar (12,936 kg DM ha-1). The highest CP (10.6%) was found in the buffel forage at 50 dar; in the rhodes forage, both values (80 and 110) were lower than 7%. The highest RFV was obtained by exotic grasses; however, higher TDN was recorded for the blue grama grass. Rhodes grass obtained the lowest ME at 80 dar (1.76), while the highest ME was obtained by buffel at 50 dar (1.91). The three varieties can be defoliated when the plants show intermediate yield and nutritional value, i.e., 80 days after the regrowth begins; nevertheless, buffel and rhodes should be defoliated at 50 dar.

Cevizde açık tozlanmış tohumlardan elde edilen genotiplerde fenolojik ve moleküler karakterizasyonun belirlenmesi üzerine bir araştırma

This research was carried out to reveal the phenological and genetic differences between the S-1/1 walnut genotype and its 94 progenies. In the phenological observations made, it was observed that budburst in genotypes took 52 days, leafing 50 days, leaf yellowing 31 days, and defoliation date 27 days. When the mother plant (S-1/1) and the genotypes were compared, it was found that there was a phenological variation of 75.54% in budburst, 73.41% in the leafing, 34.05% in leaf yellowing, and 93.62% in defoliation date, while the average variation was 69.15%. In the dendrogram obtained using phenological data, 2 main and 5 subgroups were obtained. In molecular genetic analyzes, 7 ISSR primers were used to determine genetic variations, as a result, 7 monomorphic and 45 polymorphic bands were obtained, and the rate of polymorphism was found to be 86.53%. The average number of alleles was calculated to be 7.42. In genotypes, the polymorphism information content (PIC) value varied between 0.48 and 0.95, while the average PIC value was calculated to be 0.73. As a result of cluster analysis, it was seen that genotypes were divided into 2 main clusters and 2 subsets. At the end of the study, it was determined that the S-1/1 and its progenies have a significant variation both phenologically and genetically.

Variation in Onset of Leaf Unfolding and Wood Formation in a Central African Tropical Tree Species

A diversity of phenological strategies has been reported for tropical tree species. Defoliation and seasonal dormancy of cambial activity inform us on how trees cope with water stress during the dry season, or maximize the use of resources during the rainy season. Here, we study the matching between leaf phenology (unfolding and shedding) and cambial activity for Prioria balsamifera, a key timber species in the Democratic Republic of Congo. In particular, we (i) evaluated the seasonality of cambial activity and synchrony of phenology among trees in response to climate and (ii) identified the seasonality of leaf phenology and its relation with cambial phenology. The study was conducted in the Luki Man and Biosphere Reserve, located in the Mayombe forest at the southern margin of the Congo Basin. Historic defoliation data were collected every ten days using weekly crown observations whereas recent observations involved time-lapse cameras. Cambial pinning was performed on ten trees during 20 months and radius dendrometers were installed on three trees during 13 months. Tree rings were measured on cores from 13 trees and growth synchrony was evaluated. We found that P. balsamifera defoliates annually with a peak observed at the end of the dry season and the beginning of the rainy season. The new leaves unfolded shortly after shedding of the old leaves. The peak defoliation dates varied across years from September 12 to November 14 and the fraction of number of trees that defoliated at a given time was found to be negatively correlated with annual rainfall and temperature; during the dry season, when precipitation and temperatures are the lowest. Wood formation (radial growth), was found to be highly seasonal, with cambial dormancy occurring during the dry season and growth starting at the beginning of the rainy season. Individual ring-width series did not cross date well. The within species variability of leaf phenology and cambial rhythms provides indication about resistance of the population against climatic changes.

Regional-scale data assimilation with the Spatially Explicit Individual-based Dynamic Global Vegetation Model (SEIB-DGVM) over Siberia

This study examined the regional performance of a data assimilation (DA) system that couples the particle filter and the Spatially Explicit Individual-based Dynamic Global Vegetation Model (SEIB-DGVM). This DA system optimizes model parameters of defoliation and photosynthetic rate, which are sensitive to phenology in the SEIB-DGVM, by assimilating satellite-observed leaf area index (LAI). The experiments without DA overestimated LAIs over Siberia relative to the satellite-observed LAI, whereas the DA system successfully reduced the error. DA provided improved analyses for the LAI and other model variables consistently, with better match with satellite observed LAI and with previous studies for spatial distributions of the estimated overstory LAI, gross primary production (GPP), and aboveground biomass. However, three main issues still exist: (1) the estimated start date of defoliation for overstory was about 40 days earlier than the in situ observation, (2) the estimated LAI for understory was about half of the in situ observation, and (3) the estimated overstory LAI and the total GPP were overestimated compared to the previous studies. Further DA and modeling studies are needed to address these issues.

The Impact of Herbage Mass on Perennial Ryegrass Swards in Autumn on Autumn and over Winter Production and Characteristics

Accumulating herbage mass to facilitate the extension of the grazing season in autumn is commonly practised. The objective of the current study was to investigate the effect of accumulating varying target herbage masses (THM) in autumn and imposing different defoliation dates (DD), on herbage mass, sward quality and water-soluble carbohydrates in autumn and the subsequent spring. A 4 × 3 factorial split plot design was assigned with four THM (Low ≈ 500 kg·DM·ha−1, Medium ≈ 1500 kg·DM·ha−1, High ≈ 2000 kg·DM·ha−1 and Very high ≈ 3000 kg·DM·ha−1) and three DD (DD1—15 October, DD2—7 November and DD3—21 November), across two years. Measurements were carried out at each DD and in spring. Differences in sward quality were found between each THM on different DD. Sward quality reduced from DD2 to DD3 in the high THM (−13 g·kg−1 DM CP, p < 0.001). The very high THM had the lowest sward quality from DD1 (206 g·kg−1 DM CP, p < 0.001 and 787 g·kg−1 DM DMD, p < 0.05). This study has identified the defoliation date of THM in autumn as key to improving autumn management strategies for increased utilisation and sward quality.

The effects of first defoliation and previous management intensity on forage quality of a semi-natural species-rich grassland.

Semi-natural grasslands occupy large parts of the European landscape but little information exists about seasonal variations in their nutritive value during the growing season. This paper presents results of novel data showing the effect of 13 years of previous contrasting management intensities on herbage nutritional value in relation to different dates of first defoliation (by grazing or haymaking). The treatments were: extensive management and intensive management from previous years (1998–2011). Both treatments were cut in June followed by intensive/extensive grazing for the rest of the grazing season (July–October). To evaluate forage quality in the first defoliation date, biomass sampling was performed in the year 2012 for 23 weeks from May to mid-October, and in 2013 for seven weeks from May to mid-June. Sampling was performed from plots that were not under management during the sampling year. Previous extensive management was associated with significantly reduced forage quality for in vitro organic matter digestibility (IVOMD), crude protein, neutral detergent fibre, acid detergent fibre and reduced divalent cations (Ca, Mg) and Na during the first seven weeks of the grazing season and the forage was suitable only for beef cattle. Due to low forage IVOMD, the forage is suitable only for cattle maintenance or for low quality hay when the start of grazing was postponed from seven weeks of vegetative growth to 13 weeks, regardless of the previous intensity. Herbage harvested after 13 weeks of the grazing season was of very low quality and was unsuitable as a forage for cattle when it was the only source of feed. Agri-environmental payments are necessary to help agricultural utilisation to maintain semi-natural grasslands by compensating for deterioration of forage quality, not only for the postponement of the first defoliation (either as cutting or grazing) after mid-June, but also when extensive management is required.

Severe insect defoliation at different timing affects cell wall formation of tracheids in secondary xylem of Larix kaempferi

Severe insect defoliation induced non-lignified tracheids and thin-walled latewood tracheids in the defoliation year in Larix kaempferi. These anomalous tracheids had heterogenous lignin deposition of S2 layer. Severe insect defoliation induces the formation of light ring consisted of thin-walled latewood tracheids in larch trees, which would affect wood quality. However, the identity of the secondary wall layers affected by insect defoliation remains unclear. The aim of this study is to clarify the effects of insect defoliation on the formation of secondary cell walls of tracheids in Larix kaempferi with focus on the defoliation timing. We observed by light microscopy and transmission electron microscopy (TEM) the secondary cell walls of tracheids produced in a defoliation year in L. kaempferi trees on which needles were attacked in July (gypsy moth, GM samples) or August (larch sawfly, LS samples). Light microscopy observations revealed that GM samples produced non-lignified tracheids in the transition zone between earlywood and latewood, as well as thin-walled latewood tracheids, and non-lignified tracheids were observed near the cambial zone in LS samples following defoliation for two consecutive years. TEM observations revealed that the tracheids in both samples had anomalous layers in secondary walls such as thin S2 layers with heterogeneous lignin deposition. The production of non-lignified tracheids corresponded to defoliation date in both samples, unlike changes in the layer composition of anomalous tracheids. Our study suggests that changes in wood structure depend on the date of insect defoliation and that insect defoliation affects the formation of secondary cell walls of tracheids, presumably in response to inadequate photosynthate supply as a result of defoliation.

Phenological response of peach to climate change exhibits a relatively dramatic trend in China, 1983–2012

The climate has warmed dramatically over recent decades, especially since the 1980s. Global climate change impacts can already be tracked in many biological systems. Plant phenology is often used as a good indicator for climate changes. Previous analyses have stated an advance of spring events, a delay of autumn events, and a lengthening of the growing season over recent decades. Here, we continuously observed the bloom date and defoliation date of seven peach varieties from 1983 to 2012. We demonstrate that the bloom date of peach has advanced by 11.1 days, whereas the defoliation date has been delayed by 8.7 days. These observations suggest that the growing season has lengthened by 19.8 days over 30 years. The climate data indicates that the annual mean temperature has warmed by 0.67°C per decade at NPGRC since the 1980s. The bloom date was negatively correlated with the average air temperature from February to April (r=−0.83, p<0.01), indicating that higher temperatures from February to April may accelerate developmental processes and finally lead to an advance of spring events. To verify how winter warming influenced the spring event, we designed different winter chill units treatments to simulate different degrees of warming conditions. Simulative experiments indicated that the decrease in winter chill delayed the bloom date, which also suggests that the warming from February to April was the major reason for the advancing bloom date. Our results show a more dramatic trend of phenological change than other previous studies. This difference may be because our observations were focused on local climate changes and a relatively temperature-sensitive species. Our studies found a different pattern of phenological changes and could provide insights into local climate change research.

Leaf damage effects on leaf expansion timing in Mallotus japonicus (Euphorbiaceae)

AbstractFor many plant species, biotic factors determining the timing of leaf expansion have not been elucidated sufficiently. We investigated the effects of leaf damage on the timing of leaf defoliation and on the timing of leaf expansion in Mallotus japonicus (Euphorbiaceae). The degree of leaf herbivory of M. japonicus in the field was examined in summer, with subsequent investigation of the date of leaf defoliation in autumn and that of leaf expansion the following spring. Effects of artificial leaf damage on the timing of leaf defoliation and of leaf expansion were also examined in a greenhouse. In the field, the degree of leaf herbivory sustained by M. japonicus did not affect the timing of defoliation. However, the timing of leaf expansion the next spring advanced earlier, concomitantly with the increase of leaf damage during the previous year. Fifty per cent artificial leaf damage in M. japonicus saplings in summer had no marked effect on the timing of leaf defoliation in autumn. However, the effects of leaf damage on the timing of leaf expansion were apparent the following spring: the damaged plants expanded new leaves ca. 8 days earlier than control plants did. Plants that showed earlier leaf expansion tended to have a higher shoot/root ratio. Our results demonstrate that the plants advance the timing of their leaf expansion in response to leaf damage sustained during the previous year, suggesting that the shoot/root ratio is a determining factor.

Early season dry matter production of three hybrid ryegrass (Lolium boucheanum) and two perennial ryegrass (Lolium perenne) cultivars

Hybrid ryegrass (HRG) is purported to have high overwinter growth characteristics and may have a role in Irish grassland systems. In a 3-year small-plot study at Fermoy, Cork, Ireland, the dry matter (DM) production and herbage quality of HRG cultivars in early spring were compared with diploid and tetraploid perennial ryegrass (PRG) cultivars. Treatments comprised ryegrass cultivars × defoliation managements × defoliation heights in a 5 × 3 × 2 factorial arrangement. The cultivars were Navan (tetraploid PRG), Foxtrot (diploid PRG), Foyle (tetraploid HRG), Brutus (diploid HRG) and Belleek (tetraploid HRG). Defoliation dates were: (A) 2 February and 20 April; (B) 2 February, 3 March and 20 April; and (C): 2 February, 23 March and 20 April. Defoliation heights were 4 and 7 cm, applied across each management. The HRG cultivars yielded similarly to cv. Navan on 2 February. Cumulative spring DM production was higher in cvs. Brutus and Belleek than all other cultivars. During this period, the managements A and B had similar DM yields (2765 kg DM ha−1) but were greater than management C. Defoliating swards to 4 cm height increased DM yield compared with 7 cm height. Organic matter digestibility of grass at the silage-harvest growth stage was greater for PRG cultivars. The study indicates that early spring growth of the three HRG cultivars evaluated here was not sufficiently greater than the tetraploid PRG cv. Navan to compensate for their poorer midseason sward quality.

The effects of simulated summer‐to‐winter grazing management on herbage production in a grass–clover sward

AbstractThe effects of summer‐to‐winter simulated grazing management factors, namely defoliation interval (INT: 21, 42, 56 or 84 d), defoliation height (DH: 2·7, 3·6, 5·3 or 6·0 cm) and final defoliation date (FIN: 23 September, 4 November or 16 December) on herbage production in a grass–clover sward were studied. Treatments were imposed between July and December 2008, with all plots under common management in the following March to June 2009. The 42‐d INT achieved the highest (P &lt; 0·001) total herbage yield at 11·00 t DM ha−1. Shorter (21 d) and longer (56–84 d) intervals reduced annual clover herbage yield and biological nitrogen fixation estimates. Lowering DH from 6·0 to 2·7 cm in the summer‐to‐winter period increased sward clover content and clover herbage yield through to the following June, 6 months after treatments ended. Delaying FIN from 23 September to 16 December had no significant effect on annual clover, grass or total herbage yield. Spring–summer clover herbage yield was positively correlated with spring–summer clover stolon mass (R2 = 0·54, P &lt; 0·001) and, to a lesser extent, light penetration through the sward in the previous winter (R2 = 0·16, P &lt; 0·05). A 42‐d INT with low DH (2·7–3·5 cm) is therefore recommended for grass–clover swards.

Individual Shoot Defoliation Minimally Affects Pecan Return Bloom

Alternate bearing of pecan [Carya illinoinensis (Wangenh.) K. Koch] remains the leading problem of the industry. Several cultural practices have been developed or improved to mitigate alternate bearing. Premature defoliation was one problem identified that substantially decreased return bloom. The objective of this study was to determine the response of individual shoots exposed to various defoliation treatments. In one study, individual vegetative or bearing shoots were hand-defoliated in mid-September. Defoliation was the basal one-half, distal one-half, entire shoot, or not defoliated. Another study applied the same defoliation treatments to bearing shoots in July, August, or September. Defoliation had minimal effects on return bloom and rarely affected the percentage of current-season shoots fruiting the next year. Defoliation date also had little effect on return bloom. These data indicate that individual shoot response to defoliation was not autonomous and has implications for determining crop overload and needed mechanical fruit thinning.

A general Landsat model to predict canopy defoliation in broadleaf deciduous forests

Defoliation by insect herbivores can be a persistent disturbance affecting ecosystem functioning. We developed an approach to map canopy defoliation due to gypsy moth based on site differences in Landsat vegetation index values between non-defoliation and defoliation dates. Using field data from two study areas in the U.S. central Appalachians and five different years (2000, 2001, 2006, 2007, and 2008), we fit a sigmoidal model predicting defoliation as a function of the difference in the vegetation index. We found that the normalized difference infrared index (NDII, [Band 4−Band 5]/[Band 4+Band 5]) and the moisture stress index (Band 5/Band 4) worked better than visible-near infrared indices such as NDVI for mapping defoliation. We report a global 2-term fixed-effects model using all years that was at least as good as a mixed-effects model that varied the model coefficients by year. The final model was: proportion of foliage retained=1/(1+exp(3.057−31.483∗[NDIIbaseyear−NDIIdisturbanceyear]). Cross-validation by dropping each year of data and subsequently refitting the remaining data generated an RMS error estimate of 14.9% defoliation, a mean absolute error of 10.8% and a cross-validation R2 of 0.805. The results show that a robust, general model of percent defoliation can be developed to make continuous rather than categorical maps of defoliation across years and study sites based on field data collected using different sampling methods.

The timing of leaf fall affects cold acclimation by interactions with air temperature through water and carbohydrate contents

Three parameters (i.e. the water content, soluble sugar content and minimal air temperature) can be used to predict the cold acclimation process of walnut trees. In order to test this assumption, two-year-old walnuts were defoliated at two different dates, i.e. mechanical defoliation in early October (early leaf fall, EF) or natural defoliation in early November (natural leaf fall, NF) and conditioned in either outdoor freeze-deprived or cold-deprived ( T min > 13 °C) greenhouses over winter. Even if early defoliation date could have affected short day signal perception (SDSP), water balance and carbohydrate metabolism were more altered. EF treatment, by stopping transpiration, significantly increased tree's water content and at warm temperature high root activity stopped normal winter dehydration. Starch content decreased in all treatments, but there was only a significant increase in soluble sugar content when water content had sufficiently decreased. Thus, depending on date of defoliation, cold-deprived trees were or were not able to acclimate to frost (minimal frost hardiness = −21.8 °C vs. −22.1 °C in controls (freeze-deprived) for NF and −13.7 °C vs. −25.3 °C in controls for EF). Different treatments showed the relationship between minimal water content observed during winter and maximal soluble sugars synthesized. Thus, the cold acclimation process appeared dependent on these physiological parameters (water and soluble sugar contents) through the interaction between air temperature and timing of leaf fall.

Investigations of alternative kale management: Production, regrowth and quality from different sowing and defoliation dates

Gruner kale showed a linear increase (8.0 kg DM/ha per °Cd) in biomass with sowings on 1 October, 3 November and 1 December producing 23, 19 and 17 t DM/ha (respectively) by the 29 May. Regrowth following mid season defoliation was slow (5.3 kg DM/ha per °Cd) reducing total production (relative to undefoliated) by 7.5 and 5.5 t DM/ha for treatments defoliated on the 29 January and 14 March, respectively. Keyworks: defoliation, kale, quality, regrowth, sowing date, thermal time, yield

Freezing temperature treatment induces bud dormancy in ‘Granny Smith’ apple shoots

One-year-old, ‘Granny Smith’ apple ( Malus × domestica Borkh.) shoots were selected randomly from commercial orchards in 1999 and 2003, cold stored at temperatures between 1 and 13 °C for varied periods, following a 12/12 h freezing temperature pre-treatment of −1/13 °C (supposedly non-chilling temperatures) for 1 or 2 weeks. The rate of budburst, determined after forcing at 25 °C, was used to follow the progression of bud dormancy. The freezing pre-treatment clearly enhanced (deepened) bud dormancy in all experiments. Clearly definable influences due to the presence of leaves during the pre-treatment were not observed on shoots cold stored at chilling temperatures. Trees were defoliated before leaf drop in late summer/autumn under field conditions by hand (2001) and using a spray containing 3% urea and 1.8% zinc sulphate in 2003. In 2001 the first two hand defoliation dates significantly enhanced dormancy, while in 2003 chemical defoliation had no effect. These data indicate that the leaves are not clearly involved in the perception of factors that are responsible for dormancy induction. As with dormancy release it is possible that the perception of induction factors, i.e. low temperatures, and as shown in these data, freezing temperatures, occurs within the buds themselves.

Defoliation Impacts on Festuca campestris (Rydb.) Plants Exposed to Wildfire

Wildfires commonly occur in the Fescue Prairie of Alberta, but little information exists to provide a basis for making grazing recommendations after burning. A wildfire in April 1999 provided an opportunity to study the effect of season and intensity of post-burn defoliation on foothills rough fescue (F. campestris Rydb.) in southwestern Alberta. A 3 (date of defoliation) x 2 (defoliation intensity) factorial experiment with 10 replicates (plants) was established in both a burned and a non-burned grassland and analyzed as a nested design. Plants were defoliated once during active vegetative growth (17 May), inflorescence development (2 July), or dormancy (30 September), at either 5 or 15-cm clipped stubble heights in the first growing season after fire. Burning increased tiller numbers by 54% compared to non-burned plants but reduced plant ANPP by 51% in the second growing season. While a single defoliation of burned plants, particularly early in the year, had little effect on growth, delaying defoliation into July decreased tillers 1 year later. Increasing defoliation intensity had the greatest impact on non-burned plants, reducing plant height (15%) as well as tiller (21%) and plant (32%) ANPP in the second year. May defoliation reduced etiolated growth 1 year later regardless of burn treatment. A single grazing event after wildfire does not necessarily appear to detrimentally affect rough fescue; however, the low herbage available immediately after fire may not justify the increased risk to the plant with subsequent grazing. DOI:10.2458/azu_jrm_v56i4_bogen

Defoliation Regime Effects on Accumulated Season‐long Herbage Yield and Quality in Boreal Grassland

AbstractWithin boreal grasslands, little information exists on the effects of initial defoliation date, frequency, and intensity on accumulated herbage yield and quality. We investigated the effects of initial defoliation in May, June or July, at heights of 5, 10 or 15 cm, and repeated at 2‐, 4‐ or 6‐week intervals throughout the growing season. Harvested material was combined with year‐end residual biomass, and examined for herbage removed, crude protein (CP), crude protein yield (CPY) and neutral detergent fibre (NDF). Compared to single defoliated check plots, total, grass and forb dry matter (DM) yields were lower under repeated defoliation by 25, 38 and 17 %, respectively. The majority of total herbage produced was harvested in the spring clipping. Total, grass and forb DM yields peaked under moderate (10 cm) clipping. Total DM and grass biomass were maximized with long (6 week) recovery periods. In contrast, forb biomass was greatest with May defoliation followed by a 4‐week interval. While maximum grass CPY was found under 10‐cm defoliation, forb CPY peaked with early and moderate to intensive defoliation. These results indicate that season‐long herbage biomass, along with CPY, can be maximized in boreal grasslands through controlled defoliation.